A color photo-catalogue of all known African species and subspeciesof Stag Beetles. This book treats the whole African continent, includingMadagascar and small islands: 125 species of 16 genera from tropicalAfrica and North Africa are listed and figured. For all species andsubspecies the detailed distribution is given, and for some speciesphotos of habitats and living specimens in the field are included.In addition, portraits and short biographical information aboutentomologists and explorers who worked particularly with the Africanlucanid fauna are also given. The species richness of the variouscountries and the perspectives for further research are discussed.For each African country all the known species are listed, togetherwith the number of endemisms. Author, year, publication, synonyms,size, type locality, abundance, remarks, and distribution are given forevery taxon.The label data of all the illustrated specimens are cited,as well as the acronym of the collection where the specimen is held.Many of the type specimens from the most important entomologicalmuseums (Berlin, Pretoria, Eberswalde, London, Paris, Cape Town,Munich, Dresden, Tervuren, Vienna, Florence, Genova and Basel)are illustrated. A large amount of bibliographical references are givenat the end.

More then 400 color photographs of all the speciesand subspecies from the African continent,as well as photos of living specimens and habitats- 192 pages, luxurious hardcover edition, largeformat 23x31, text in English

10

fig. 1 fig. 2

fig. 3 fig. 5fig. 4

Hermann Julius KolbeHenry BoileauEugene SéguyBernard BeneshSebastian Endrödy-YoungaHugues Eli BomansMarc DesfontainePhilippe MorettoJean Michel MaesTetsuo MizukamiShinya Kawai

`

fig. 1 - Hermann Julius Kolbe, fig. 2 - Henry Boileau, fig. 3 - Eugene Séguy, fig. 4 - Bernard Benesh, fig. 5 - Sebastian Endrödy-Younga`

11

fig. 6

fig. 7 fig. 8

fig. 9 fig. 10

fig. 6 - Jean Michel Maes, fig. 7 - Hugues Eli Bomans, fig. 8 - Marc Desfontaine (on left) and Philippe Moretto, fig. 9 - Tetsuo Mizukami,fig. 10 - Shinya Kawai (on right) here together with Robert Lízler (on left) and Karl Werner

22

Oonotus ( Macroonotus ) rex Endrödy-Younga

Publication: ENDRÖDY-YOUNGA, 1993: 34

Type locality: South Africa, Transkei, Port St. Jones

Size: m 19.1, f 15.7

Rarity: very rare

Remarks: only the type specimens are known.

The genus Oonotus has been revised by Endrödy-Younga (1993). R. Müller (pers.

comm.), who was with S. Endrödy-Younga when he collected the holotype, told us

that the specimen was slowly wandering on the bark of a tree in thick forest.

Distribution: South Africa (Transkei)

fig. 17 - m, 19.1 mm, South Africa, Transkei, Port St. Jones, 2.XII.1987, Endrödy-Younga leg., TMP, HOLOTYPUS

fig. 18 - f, 15.7 mm, South Africa, Transkei, Port St. Jones, 2.XII.1987, Endrödy-Younga leg., TMP, ALLOTYPUS

fig. 17 fig. 18

37

Figulus sublaevis (Palisot de Beauvois)

Publication: PALISOT DE BEAUVOIS, 1805: 3 (Lucanus)

Synonyms: anthracinus Klug, 1833: 85

ebenus Westwood, 1834: 119

nigritus Westwood, 1838: 261

monilifer Parry, 1862: 113

vulneratus J. Thomson, 1862: 433

lettowi Kriesche, 1921: 113

nossibenus Kriesche, 1921: 114

Type locality: “Regne Oware”

Size: 11.4 -17.4

Rarity: common

Remarks: according to Maes (1992a: 53) F. decipies is a subspecies of F. sublaevis

inhabiting only Sao Thomé Island, but according to Bomans (1992: 251)

and Monte & Bartolozzi (2004, in press) F. decipies is a valid species.

F. anthracinus seems to be a valid species too, but further studies are required.

On several occasions individuals of Figulus sublaevis were collected at light

by the second author and R. Lizler in open forests of southern and eastern Africa.

Distribution: Cameroon, Democratic Republic of the Congo (ex Zaire),Gabon,

Guinea, Malawi, Mozambique, Madagascar, Senegal, Tanzania, Togo

fig. 42 - 15 mm, Principe Isl., AKC fig. 43 - m, 17 mm, ex Coll. Borel, AKC

fig. 44 - m, 13 mm, Tanzania, Morogoro Prov., Uluguru Mts., 8. - 9.XII.1996, Werner & Lizler leg., KWC

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Figulus

fig. 42 fig. 43 fig. 44

26

Amneidus godefroyi Coquerel

Publication: COQUEREL, 1866: 326

Type locality: Is. Bourbon

Size: m 13.0 -15.0, f 12.0-15.0

Rarity: rare

Remarks: interesting remarks on this species are given by Paulian (1959, 1973).

Distribution: La Réunion Is.

fig. 26 - m, 15 mm, Réunion, I.1996, AKC fig. 27 - f, 15 mm, La Réunion, Kerveguen, 10.I.1962, AKC

fig. 26 fig. 27

53

Nigidius gnu Kriesche

Publication: KRIESCHE, 1921: 109

Type locality: D. Ost Afrika, Pangani, Pugu c/o Daressalam

Size: 13.0 -18.0

Rarity: very rare

Remarks: close to N. bubalus. Bomans redescribed this species

and established the lectotype.

Distribution: Tanzania

fig. 73 - 17.5 mm, D. O. Afrika, Pangani, ZMB, LECTOTYPUS

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Nigidius

fig. 73

82

fig. 119 - Madagascar, Marojejy, March 2001 fig. 120 - Madagascar, Ankarana, March 2001

fig. 121 - Madagascar, “Andasibe (Périnet)”, May 1998

fig. 119

fig. 120 fig. 121

83

fig. 122 - Madagascar, Morondava, April 1998 fig. 123 - Madagascar, Beza Mahafaly, April 1998

fig. 124 - Madagascar, Ranomafana, March 1998 fig. 125 - Madagascar, Ranomafana, March 1998

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Ganelius

fig. 124

fig. 123

fig. 122

fig. 125

84

fig. 126 - m, 25 mm, Ruanda, Ihembe, 11 km E de Dendezi, 2000km, 19.IX.1951, H. RENIER leg, IRSNB, PARATYPUS

fig. 127 - m, 24 mm, Zaire, Kahuzi, km 82, 1-15.IX.1951, legit H. Bomans (in coll. G. Taroni)

Dendezia renieri Basilewsky

Publication: BASILEWSKY, 1952: 37

Type locality: Rwanda, Dendezi

Size: m 23.0 -25.0, f [size not indicated in the description]

Rarity: very rare

Remarks: this species is very peculiar for the presence of a medial cephalic horn.

We are grateful to A. Drumont (IRSNB) for the fine pictures of this rare species.

B. Gill (pers. comm.) found a dead specimen of Dendezia renieri on a dirt road in

the Kibale forest of Uganda in October, 2001.

Distribution: SE Democratic Republic of the Congo (ex Zaire), Rwanda, Uganda

fig. 126

fig. 127

90

fig. 137 - m, 19 mm, Rep. of S. Africa, Cape, Swartberg, IX.1993, Lizler leg., AKC

fig. 137

Colophon berrisfordi Barnard

Publication: BARNARD, 1932: 173

Type locality: South Africa, Zwartberg Range, Meiringspoort Berg

Size: m 19.0 -22.0

Rarity: very rare

Remarks: the genus Colophon has been revised by Endrödy-Younga (1988)

and Mizukami & Kawai (1997). R. Lizler (pers. comm.) collected a single male

of C. berrisfordi among several specimens of C. whitei under stones.

Distribution: South Africa (Swartberg)

92

fig. 140 - m, 17.5 mm, Rep. of S. Africa, Cape, Meiringspoortberg, I.1994, AKC

fig. 141 - f, 16 mm, Rep. of S. Africa, Cape, Meiringspoortberg, I.1994, AKC

fig. 141fig. 140

Colophon cassoni Barnard

Publication: BARNARD, 1932: 172

Type locality: South Africa, Zwartberg Range, Meiringspoort Berg

Size: m 16.0 - 24.0, f 16.0 -17.0

Rarity: very rare

Remarks: the genus Colophon has been revised by Endrödy-Younga (1988)

and Mizukami & Kawai (1997). This species is active only during proper weather

conditions.

Distribution: South Africa (Swartberg)

93

fig. 142 - Swartberg Mts., Rep. of S. Africa, March 1993 fig. 143 - C. haughtoni - m, Rep. of S. Africa, Cape, Hexriver Mts., Matroosberg, I. 1993

fig. 144 - C. eastmani nagaii - m, Rep. of S. Africa, Cape Province, Langeberg, Dassie'shoek, 29. I. 1993,

fig. 145 & 146 - C. izardi - m, Rep. of S. Africa, Cape Province, Langeberg, Boosmansbos, 27. XII. 1992

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Colophon

fig. 142

fig. 145 fig. 146

fig. 143 fig. 144

142

fig. 268 - m, 32 mm, Congo Belge, Kasai, Kondue, Leohard leg., MRAC, PARATYPUS

fig. 269 - m, 28 mm, Congo Belge, Kasai, Kondue, Leohard leg., MRAC, PARATYPUS

Prosopocoilus kasaiensis Maes

Publication: MAES, 1990: 13

Synonym: swanzianus sensu Auctorum (partim)

Type locality: Congo Belge, Dekese, Kasai

Size: m 28.0 -32.0, f 20.0 - 22.0

Rarity: very rare

Remarks: this species was described as a form of P. swanzianus, then elevated

to subspecies by Maes (1992b); according to Desfontaine & Moretto (2003)

it is a valid species, very close to P. demeyeri, and can be separated only

by the examination of the genitalia.

Distribution: Democratic Republic of the Congo (ex Zaire)

fig. 268 fig. 269

143

fig. 270 - m, 34.5 mm, Kamerun, 1968, AKC

fig. 271 - f, 26 mm, Kamerun, 1968, AKC

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Prosopocoilus

Prosopocoilus katanganus Bomans

Publication: BOMANS, 1967a: 392

Type locality: Rép. du Congo, Kafakumba

Size: m 31.0 -34.5, f 20.0 - 26.0

Rarity: very rare

Remarks: this species is close to P. congoensis.

Distribution: Cameroon, Democratic Republic of the Congo (ex Zaire)

fig. 270 fig. 271

170

fig. 334 - m, 69 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC

fig. 335 - f, 38 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC

Prosopocoilus umhangi (Fairmaire)

Publication: FAIRMAIRE, 1891b: 192 (Cladognathus)

Synonyms: hacquarti Oberthür, 1880: 119

wentzelheckmannae Kolbe, 1901: 71 (Metopodontus)

elizabethae Waterhouse, 1905: 375 (Metopodontus)

Type locality: Zanzibar

Size: m 28.2 - 69.1, f 26.0 - 38.0

Rarity: not rare

Remarks: P. umhangi specimens were collected in numbers by the second author

and by R. Lizler on trees of rain forest in central Tanzania.

Distribution: Malawi, Tanzania

fig. 334 fig. 335

171

fig. 336 - m, 34 mm, Tanzania, Usambara, AKC fig. 337 - f, 26 mm, Tanzania, Usambara, AKC fig. 338 - m, 37 mm, Tanzania,

Uluguru Mts., IV.1997, Werner & Lizler leg., KWC fig. 339 - m, 67 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC

fig. 340 - f, 32.5 mm, D.O.A., Nkonda Uguro, 1910, AKC fig. 341 - m, 57 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Prosopocoilus

fig. 339 fig. 341fig. 340

fig. 336 fig. 338fig. 337

180

fig. 360 - m, 45 mm, Italia, Basilicata, Gallipoli-Cognato, 7.VII.1997, B. Cecchi & B. Carletti leg., LBC

fig. 361 - f, 32 mm, Sicilia, San Fratello, Messina Prov., 1.VII.1984, Aliquó leg., LBC

Lucanus tetraodon Thunberg

Publication: THUNBERG, 1806: 188

Synonyms: for the long list of synonyms see Krajcik (2001)

Type locality: Italy

Size: m 26.1-50.0, f 25.8 - 37.2

Rarity: common

Remarks: L. tetraodon is a palaearctic species occurring in oak forests. In North

Africa the species is quoted only from Algeria.

Distribution: Algeria; Balkan Peninsula, France (including Corsica),

S Italy (including Sicily)

fig. 360 fig. 361

181

fig. 362 - m, 38 mm, Italia, Calabria, Capistrano, Catanzaro Prov., 800 – 1000 m, VIII.1990, I. Bonato leg., LBC

fig. 363 - m, 28 mm, Italia, Calabria, Capistrano, Catanzaro Prov., 800 – 1000 m, VIII.1990, I. Bonato leg., LBC

fig. 364 - m, 42 mm, Sicilia, Messina Prov., P. Femmina Morta, 9.VII.1983, Aliquó leg., LBC

Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Lucanus

fig. 364fig. 362 fig. 363