FRAGMENTA PALAEONTOLOGICA HUNGARICA 24-25, BUDAPEST, 2007

Init ia l notes to a revision of Late Triassic gastropods from Budapest (Hungary): Hungariella K U T A S S Y , 1933 (Neritopsidae)

by János S Z A B Ó

Abstract — During the early years of the twentieth century, the Transdanubian Central Range in Hungary became a classical area for the exploration of the Triassic faunas. Several papers on gastropods were also published. As initial part of a series, planned on results about a revision of the gastropod faunas, this paper updates knowledge on Hungariella KUTASSY, 1933 (Neritopsidae) with some corrections. New name is given instead of "Neritopsis/Hungariella spinosa KUTASSY, 1927" that is a primary junior homonym of a iShddle Jurassic gastropod. Lectotypes and a neotype have been selected to the species, respectively.

Keywords — Hungariella, Neritopsidae, Gastropoda, Late Triassic.

SZABÓ, J.: Initial notes to a revision of Late Triassic gastropods from Budapest (Hungary): Hungariella KUTASSY, 1933 (Neritopsidae). — Fragmenta Palaeontologica Hungarica, 24-25: 69-75.

Introduction

Rather few localities are known with well preserved megafossils from the Late Triassic. Their significance has been recently raising with the increasing interest of the earth science researchers in the end-Triassic mass extinc­tion. The youngest fossiliferous sediments, deposited before the extinction, have become important sources in collect­ing information about the last, undamaged state of the biota, therefore the value of the previously published faunas is also increasing.

In Hungary, there are several prospective localities and faunas corresponding to the above criteria, however, from the point of view of gastropods, the importance of the north­western edge of Budapest and the adjacent western area has already been demonstrated. Gastropods were collected from two localities (Figure 1) of Late Triassic Dachstein Limestone, exposed in quarries of Remete-hegy (Norian) and Fazekas-hegy (Carnian), respectively. Larger part of Remete-hegy with fossiliferous outcrops extends westwards beyond the boundary- of Budapest to the area of Remete­szőlős, a neighbouring village. Megataunas of these localities, nearby each-other, were published by K U T A S S Y (1927, 1933, 1936)', B A R T K O (1939) and B É R C Z I N F M A K K (1969). These papers added lots of new information to the knowledge of the Late Triassic "Alpine" type (= Tethyan) faunas.

These faunas are principal sources also to find the roots of the post-Triassic (Early Jurassic) recovery, that is a not less interesting and important field of research. The author is especially interested in the latter aspect of studies within Class Gastropoda; that is why he has started working on Triassic gastropods. One part of his project aims to revise the originals that have become basic documents for the Triassic palaeontology with their publication from Hungary before 1940. Some ten years after their publications, usually even a simple systematic revision itself can result in new

data, however, experience shows that the original collec­tions usually contain lots of information that have still remained hidden.

The typical Dachstein Limestone developed in shallow water of high energy platform environment that was dis­advantageous for fossilisation. Normally megafossils are rare and hardly removable from the massive matrix. In the area of Remete-hegy and Fazekas-hegy, a local Dachstein Lime­stone variety lost the usual hardness and the fossils became relative easily cleanable from the chalky matrix. Another speciality of this facies is the rich, shelly, relatively well-

Figure 1 — Sketch map of the localities. — Asterisks indicate the abandoned quarries; dotted Une is the boundary of Budapest in the NW part of the cin. Distance of localities is about one kilometre.

70 SZABÓ, J .

preserved fauna, suggesting also a sedimentary environ­ment, atypical for the Dachstein Limestone. Large colonies of ramified corals indicate that this special component was most probably a platform margin position of the deposi-tional area with deeper, calm water bottom parts. However, the high abundance of the neritoideans shows shallow water platform origin at least for a part of the gastropod fauna that must have been allochtonously embedded here.

K U T A S S Y (1927, 1933, 1936) introduced abundant and diverse gastropod assemblages with numerous new taxa. Most of the new elements provide the fauna with endemic

1. Amauropsis macra]. Böl IM 2. Coelochrysalis rotundas Kl'TASSY 3. Coelostylina solida KOKEN et WOEHRMANN 4. Delphinulopsis triadica KUTASSY 5. Dicosmos (Fedaielld) dec/iris KlTTL 6. Dicosmos {Fedaielld) sp. 7. Dicosmos sigaretoides Kl'TASSY 8. Fossariopsis binodosa MÜNSTER 9. Fossariopsis pdljyi KUTASSY 10. F usus noricus BARTKÓ 11. Hologyra cassiana WlSSMANN 12. Hungariella pappi KUTASSY 13. Hunga rie/la spin osa KUTASSY 14. Hungariella stredae KUTASSY 15. Kokeniella pálfyi KL'TASSY 16. Callotrochus triadicus KUTASSY 17. Callotrochus triadicus Kl'TASSY var. elegantula Kl'TASSY 18. Katicella sublineata MÜNSTER 19. Neritaria turriculata KUTASSY 20. Neritaria plicatilis KLIPSTEIN 21. Xeritaria sp. 22. Omphaloptycha {Coelostylind) conica MÜNSTER 23. Omphaloptycha {Coelostylind} platistoma H AB ERLE 24. Omphaloptycha bacchus KlTTL 25. Omphaloptycha pupoides Sri )PPAN1 26. Omphaloptycha qtteli BÖHM 27. Ooniagappi KlTTL 28. Palaeonarica (?) rugoso carinata (KLIPSTEIN) 29. Palaeotriton (?) globu/aris Kl'TASSY 30. Parangularia hungarica KUTASSY 31. Platychilina Wöhrmann/ K( )KEN var. maior Kl'TASSY 32. Pleurotomaria sp.

Although the most important reason of the complete revision is the advance in gastropod systematics since the end of the 1930's years when KUTASSY has prepared his last publications, this work is urged also because KUTASSY's original material became rather decimated while it was (not really carefully) kept in one of the depositories, indicated in the publications. Recendy none of the speci­mens can be found in the collection of University Eötvös Loránd, Budapest (= University7 Pázmány Péter in the papers) because of rationalization. Major part of die material has been rescued to the Hungarian Geological Museum (part of the Hungarian Geological Institute). Further ruins of the original collection were presented to the Department of Geolog}7 and Palaeontology of the Hungarian Natural 1 liston Museum in another rescue action.

One of the new genera, established by KUTASSY (1933), is Hungariella, represented by three species in the faunas. Because no types were selected to the species, one of the aims of this paper is to find cither lectotypes or neotypes. The sparsest one of the species, based on a

characters but the major part of the supra-specific taxa clearly shows Alpine/Tethyan relation. B A R T K ( ' ) (1939) added the description of a single specimen of a peculiar new species, having siphonostomatous shell ("Fusus" noricus). The number of the published species level gastropod taxa arose up to around sixty7. The faunal list below is given in alphabetic order to demonstrate the most advanced taxo-nomical state [ K U T A S S Y ' s (1940); Glossophora Triadica II.)], however, there is obvious need of a revision both on the genus level (e.g. "Fttsus, "Tectus") and on the species level (e.g. to clear up what the "varieties" actually mean).

33. Protonerita mandelslohi KOPSTEIN 34. Prorermicularia sp. 35. Purpurina minima K i "i ASSY 36. Purpurina plicata KUTASSY 37. Purpuroidea excelsior Kc )KEN 38. Purpuroidea fereneçi Kl'TASSY 39. Purpuroidea nassaeformis DI STEFANO 40. Purpuroidea taramelli ST( )PPAN I 41. Purpuroidea turriculata KUTASSY 42. Solariella nodifera K l ITASSY 43. Stephanocosmia {Tyrsoecus) dolomiticus KlTTL 44. Stephanocosmia corona ta K O K E N 45. 'Pectus sp. 46. Te/leria gigantea KUTASSY 47. Felleria sp. 48. Trachynerita cf. nodifera KlTTL 49. Trachynerita nodifera KlTTL var. donga ta Kl'TASSY 50. Trachynerita quadrata (STOPPANI) 51. Trachynerita quadrata STOPPANT var. canaliculata AHLBURG 52. Trachynerita quadrata STOPPANI var. praeacuta KUTASSY 53. fretospira cf. fasciafa KlTi'l . 54. Undu/aria (Toxoconcha) cf. brocchii STOPPANI 55. If oehrmannia decora ta KUTASSY 56. If orthenia escheri S'l'( >PPANI 57. W'ortheuia gigas Kl'TASSY 58. W'ortheuia infraornata Kl'TASSY 59. W'ortheuia infini ST( )PPANI var. rotundocariuata Kl'TASSY 60. W'orthenia ornata Kl'TASSY 61. II ortheniopsis budensis Kl'TASSY 62. Zygopleura arcteocostata MÜNSTER 63. Zygopleura cf. a/padis KlTTL

single, not really well-preserved specimen, badly needs also a nomenclatorical revision. Correction of the name is the other purpose of this paper, planned as the first part of a full revision.

All available Hungariella specimens have been studied, but the types are selected from the collection of the Hun­garian Natural History7 Museum because it contains the only, doubtlessly recognisable originally figured specimen. K U T A S S Y (1927) established his "Neritopsis spinoscP on a single specimen, having characteristic shell damages, visible also in the original figures ( K U T A S S Y 1927, p. 153, pl. I l l , Figs 9a—b, and see them in Plate I : 18-20). The possibility for selection of a lectotype has been investigated also in cases of the other species, respectively. If existence of syntypes were not proven or probable, further specimens from the type locality7 are considered to select as neotype.

The chance to collect new material is low and decreasing, because the quarries in the area of the localities have been long inactive, and the other outcrops have become either a natural conservation area or built up.

Revision of Hungariella KUTASSY, 1933

Systematics

Abbreviations to measurements — H = total height; H L = height of last whorl; HP = height of peristome; D = diameter of last whorl; WP = width of peristome; AA = apical angle; AL = coiling angle of last whorl (if different from apical angle). Asterisks indicate reconstructed measurements taken on more (**) or less (*) damaged shell parts.

Class Gastropoda CUVIER, 1797

Subclass Neritaemorphi K O K E N , 1896 Order Neritopsina Cox & K N I G H T , 1960

Family Neritopsidae GRAY, 1847 Genus Hungariella KUTASSY, 1933

non: 1946: Hungariella ERDŐS

Type species —Neritopsis Pappi KUTASSY, 1927 = Hungariellapappi (KUTASSY, 1927).

(Re-)Description — Low turbiniform—subglobose shell, consisting of few (4—5), rapidly expanding whorls, separated by impressed, distinct suture. Spire rather pro­truding and gradate in outline. Whorl cross-section slightly ovate; axialfy elongated. Subsutural belt forms narrow, feebly concave to feebly convex shoulder with rounded abaxial edge. Aperture half circle-shaped with almost straight inner lip line, and angulation at suture. Peristome continuous, outer lip sharp, inner lip appears as moderately wide parietal callosity, extending wider over central part of base then turning into short columellar lip, having broad outer face with shallow, axial concavity7. Callosity7 has rounded thicken­ing at parietal region but thinner and feebly concave below; apertural edge sharply angular and feebly arched with abaxial concavity. Actually, columellar part of angula­tion forms narrow, rudimentary plate. Single dent-like process develops deeply behind peristome on wall of penultimate whorl. At suturai angle, narrow canal found. KUTASSY (1933) notes one or two smaller denticles that may appear also at foot of columella. Operculum unknown.

Ornament consists of low and wide transversal (but not collabral) costae or undulations, low spiral folds, spiral cords and threads; some of spiral ones corrugated by nodes of different size that may change also into spines. Nodes and spines may be arranged into opisthocline rows, resulting in wide, low, undulating costae. Nodes, spines and costae strongest at abaxial edge of shoulder, weaken towards suture and peripher)7; they may extend also onto base. In different forms but two more or less marked rows of nodes at and above outer edge of ramp appear in all known species.

Growth-lines usually delicate but thread-like in last growth stase.

Remarks — Hungariella resembles to Neritopsis GRATE-LOUP, 1832, however, there are important differences: — whorl cross-section and aperture are axially ovate in Hungariella but rather circular in Neritopsis; — aperture "D"-shaped in Hungariella but subcircular in Neritopsis; — columellar hp has sharp apertural rim (rudimentary plate) in Hungariella but rounded in Neritopsis.

Significant differences between the two genera are in the characters of the ornament. The rows of nodes and nodulae, especially at rim of ramp, are specific elements in the sculpture of Hungariella. Comparable ornament was found in the Carboniferous and Permian members of the family (Jrachydomia M E E K & WORTHEN, 1866 and Trachj-spira GEMMELLARO, G. G., 1889). These genera, similarly to Hungariella, have also usually more protruding and/or higher spired than the remaining Triassic neritopsids.

[For a new Hymenoptera genus, ERDŐS (1946) also applied the name "Hungariella" that has to be obviously substituted.)

Distribution — Hungariella species were found in Carnian and Norian strata of the Dachstein Limestone Formation in the Budai-hegység (= Buda Hills, the western part of Budapest, and the adjacent Hills along the W boundary of the city7). Three species has been established on the material from two localities (Remete-hegy7, Faze­kas-hegy), the two frequent ones (H. pappi and H. stredae) are known from Carnian and Norian beds and they have been identified also from the Dachstein Limestone of the South Karawanke Mountains near Trzic in Slovenia ("St. Anna bei Neumarkti"; KUTASSY 1934). The third, sparse species (H. "spinosa') is justifiable only from the Fazekas­hegy Carnian.

Hungariella pappi (KUTASSY, 1927)

(Plate I : 1-9)

1927: Neritopsis Pappi n. sp. — KUTASSY, p. 153, Pl. I l l , Figs lOa-b (non: 10c-d). 1933: Hungariella pappi KUTASSY — KUTASSY, p. 240, Pl. I , 7-9.

cf. 1934: Hungariella cfr. pappi Kl'TASSY — Kl'TASSY, p. 74.

Neotype — M 2007.8.L, Plate I : 6-9. Type locality — Remete-hegy, Budapest. Type strata — Upper Triassic (Norian) Dachstein Limestone Formation.

Material — Thirty-eight specimens (M 97.34. inventory N° refers also a H. stredae specimen, being inseparable).

72 SZABÓ , J.

Measurements H H L HP D WP AA AL M 97.34. Plate I : 1-2 *38 *36 *34 41 - 125° 110° M 2007.8.1. Plate 1: 6-9 *31 *28 *22 29 22.6 - 105° M 97.31. Plate I : 3 *29 *26.5 *24 *26.8 — 120° 115° M 2007.5.1. Plate 1: 4-5 *28.5 25 22 23.4 119° 110°

Description — Axially ovate, neritiform shell, con­sisting of 3—5 whorls, separated by incision-like suture. Protoconch unknown. Whorls convex, disregarding sub-suturai belt of flat or concave shoulder. Axial edge of ramp bends upward to previous whorl, thus resulting in narrow concave subsutural belt of variable distinctness (Plate I). Outer (abaxial) edge of ramp usually not marked, but frequently indicated by some change in most "prom­inent" ornamental elements being wide, low, orthocline costae, sometimes wrinkles or just weak undulation. They may run from suture to suture, or to outer edge of ramp from lower suture. In previous case, costae consisted of few indistinct nodulae on ramp. Costae and corresponding orna­mental elements cross growth-lines (—30°), being rather prosocline. Costae disappear at initial part of last whorl of largest (Padult/gerontic) specimens (Plate I : 1—2), only pen­ultimate whorl bears costae. Smaller specimens with ribbed shell (subadult? forms) seem more frequent (Plate I : 3—9).

Peristome and aperture of nearly half-circle shaped, slighdy ovate in suture to foot of columella clirection. Inner lip rather wide and extends as callosity, forming thick, rounded elevation in parietal region, and thin, feebly concave outer face at basal-columellar parts. Apertural edge of inner lip sharp, feebly concave line; basal-columellar part of inner Hp can be regarded as rudimentary aperture plate. Deeply behind inner Hp, weak, tooth-Hke process found on wall of penultimate wTiorl. Outer lip evenly

arched, its outer edge sharp. Distinct, narrow canal visible in suture angle of peristome.

Remarks — Unfortunately, the characteristically dama­ged original specimen, figured in K U T A S S Y 1927, Pl. I l l , figs 10 a-b, can not be found. Neritopsis pappi was estabHshed on four specimens; one of them (Pl. I l l , figs 10 c-d) belongs to H. stredae ( K U T A S S Y 1933, p. 241), the other ones are not identifiable amongst the specimens, available from the two collections. In lack of a verified syntype, selection of a neotype for Hungariella pappi has become necessary. Many of the subsequently found specimens are better preserved than the figured one. The available speci­mens are surely topotypic and a part of them might be identified by K U T A S S Y (1933) when Hungariella was outiined as new genus. The selected neotype shows most of the shell morphological characters, the other figured specimens give an impression about the variability7 of the sheU form and the ornament (Plate I : 1—9).

Specimens with weak ornament may be hardly separable from smooth genera of Naticopsinae. Other known species of Hungariella bear marked ornament of different com­ponents.

Distribution — Hungariella pappi has been found in both Budapest locaHties and (questionably) also in the South Karawanke Mountains (Slovenia, K U T A S S Y 1934); it occurred in Carnian and Norian strata of the Dachstein limestone.

Hungariella stredae KUTASSY, 1933

(Plate I : 10-17)

1927: Neritopsis Pippin, sp. — KUTASSY, p. 153, Pl. I I I , Figs lOc-d (non:10a-b). 1933: llitngariella stredae nov. sp. — KUTASSY, p. 241, Pl. I , Figs 10-14. 1934: Hungariella stredae KUTASSY—Kl'TASSY, p. 73, PL 11, Figs 8-10.

Lectotype — M 2007.4.1., Plate I : 12-13. Type locality — Remete-hegy, Budapest. Type strata — Upper Triassic (Norian) Dachstein Limestone Formation.

Material — Thirty7 specimens (M 97.34. inventory J\r° refers also a H. pappi specimen, being inseparable).

Measurements H H L HP D WP AA AL M 97.34. Plate I : 16-17 - - 34.2 102° 102e

2007.4.1. Plate I : 12-13 31 28.7 22.5 27 20.5 98° 98° 2007.7.1. Plate I : 10-11 - 31.3 - 100° 100e

2007.6.1. Plate I : 14-15 22.2 18.7 16.8 19 - 95° 95°

Description — Rather high spired, low turbiniform shell with rapidly expanding 4—5 whorls. No protoconch found. Whorls arched, disregarding narrow, flatten sub­sutural ramp having rounded outer angulation. Second, weak angulation visible at peripher}7, followed by suture of subsecjuent whorl. Base convex, narrow (false?) umbi-Hcus appears along basal Hp. Peristome shape and its structure similar to that of Hungariella pappi.

Differently from H. pappi, H. stredae has marked, charac­teristic ornament of considerable variability. Whole shell surface ornamented by spiral cords, and, in their inter­spaces, threads. AH cords and threads bear spiraUy elongat­ed nodes and nodulae or, at least, thickenings (Plate I : 10—17). Actually nodes and nodulae correspond morpho­logically to short, spiraUy arranged ribs and riblets. Nodes strongest at and somewhat below outer edge of subsu-

tural ramp. On penultimate whorl and at beginning of body-whorl (same shell part as in Hungariella pappi) nodes arranged into transverse costae that may persist to peristome in some specimens (Plate I : 16—17).

Remarks — The shape and the characteristic orna­ment distinguishes Hungariella stredae from the other species, discussed here (see also below).

The specimens, figured in KUTASSY (1933) can not be found in the remnants of the original collection. Therefore another gastropod must be selected as type. A part of the material stored in the collection of the Hungarian Natural History Museum (received from the Geological Depart-

ment of the Eötvös Loránd University) must belong to the syntypes. Based on the accompanied label, the speci­mens are topotypic (Remete-hegy, Norian), year of the collecting is 1919, and the collector is VADÁSZ, E. who has been employed also at the Geological Department of the Pázmány Péter University = Eötvös Loránd University. The specimens must have been available for KUTASSY in 1933 when Hungariella stredae was published.

Distribution — Hungariella stredae is found in both Budapest locality and also in the South Karawanke Mountains (Slovenia); this species occurred in Carnian and Norian strata of the Dachstein Limestone Formation.

Hungariella kutassyi nom. nov. (Plate I : 18-20)

non: I860: Neritopsis spinosa (nov. sp.) — H i - B E R T & D E S L O N G C H A M P S , p. 184, Pl. I , Figs 5a-d. 1927: Neritopsis spinosa n. sp. — K U T A S S Y , p. 153, Pl. I l l , Figs 9a—b. 1933: Hungariella spinosa K U T A S S Y — p. 243.

Lectotype — M 2007.2.1, Plate I : 18-20 (originally figured by KUTASSY (1927) as Neritopsis spinosa] Type locality — Fazekas-hegy, Budapest. Type strata — Upper Triassic (Norian) Dachstein Limestone Formation.

Material — Single damaged specimen.

Measurements H H L HP D WP AA AL 2007.2.1. - 26.2 23 31 20.2

Description — Damaged last whorl of neritiform shell with remnants of spire. Shallowly impressed suture sepa­rates whorls. Imprint on inner mould indicate same peristome and aperture morphology type, seen in other two species of Hungariella: half-circle ("D-shaped") outline with inner lip having almost straight apertural rim. Traces of inner lip on steinkern indicate rather wide, plate-like apertural edge.

Ornament consists of spiral threads, cords, and nodes that may become short, thick spines. Latter elements arranged into three rows on subsutural ramp; strongest row of spines visible along abaxial edge of ramp, weaker and weakest rows can be found adaxially from the first one in order of decreasing strength. Spines connected by carinae of different strength, correlating with size of spines. Irregularly some spines appeared also on adaxial shell edge bending up penultimate whorl. In KUTASSY's figure (1927, Pl. I l l , Figs 9a—b) spine present on strongest keel, running along ramp edge, but subsequently all spines broken away, now only scars visible at their place (Plate I :

19—20). Between ramp and base, regularly repeating, sparse

spiral cords and threads alternate. T w o cords mark edge o f

base, upper one w i t h stronger nodes, lower one w i t h smaller

nodulae. Subregulaiiv granulae may appear on all cords o f

preserved shell part o f last whor l . Growth-lines prosocline,

rather strong, thread-like.

Remarks — The spiny keels are distinctive characters

in a comparison either to H. pappi orH. stredae. The original name o f this species ('Neritopsis spinosa"),

given by KUTASSY (1927), is a junior pr imary h o m o n y m

o f the Midd le Jurassic Neritopsis spinosa HEBERT & DES­

LONGCHAMPS, 1860, therefore inva l id . Doubt less ly ,

KUTASSY's single specimen (monotype) represents a wel l

separable species therefore "'kutassyP is proposed here to

substitute the species name. The shape o f the damaged

shell parts, visible also in KUTASSY's figures, made the

identification o f the type specimen doubtless.

Distribution — The lectotype was found i n the Faze­

kas-hegy Carnian Dachstein Limestone.

Hungariella aff. kutassyi

(Plate I : 21-22)

Material — Single damaged specimen.

Measurements H H L HP D WP AA AL 2007.3.1 - 31.5 25.8 34.5 23.5

Description — Shape and measurements close to those of H. kutassyi, but shell has also remnants of penultimate whorl indicating spire height like in 17. pappi, being lower than in H. stredae. On ramp, this specimen has one more

(four, all together) spiny spiral cords and six to eight cords with coarse nodes between abaxial edge of ramp and centre of base. Number of nodosed cords increasing with growth of shell. These latter nodes also become spines

near last peristome. Between pairs of stronger spiral elements (cords), two threads visible, but only one in H. kutassyi.

Remarks — The morphology of the shell and the orna­ment shows close relation to Hungariella kutassyi, however, there are also important differences (more carinae with spines on the ramp; strongly nodosed cords on the last whorl and the base). These differences might be enough to establish a new species. However, decision must be post­poned because the specific variability7 of Hungariella kutassyi,

being unknown owing to the low specimen number, may cover also the morphology7 of Hungariella aff. kutassyi.

Distribution — The specimen was in the rescued material without label. The infilling of the shell and the preservation show that its locality is somewhere in the narrow distribution area of the chalky Dachstein Lime­stone facics in NW Budapest and the adjacent hills, most probably Remete-hegy or Fazekas-hegy; the age of the only Hungariella aff. kutassyi specimen must be regarded as Carnian—Norian.

* * * Acknowledgements — The study of old and recendy collected Triassic gastropods from the Transdanubian Central Range is supported by the Hungarian Scientific Research Fund, project Ns: OTKA T 042739.

References

B A R T K Ó , L. (1939): Fusus noricus nov. sp. a remetehegyi dachstein mész­kőből. (Fusus noricus n. sp. aus dem Dachsteinkalkstein des Remete-gebirges.) — Földtani Közlöny, 69: 196-198.

BÉRCZINÉ MAKK, A. (1969): A fazekas-hegyi felsőtriász ammonoideák (Die obertriadischen ammonoiden des Fazekas-Berges). — Földtani Közlöny, 99: 351-367.

E R D Ő S J. (1946): Genera nova et species novae Calcioidarum (Hym.). (Figura 14 originales in textu.) —Annales historico-naturales Musei nationalis hungariä, 39: 131-165.

HF.BF.RT, E . & DFSLOXGCHAIWS, E . (1860): Mémoire sur les fossiles de

Monu'euil—Bellay (Maine-et-Loire). — Bulletin, Société linnéenne de 'Nor­mandie, 5: 153-240, Pis 1-9.

KUTASSY, A. (1927): Beiträge zur Stratigraphie und Paläontologie die Alpinen Triasschichten in der Umgebung von Budapest. — A magyar királyi Földtani Intézet Evkönyve, 27: 107—175, Pis 1—6.

KUTASSY, A. (1933): Újabb adatok a Budapest környéki dachstemi mészkő faunájának ismeretéhez. (Weitere Beiträge zur Kentniss der Fauna des Dachsteinkalkes in der Umgebung von Budapest.) — Matematikai és Természettudományi Értesítő, 49: 222—250, Pis 1—2.

KUTASSY, A. (1934): A nóri dachsteinmész faunája Szt. Annán Neumarkd közelében (Felső Krajna) [Die Fauna des norischen Dachsteinkalkes von St. Anna bei Neumarkd (Oberkrain)]. — Földtani Közlöny, 56(4—6): 65-81, Pis 2-4.

KUTASSY, A. (1936): Fődolomit és Dachsteinmészkő faunák a Budai hegységből. (Faunen aus dem Hauptdolomit und Dachsteinkalk des Budaer Gebirges) — Matematikai és Természettudományi Értesítő, 54: 1006-1050, Pis 11-13.

KUTASSY, A. (1940): Glossophora triadica I I . — Fossilium Catalogus. F. Animalia., 81: 1- 477.

Authors' address Dr. J á n o s SZABÓ Department of Geology and Palaeontology Hungarian Natural History Museum Budapest, V I I I , Ludovika tér 2 Mail 1431 Budapest, pf. 137

Hungary E-mail: jszabo@nhmus.hu

Explanations to Plate I

1-2 Hungariella pappi ( K U T A S S Y , 1927) — x l ; spire (1) and back (2) view of largest specimen (M 97.34.) to show weakly developed costae.

3 Hungariella pappi ( K U T A S S Y , 1927) — x i ; spire view of specimen M 97.31. with sparse and strong costae.

4 - 5 Hungariella pappi ( K U T A S S Y , 1927) — x l ; spire (4) and back (5) view of specimen 2007.5.1. with denser costae.

6 - 9 Hungariella pappi ( K U T A S S Y , 1927) — x l ; neotype (M 2007.8.1.) in spire (6), back (7) and apertural (8) view; oblique apertural view (9)

shows place of inner denticle.

10-11 Hungariella Stredae K U T A S S Y , 1933 — x l ; back (10) and spire (11) view of 2007.7.1. specimen; nodes are arranged into costae only on juvenile shell.

12-13 Hungariella Stredae K U T A S S Y , 1933 — x l ; lectotype (2007.4.1.) in back (12) and apertural (13) view; nodes elongated on last whorl and not arranged into costae.

14-15 Hungariella Stredae K U T A S S Y , 1933 — X l ; back (14) and spire (15) view of 2007.6.1. young specimen; nodes more globular and arranged into costae.

16-17 Hungariella Stredae K U T A S S Y , 1933 — x l ; spire (16) and back (17) view of 97.34. specimen to demonstrate nodes are sometimes

arranged into costae also in adults.

1 8 - 2 0 Hungariella kutassyi nom. nov. — x l ; lectotype (2007.2.1.) in aperture (18), back (19) and spire (20) view.

2 1 - 2 2 Hungariella aff. kutassyi — x l ; the 2007.3.1. specimen m back (21) and spire view (22).