Herzogia 27 (2), 2014: 237–256 237

Interesting records of lichenized, lichenicolous and sapro-phytic fungi from northern Germany

Ulf Schiefelbein, Christian Dolnik, Uwe de Bruyn, Matthias Schultz, René Thiemann, Regine Stordeur, Pieter P. G. van den Boom, Birgit Litterski &

Harrie J. M. Sipman

Abstract: Schiefelbein, U., Dolnik, C., de Bruyn, U., Schultz, M., Thiemann, R., Stordeur, R., van den Boom, P. P. G., Litterski, B. & Sipman, H. J. M. 2014. Interesting records of lichenized, lichenicolous and sapro-phytic fungi from northern Germany. – Herzogia 27: 237–256.Details of 50 lichen species, 19 lichenicolous fungi and two saprophytic fungi from northern Germany are presented. Chrysothrix flavovirens, Opegrapha viridipruinosa and Sclerococcum tephromelarum are new to Germany. Arthonia digitatae, A. molendoi, Calicium parvum, Caloplaca limonia, Cornutispora intermedia, Endococcus rugulosus, Intralichen lichenicola, Lecidea nylanderi, Muellerella erratica, Pycnora sorophora and Verrucaria tristis are record-ed for the first time from northern Germany. 38 species are added to the lichen flora of the federal state Mecklenburg-Western Pomerania, 12 species to Schleswig-Holstein, five species to Lower Saxony and one species to Brandenburg.

Zusammenfassung: Schiefelbein, U., Dolnik, C., de Bruyn, U., Schultz, M., Thiemann, R., Stordeur, R., van den Boom, P. P. G., Litterski, B. & Sipman, H. J. M. 2014. Interessante Funde von lichenisierten, lichenicolen und saprophytischen Pilzen aus Norddeutschland. – Herzogia 27: 237–256.Informationen zu 50 Flechten, 19 flechtenbewohnenden und zwei saprophytischen Pilzen werden gegeben. Chrysothrix flavovirens, Opegrapha viridipruinosa und Sclerococcum tephromelarum sind neu für Deutschland. Arthonia digita-tae, A. molendoi, Calicium parvum, Caloplaca limonia, Cornutispora intermedia, Endococcus rugulosus, Intralichen lichenicola, Lecidea nylanderi, Muellerella erratica, Pycnora sorophora und Verrucaria tristis wurden das erste Mal in Norddeutschland nachgewiesen. 39 Arten können zur Flechtenflora von Mecklenburg-Vorpommern, 12 Arten zur Flechtenflora von Schleswig-Holstein, fünf Arten zur Flechtenflora von Niedersachsen und eine Art zur Flechtenflora von Brandenburg addiert werden.

Key words: Biodiversity, distribution, Brandenburg, Lower Saxony, Mecklenburg-Western Pomerania, Schleswig-Holstein.

IntroductionThis paper is a further contribution to the lichen flora of northern Germany and its federal states. During several field trips of the authors and the revision of herbarium material interest-ing lichenized, lichenicolous and non-lichenized fungi were found. In this paper, records of species new to northern Germany or one of its federal states, rediscoveries of species not ob-served for an extended period and rare species which were found for the last time before 2000 in northern Germany or one of its federal states are presented.

238 Herzogia 27 (2), 2014

Material and methodsThe nomenclature of lichens follows Wirth et al. (2013) and that of lichenicolous fungi Wirth et al. (2011). Cited specimens are kept in the private herbaria of the authors: P. P. G. van den Boom (Bo), U. de Bruyn (Br), C. Dolnik (Do), M. Schultz (Schu) and U. Schiefelbein (Schi) as well as in the herbaria of the Ernst-Moritz-Arndt-University Greifswald (GFW), Martin-Luther-University Halle (HAL), Christian-Albrechts-University Kiel (KIEL) and Übersee-Museum Bremen (BREM). Characteristic lichen substances of some specimens were analysed by thin-layer chromatography (TLC) in solvent system A, B’, C (Orange et al. 2001) by C. Dolnik, H. J. M. Sipman and U. Schiefelbein.

All presented localities are provided with the number of the German ordnance map (Messtischblatt) plus quadrant and geographic latitude and longitude coordinates.

The following signs are used: * lichenicolous fungus, + saprophytic fungus.

Species listAgonimia allobata (Stizenb.) P.James

Mecklenburg-Western Pomerania: 1448/1, Isle of Rügen, Jasmund National Park, 200 m inland from the coast, 13°40'07''E/54°33'36''N, alt. 96 m, old coastal beech forest, on Fagus sylvatica, 9 June 2013, leg. & det. R. Thiemann, conf. R. Stordeur & U. Schiefelbein (HAL 4887).

New to Mecklenburg-Western Pomerania. Agonimia allobata is a rare species in the northern German lowland (Hauck & de Bruyn 2010) as well as in the whole of Germany (Wirth et al. 2013). In northern Germany it is hitherto known only from the Bentheimer Wald, Lower Saxony (de Bruyn 2005), the Spreewald (Otte et al. 1997) and the Havelluch (Rätzel et al. 2005) in Brandenburg. The record from the Pobüller Bauernholz in Schleswig-Holstein (Dolnik 2005) belongs to the newly described A. flabel-liformis (Guzow-Krzemińska et al. 2012), a species very similar to A. allobata. According to Guzow-Krzemińska et al. (2012) the remaining A. allobata probably still represents two sibling species.

Agonimia tristicula (Nyl.) Zahlbr.Mecklenburg-Western Pomerania: 1447/2, Isle of Rügen, Jasmund National Park, Victoria-Sicht, 13°39'54''E/54°34'15''N, alt. 70 m, old coastal beech forest, on Fagus sylvatica, over moss, 2 September 2010, leg. & det. P. P. G. van den Boom (Bo 44972).

New to Mecklenburg-Western Pomerania. According to Wirth et al. (2013) the species might be fairly common in Germany, but in the northern part it is only known from the Harz Mountains, Lower Saxony (Hauck et al. 2009) and three localities in Brandenburg (Otte et al. 2006, Rätzel et al. 2005).

Anisomeridium biforme (Borrer) R.C.HarrisLower Saxony: 2409/2, Ostfriesisch-Oldenburgische Geest, Halbemond 5 km SE Norden, Abbingaster Tief, 07°17'00''E/53°34'07''N, alt. 1 m, on old Quercus robur along a creek, 01 April 2014, leg. & det. U. de Bruyn (Br 4950); Mecklenburg-Western Pomerania: 1447/1, Isle of Rügen, coast between Kluckow and Nardewitz, c. 2.8 km W of Nardewitz, 13°31'36''E/54°34'36''N, alt. 2 m, foot of the cliff, on Fraxinus excelsior, 25 October 2013, leg. & det. C. Dolnik, U. de Bruyn & U. Schiefelbein (Do 2772); 1935/1, peninsula Wustrow, parkland of the for-mer manor, 11°34'16''E/54°05'N, alt. 7 m, abandoned parkland, on Acer pseudoplatanus, 20 October 2012, leg. M. Schultz & U. Schiefelbein, det. U. Schiefelbein (Schi 3693).

Rediscovered in Lower Saxony after more then 85 years. Anisomeridium biforme is an extremely rare and strongly endangered species in Germany (Wirth et al. 2011, Wirth et al. 2013). In north-western Germany only Dolnik (2005) found this species recently. The most recent records in Mecklenburg-Western Pomerania were from the 1990s by B. Litterski from the Stubnitz on the isle of Rügen and by C. Tirkot from the Ahrenshooper Holz on the Vordarß. The last record in Lower Saxony dates back to 1928 from the Elbe-Weser-triangle (Hauck & de Bruyn 2010).

Schiefelbein et al.: Records of lichenized, lichenicolous and saprophytic fungi from northern Germany 239

*Arthonia apotheciorum (A.Massal.) Almq.Lower Saxony: 2912/3, Ostfriesisch-Oldenburgische Geest, Sedelsberg, 07°44'31''E/53°02'18''N, alt. 6 m, concrete wall at railway station, on Lecanora albescens, 19 January 2011, leg. A. Willenborg, det. U. de Bruyn (BREM); Mecklenburg-Western Pomerania: 1747/2, Isle of Rügen, peninsula Groß Zicker, cliff on the western side, 13°39'17''E/54°17'23''N, alt. 1 m, boulder beach, on Lecanora albescens over sandstone, 27 October 2013, leg. C. Dolnik, U. de Bruyn & U. Schiefelbein, det. U. Schiefelbein (Schi 3716); 1935/1, peninsula Wustrow, Baltic Sea coast, 11°32'19''E/54°04'52''N, alt. 2 m, ruins on the cliff, concrete, on Lecanora albescens and L. dispersa, 20 October 2012, leg. & det. C. Dolnik & U. de Bruyn (Do 2822); 2246/3, Kleines Landgrabental, Binsenberg, c. 2,3 km SSE of Siedenbollenthin, 13º23'86''E/53º42'40''N, alt. 16 m, concrete pillar at edge of a meadow, on Lecanora albescens, 02 November 2012, leg. & det. U. Schiefelbein (Schi 3383); Schleswig-Holstein: 1324/2, Anglia, Norderbrarup, 09°46'03.7''E/54°39'23''N, alt. 28 m, church wall, on concrete, on Lecanora antiqua, 09 July 2013, leg. & det. C. Dolnik (Do 2698); 1813/1, Helgoland, Düne, pond in the centre of the Düne, 07º54'56''E/54º11'04''N, alt. 2 m, on a concrete road, on Lecanora albescens, 24 September 2011, leg. C. Dolnik & U. Schiefelbein, det. U. Schiefelbein (Schi 3434); 1813/1, Helgoland, Mittelland, southern cliff “Am Kringel”, 07º53'14''E/54º10'39.6''N, alt. 12 m, on Lecanora albescens over sandstone, 23 September 2011, leg. C. Dolnik & U. Schiefelbein, det. C. Dolnik (KIEL, Do 2552).

New to Mecklenburg-Western Pomerania and Lower Saxony. According to Wirth et al. (2011) rare in Germany, but probably overlooked and rather common. In northern Germany the species is already known from Brandenburg (Otte & Rätzel 2004, as A. clemens) and Schleswig-Holstein (Dolnik et al. 2010).

*Arthonia digitatae HafellnerMecklenburg-Western Pomerania: 2645/1, Müritz National Park, 7 km E of Neustrelitz, W of Carpin, SE of Zinow, north side, marked path in forest, to Serrahn, near bird-spotting place, 13°11'47''E/53°21'26''N, alt. 70 m, mixed trees such as Betula, Pinus, Quercus, Fagus and mature Vaccinium shrubs, on Pinus, on Cladonia digitata, 21 October 2009, leg. & det. P. P. G. van den Boom (Bo 43587).

New to northern Germany. Arthonia digitatae is fairly common in Germany (Wirth et al. 2011), but for its northern range the validity of this statement still needs to be proved.

*Arthonia molendoi (Heufl. ex Frauenf.) R.Sant.Schleswig-Holstein: 1532/1, Isle of Fehmarn, Petersdorf, 11°04'06''E/54°28'48''N, alt. 3 m, west exposed church wall, on Caloplaca saxicola, 19 February 2013, leg. & det. C. Dolnik, conf. U. Schiefelbein (KIEL, Do 2692).

New to northern Germany. A rather rare lichenicolous fungus in Germany (Wirth et al. 2011). Records have been only published from Bavaria (e.g. Triebel & Scholz 2001, Brackel 2009, 2014), Hesse (Cezanne et al. 2013) and Rhineland-Palatinate (John et al. 2011).

Arthonia punctiformis Ach.Mecklenburg-Western Pomerania: 1447/1, Isle of Rügen, Uferkaten, c. 0,8 km WNW of Nardewitz, 13°33'26''E/54°34'46''N, alt. 40 m, old coastal beech forest, 25 October 2013, leg. & det. C. Dolnik, U. de Bruyn & U. Schiefelbein (Schi 3795); 1935/1, Mecklenburg bay, Wustrow peninsula, Baltic Sea coast, on the northern side of the peninsula, 11°32'09''E/54°04'27''N, on Sorbus aucuparia, 20 October 2012, leg. & det. C. Dolnik & U. de Bruyn (Br 4957).

Whereas A. punctiformis seems to be fairly common in the western part of the northern German low-land (Dolnik et al. 2010, Hauck & de Bruyn 2010) it is only rarely reported in the eastern part (Litterski 1999, Otte et al. 2006). The last (unpublished) record in Mecklenburg-Western Pomerania is from Central Mecklenburg by B. Litterski in 1997.

Bacidina caligans (Nyl.) Llop & HladunMecklenburg-Western Pomerania: 1447/3, Isle of Rügen, NW of Sagard, road 0,5 km E of Polchow, 13°31'36''E/54°32'50''N, alt. 35 m, roadside trees and shrubs, on Sambucus, 2 September 2010, leg. & det. P. P. G. van den Boom (Bo 44973); 1747/2, Isle of Rügen, peninsula Groß Zicker, cliff on the western site, 13°39'17''E/54°17'23''N, alt. 1 m, active coastal cliff, on Sambucus nigra, 27 October 2013, leg. & det. C. Dolnik, U. de Bruyn & U. Schiefelbein (Do 2754, Schi 3707); Schleswig-Holstein: 1223/4, Satrup, beech forest Kleine Holzkoppel, 09°36'04''E/54°42'51''N, alt. 50 m, over moss on log of beech, 01 August 2013, leg. & det. C. Dolnik (KIEL, Do 2665).

240 Herzogia 27 (2), 2014

New to Mecklenburg-Western Pomerania and Schleswig-Holstein. The distribution and frequency of B. caligans in Germany is insufficiently known, but Wirth et al. (2013) assume that it occurs quite commonly in western and north-western Germany.

Bacidia circumspecta (Nyl. ex Vain.) MalmeMecklenburg-Western Pomerania: 2645/3, Mecklenburgische Seenplatte, Müritz National Park, c. 1 km S of Serrahn, 13°12'07''E/53°20'13''N, alt. 112 m, old beech forest, on Fagus sylvatica, 18 August 2012, leg. & det. R. Thiemann, conf. H. J. M. Sipman, R. Stordeur & U. Schiefelbein (HAL 4888).

New to Mecklenburg-Western Pomerania and rediscovered in northern Germany after more than 70 years. Bacidia circumspecta is a rather rare and strongly endangered species of old-growth forests (Wirth et al. 2011), in northern Germany hitherto identified only by Zschacke (1909) from the Harz Mountains, Lower Saxony and by Erichsen (1939) from Oldenburg in Holstein, Schleswig-Holstein.

Bacidina etayana (P.Boom & Vězda) M.Hauck & V.WirthMecklenburg-Western Pomerania: 1935/1, peninsula Wustrow, northern part of the spit Kieler Ort, 11°31'14''E/54°05'36''N, open dunes, 20 October 2012, leg. & det. C. Dolnik & U. de Bruyn (Do 2825); Schleswig-Holstein: 1326/1, Oehe-Schleimünde, Baltic Sea coast, 10°01'30''E/54°41'41''N, alt. 2 m, plant debris of Ammophila, 10 December 2013, leg. & det. C. Dolnik (KIEL, Do 2808); 1813/2, Helgoland, dunes E of the airport, 07°55'01''E/54°11'09''N, alt. 2 m, debris of coastal dune grasses, 24 September 2011, leg. & det. C. Dolnik & U. Schiefelbein (KIEL, Do 2521).

New to Mecklenburg-Western Pomerania. Bacidina etayana probably is an overlooked species. Most records from Germany are from plant debris on coastal dunes and only few from bark. The first record for Germany has been published by Dolnik (2004) from Schleswig-Holstein.

Calicium parvum TibellMecklenburg-Western Pomerania: 2051/3, Isle of Usedom, dunes between Ahlbeck and the German-Polish border, 14º13'07''E/53º55'37''N, alt. 10 m, pine forest on dunes, on Pinus sylvestris, 24 June 2007, leg. & det. U. Schiefelbein (Schi 3516; TLC: diffractaic acid).

New to northern Germany. According to Wirth et al. (2011, 2013) Calicium parvum is an extremely rare and strongly endangered species, but as the result of the small size of the apothecia it is probably overlooked.

Caloplaca albolutescens (Nyl.) H.OlivierSchleswig-Holstein: 1813/1, Helgoland, Mittelland, southern part of the island, 07°53'11''E/54°10'51''N, alt. 12 m, sandstone boulders and artificial stones on the south-facing inactive cliff, on brick, 23 September 2011, leg. C. Dolnik & U. Schiefelbein, det. U. Arup (Do 2547 in KIEL, Schi 3409, additional to Flavoplaca limonia).

New to Schleswig-Holstein. The species is rare in Germany (Wirth et al. 2013); in the northern German Lowland it has been identified so far in Brandenburg (Rätzel et al. 2004) and Lower Saxony (Schiefelbein et al. 2010).

Caloplaca arcis (Poelt & Vězda) Arup [Flavoplaca arcis (Poelt & Vězda) Arup, Frödén & Søchting]Mecklenburg-Western Pomerania: 2033/4, Nordwestmecklenburg, Wismar bay, Hohenkirchen, churchyard, 11°18'17''E/53°55'09''N, alt. 33 m, church wall, on brick, 19 October 2012, leg. & det. C. Dolnik, U. de Bruyn, M. Schultz & U. Schiefelbein (Schi 3676).

New to Mecklenburg-Western Pomerania. This species has been neglected in Germany until the recent past. The distribution and frequency can, consequently, not be ascertained. However, it is already reported from several regions in north-western Germany (Dolnik & Neumann 2010, Wirth et al. 2013).

Caloplaca atroflava (Turner) Mong.Mecklenburg-Western Pomerania: 1447/1, Isle of Rügen, coast between Kluckow and Nardewitz, c. 2,8 km W of Nardewitz, 13°31'36''E/54°34'36''N, alt. 1 m, boulder beach, on granite, 25 October 2013, leg. & det. U. de Bruyn, C. Dolnik & U. Schiefelbein, conf. J. Vondrák (Schi 3820).

First verified record in Mecklenburg-Western Pomerania. In Schleswig-Holstein few localities from siliceous rocks of churchyards and megalith monuments have been recognized (Dolnik et al. 2012). A further record was published by Weber (2001) from a church wall in Central Mecklenburg.

Schiefelbein et al.: Records of lichenized, lichenicolous and saprophytic fungi from northern Germany 241

Caloplaca cerinella (Nyl.) Flagey [Athallia cerinella (Nyl.) Arup, Frödén & Søchting]Lower Saxony: 2912/3, Ostfriesisch-Oldenburgische Geest, Sedelsberg, 07°42'55''E/53°02'42''N, alt. 6 m, aban-doned farm, on branches in crown of felled Populus ×canadensis, 27 January 2010, leg. A. Willenborg, det. U. de Bruyn (Br 4761); Mecklenburg-Western Pomerania: 1747/2, Isle of Rügen, peninsula Groß Zicker, cliff on the western side, 13°39'17''E/54°17'23''N, alt. 1 m, coastal cliff, 27 October 2013, leg. & det. U. de Bruyn, C. Dolnik & U. Schiefelbein (Do 2767, Schi 3708).

New to Mecklenburg-Western Pomerania. Northern German records of this rare and endangered spe-cies (Wirth et al. 2013) have been presented by Hobohm et al. (2004), Otte & Rätzel (2008) and Thüs (2001). The morphologically and ecologically very similar C. cerinelloides, which differs only in the number of the spores per ascus, seems to be much more common, at least in Mecklenburg-Western Pomerania and Lower Saxony.At the here presented locality the species grows, extraordinarily, on Artemisia maritima on an active coastal cliff together with Lecania cyrtella and Lecanora sambuci.

Caloplaca cerinelloides (Erichsen) Poelt [Athallia cerinelloides (Erichsen) Arup, Frödén & Søchting]Lower Saxony: 2310/2, Ostfriesisch-Oldenburgische Geest, city of Aurich, hedge bank area W of urban zone, 07°26'11''/53°27'39''N, alt. 3 m, smooth bark of Populus tremula and Fagus sylvatica, leg. & det. U. de Bruyn, 07 March 2014 (Br 4954).

New to Lower Saxony. Caloplaca cerinelloides seems to be a spreading species on Populus tremula, Fraxinus excelsior and slightly eutrophicated bark of other tree species together with Lecania cyrtella, L. naegelii and Catillaria nigroclavata.

Caloplaca limonia Nimis & Poelt [Flavoplaca limonia (Nimis & Poelt) Arup, Frödén & Søchting]Mecklenburg-Western Pomerania: 1935/1, peninsula Wustrow, former manor, 11°34'16''E/54°05'00''N, alt. 7 m, north-facing wall of the former manor, on mortar, 20 October 2012, leg. & det. M. Schultz & U. Schiefelbein (Schi 3682, Schu 01693, 01694); Schleswig-Holstein: 1813/1, Helgoland, northern part of the island, cliff S of the sports field, 07º52'46''E/54º11'10''N, alt. 18 m, inactive sandstone cliff, on sandstone, 23 September 2011, leg. C. Dolnik & U. Schiefelbein, det. U. Arup (Schi 3401); 1813/1, Helgoland, Mittelland southern part of the island, 07°53'11''E/54°10'51''N, alt. 12 m, sandstone boulders and artificial stones on the south-facing inactive cliff, on brick, 23 September 2011, leg. C. Dolnik & U. Schiefelbein, det. U. Arup (Schi 3409).

New to northern Germany. A further sterile representative of the genus Caloplaca having been neglec-ted in Germany. Because of the preference for eutrophicated artificial walls fertilized by urine it is to be assumed that the species is more widespread than indicated by Wirth et al. (2013).

Caloplaca ruderum (Malbr.) J.R.LaundonMecklenburg-Western Pomerania: 1447/1, Isle of Rügen, Bobbin, churchyard, 13°31'38''E/54°33'13''N, alt. 31 m, church wall, on mortar, 26 October 2013, leg. & det. U. de Bruyn, C. Dolnik & U. Schiefelbein (Schi 3752, host of Intralichen lichenicola); 1546/4, Isle of Rügen, Bergen, churchyard, 13°25'57''E/54°25'01''N, alt. 70 m, church wall, 24 October 2013, vid. C. Dolnik & U. Schiefelbein.

New to Mecklenburg-Western Pomerania. Caloplaca ruderum is rather common in the western part of the northern German plain (Dolnik et al. 2010, Hauck & de Bruyn 2010), and seems to be hitherto overlooked in Mecklenburg-Western Pomerania.

Candelariella medians (Nyl.) A.L.Sm.Mecklenburg-Western Pomerania: 1546/4, Isle of Rügen, Bergen, churchyard, 13°25'57''E/54°25'01''N, alt. 70 m, old memorial slabs, 24 October 2013, vid. U. de Bruyn, C. Dolnik & U. Schiefelbein; Schleswig-Holstein: 1813/1, Helgoland, Mittelland, southern part of the island, 07°53'11''E/54°10'51''N, alt. 12 m, sandstone boulders on the south-facing inactive cliff, on sandstone, 23 September 2011, vid. C. Dolnik & U. Schiefelbein.

New to Schleswig-Holstein. Candelariella medians is a rare species in northern Germany, and oc-curs mainly in anthropogenic habitats (Hauck & de Bruyn 2010, Litterski 1999, Litterski & Schiefelbein 2007, Otte & Rätzel 2004).

Chrysothrix flavovirens TønsbergMecklenburg-Western Pomerania: 1739/2, Rostocker Heide, Ribnitzer Großes Moor, c. 0.7 km NE of Graal-Müritz, 12°16'41''E/54°16'05''N, old bog woodland, on Pinus sylvestris and wood, 04 September 2012, leg. & det.

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U. Schiefelbein (Schi 3365, 3375); 2351/4, Ueckermünder Heide, eastern edge of the nature reserve “Gorinsee”, 14°14'49''E/53°36'15''N, old pine forest, on Pinus sylvestris, 02 June 2012, leg. & det. U. Schiefelbein, conf. M. Kukwa (Schi 3302).

New to Germany. Chrysothrix flavovirens is a yellow-green, crustose species regarded by Tønsberg (1994) as the sorediate counterpart of C. chrysophthalma (P.James) P.James & J.R.Laundon. In contrast to all other very similar species [C. chrysophthalma, C. chamaecyparicola Lendemer, C. candelaris (L.) J.R.Laundon], it contains diffractaic acid (= chrysophtalma unknown, Elix & Tønsberg 2004). In Europe the species usually occurs on the bark of Pinus sylvestris, Betula pendula and Picea abies, but some collections are from Quercus sp. and wood (Aptroot et al. 2003, Kowalewska & Jando 2004, Rodrigues et al. 2011, Tønsberg 1994). In Mecklenburg-Western Pomerania it was recorded on the bark of Pinus sylvestris, on wood in an open old-growth pine forest and in old bog woodland. The species may be regarded as a preferentially coastal species (Laundon 1981, who treated them as sorediate thalli of C. chrysophthalma, Kowalewska & Jando 2004, Tønsberg 1994). The localities in the Rostocker Heide (c. 200 m away from the Baltic coast) and in the Ueckermünder Heide (c. 10 km inland) confirm this statement.Detailed information on the distribution of C. flavovirens is provided by Kowalewska & Jando (2004), Rodrigues et al. (2011) and Tønsberg (1994). In Europe it is widespread in coastal areas from the Mediterranean to the northern temperate zone with a preference for temperate regions. Its latitudinal range reaches from Portugal to Estonia. It is also known from North America (Richardson et al. 2009) and Asia (Kashiwadani & Thor 1997).

*Clypeococcum hypocenomycis D.Hawksw.Mecklenburg-Western Pomerania (only selected record): 1646/2, Isle of Rügen, Putbus, S of village, 13°28'18''E/54°21'06''N, alt. 45 m, former country-seat with many different mature scattered trees, on Sequoia, on Hypocenomyce scalaris, 3 September 2010, P. P. G. van den Boom (Bo 44983); Schleswig-Holstein: 1322/4, Schleswig, Bollingstedter Moor, c. 3,8 km NE of Bollingstedt, 09º27'07''E/54º37'02''N, alt. 18 m, fence post at the bog edge, on Hypocenomyce scalaris, 26 February 2013, leg. & det. C. Dolnik, conf. U. Schiefelbein (KIEL, Do 2519); 1526/4, Dänischer Wohld, forest Stodthagen, 10º04'24''E/54º25'10''N, alt. 20 m, fencepost, on Hypocenomyce scalaris, 27 February 2014, leg. & det. C. Dolnik (Do 2843).

New to Schleswig-Holstein. Even though the distribution and frequency is not sufficiently known it can be assumed that this lichenicolous fungus is fairly common in Germany at least in sandy, nutrient poor regions.

*Cornutispora intermedia Punith. & D.Hawksw.Mecklenburg-Western Pomerania: 1740/1, Rostocker Heide, eastern edge of forest complex, c. 1.1 km NE of Körkwitz Hof, 12º20'25''E/54º16'01''N, alt. 5 m, Prunus spinosa-shrubbery at forest edge, on Xanthoria parietina, 31 October 2012, leg. & det. U. Schiefelbein (Schi 3368).

New to northern Germany. From Europe only Brackel (2009) published this species until now. It was described by Punithalingam (2003) from Ochrolechia but Brackel (2009) also recognized it on a Xanthoria species.

+Cyrtidula quercus (A.Massal.) MinksMecklenburg-Western Pomerania: 1447/1, Isle of Rügen, Uferkaten, c. 0.8 km WNW of Nardewitz, 13°33'26''E/54°34'46''N, alt. 40 m, old coastal beech forest, 25 October 2013, vid. C. Dolnik, U. de Bruyn & U. Schiefelbein; 1747/2, Isle of Rügen, peninsula Groß Zicker, western side, Nonnenloch, 13°39'10''E/54°17'39''N, alt. 34 m, old oak trees at forest edge, on Quercus sp., 27 October 2013, leg. & det. C. Dolnik, U. de Bruyn & U. Schiefelbein (Schi 3724); 2250/3, Uecker-Randow region, Ueckermünder Heide, Ueckermünde, forest W of the Zoological Garden, 14º01'22''E/53º43'50''N, alt. 5 m, at the edge of small forest, on Quercus sp., leg. & det. U. Schiefelbein, 27 March 2006 (Schi 3502).

Cyrtidula quercus is rather common in the north-western part of Germany (Dolnik et al. 2010, Hauck & de Bruyn 2010), but seems to be rare in the north-eastern part (Litterski 1999, Rätzel et al. 2005). In Mecklenburg-Western Pomerania, U. de Bruyn provided the latter record in 1998 from the Mecklenburger Schweiz (Litterski 1999).

Schiefelbein et al.: Records of lichenized, lichenicolous and saprophytic fungi from northern Germany 243

*Dacampia cyrtellae BrackelMecklenburg-Western Pomerania: 2645/1, Müritz National Park, 7 km E of Neustrelitz, W of Carpin, SE of Zinow, north side, marked path near Serrahn, 13°12'02''E/53°20'44''N, alt. 90 m, open forest with mixed trees such as Betula, Pinus, Quercus, Fagus and Sambucus, on Sambucus, on Lecania cyrtella, 21 October 2009, leg. & det. P. P. G. van den Boom (Bo 43571).

New to Mecklenburg-Western Pomerania. In Germany, this recently described lichenicolous fungus is reported from Bavaria, Brandenburg and Hesse (Brackel 2010a, b). It is also known from Belgium and Luxemburg (Diederich et al. 2012).

*Dactylospora parasitica (Flörke) ArnoldMecklenburg-Western Pomerania: 1540/2, Darß peninsula, beech forest along the Tiefenstück-Weg, 12º29'29''E/54º27'07''N, alt. 10 m, acidophilous beech forest, on Fagus sylvatica, on Pertusaria hymenea, 7 September 2013, leg. R. Thiemann, R. Stordeur & U. Schiefelbein, det. U. Schiefelbein (Schi 3540).

The species is described by Flörke (1819) as Lecidea parasitica from Berlin. It is a rather rare liche-nicolous fungus in Germany (Wirth et al. 2011) known from Baden-Württemberg (Cezanne et al. 2008), Bavaria (Brackel 2014), Brandenburg (Otte & Rätzel 1996, Rätzel et al. 2005), Hesse (Cezanne et al. 2008, Teuber 2009), Mecklenburg-Western Pomerania (Litterski & de Bruyn 1998), North Rhine-Westphalia (Heibel 1999), Rhineland-Palatinate (John 1990, John et al. 2011) and Thuringia (Brackel 2014).

Diplotomma pharcidium (Ach.) M.ChoisyMecklenburg-Western Pomerania: 1540/2, Darß peninsula, beech forest along the Tiefenstück-Weg, 12º29'29''E/54º27'07''N, alt. 10 m, acidophilous beech forest, on Fagus sylvatica, 7 September 2013, leg. R. Stordeur, R. Thiemann & U. Schiefelbein, det. R. Thiemann, conf. H. J. M. Sipman (HAL 4889); Schleswig-Holstein: 1321/4, Jörl, Rupeler Moor, 09º15'56,5''E/54º36'23''N, alt. 14 m, on Populus tremula, 19 July 2014, leg. & det. C. Dolnik (KIEL, Do 2931).

Rediscovered in Mecklenburg-Western Pomerania after more than 70 years. An extremely rare species in Germany, which has seemingly disappeared in most regions (Wirth et al. 2013). Erichsen (1936) provided the only collections from Mecklenburg-Western Pomerania.

*Endococcus rugulosus coll.Schleswig-Holstein: 1624/1, Bünsdorf, St. Katharinen church, 09º44'33''E/54º22'09''N, alt. 10 m, north exposed church wall, on Lecidella scabra over granite, 28 April 2013, leg. & det. C. Dolnik, conf. U. Schiefelbein (KIEL, Do 2594).

New to northern Germany. The genus Endococcus is in critical need of revision, because several ta-xonomical concepts exist. Whereas Sérusiaux et al. (1999) used the name E. rugulosus for species growing on Verrucaria species, Triebel (1989) and Ihlen & Wedin (2008) assign all specimens with (dark) brown, thick and smooth-walled ascospores of 13 –16 × 6 –7.5 µm, growing on a wide range of host lichens including species of Amygdalaria, Aspicilia, Ionaspis, Lecanora, Placopsis, Rhizocarpon and Verrucaria to this species.

Enterographa crassa (DC.) FéeMecklenburg-Western Pomerania: 1447/1, Isle of Rügen, Rugeshus, c. 0.6 km NW of Nardewitz, 13°33'48''E/54°34'48''N, alt. 10 m, old coastal beech forest, 25 October 2013, vid. C. Dolnik, U. de Bruyn & U. Schiefelbein; 1447/1, Isle of Rügen, Uferkaten, c. 0.8 km WNW of Nardewitz, 13°33'26''E/54°34'46''N, alt. 40 m, old coastal beech forest, 25 October 2013, leg. & det. C. Dolnik, U. de Bruyn & U. Schiefelbein (Schi 3794); 1447/2 Isle of Rügen, Jasmund National Park, N of Stubbenkammer, 13°40'07''E/54°33'36''N, alt. 96 m, old coastal beech forest, on old Fagus sylvatica, 9 June 2013, leg. R. Thiemann, det. U. Schiefelbein (HAL 4890).

First verified record in Mecklenburg-Western Pomerania. Enterographa crassa is a temperate-atlantic species (Litterski 1999, Wirth et al. 2013) which in Germany occurs predominantly, but rarely in Lower Saxony and Schleswig-Holstein. In Mecklenburg-Western Pomerania it has been con-fused with E. hutchinsiae and its occurrence was not yet clarified (Litterski 1999, Litterski & Schiefelbein 2007). The localities on the Isle of Rügen represent the eastern border of the distribution area. Noteworthy is furthermore that E. crassa is associated with E. hutchinsiae (Leight.) A.Massal. at all here published localities.

244 Herzogia 27 (2), 2014

Fellhaneropsis vezdae (Coppins & P.James) Sérus. & CoppinsMecklenburg-Western Pomerania: 1346/1, Isle of Rügen, Varnkewitz, coast between forest “Schwarbe” and “Höllen-Liet”, NE of Varnkewitz, north of the military base, 13°23'12''E/54°40'48''N, old Sambucus nigra shrub on coastal cliff, 18 March 1990, leg. & det. B. Litterski, conf. U. Schiefelbein, (GFW-Li 5238).

New to Mecklenburg-Western Pomerania. Fellhaneropsis vezdae is a temperate subatlantic species (Wirth et al. 2013). Consequently, it occurs in northern Germany, although not very frequently, main-ly in the western and north-western part and lacks distribution in the south-eastern part (Otte & Rätzel 2004, Wirth et al. 2013).

Fuscidea pusilla TønsbergMecklenburg-Western Pomerania: 1647/3, Isle of Rügen, Greifswalder Bodden, coast E of Lauterbach, 13°31'38''E/54°20'25''N, alt. 1 m, oak on the foot of the coastal cliff, 24 October 2013, leg. C. Dolnik, U. de Bruyn & U. Schiefelbein, det. C. Dolnik (Schi 3813, TLC: divaricatic acid).

New to Mecklenburg-Western Pomerania. Fuscidea pusilla has been found several times in north-western Germany (Boch & Sparrius 2006, de Bruyn et al. 2008, Dolnik 2005).

Halecania viridescens Coppins & P.JamesMecklenburg-Western Pomerania: 2645/1, Müritz National Park, 7 km E of Neustrelitz, W of Carpin, SE of Zinow, north side, marked path near Serrahn, 13°12'02''E/53°20'44''N, alt. 90 m, open forest with mixed trees such as Betula, Pinus, Quercus, Fagus and Sambucus, on Sambucus, 21 October 2009, leg. & det. P. P. G. van den Boom (Bo 43573).

New to Mecklenburg-Western Pomerania. Halecania viridescens has a temperate-subatlantic distribu-tion area (Wirth et al. 2013) and is more common in its western than in its eastern part. In northern Germany it has been reported rarely (de Bruyn et al. 2008, Dolnik et al. 2010, Wirth et al. 2013) but is certainly overlooked.

*Intralichen christiansenii (D.Hawksw.) D.Hawksw. & M.S.ColeMecklenburg-Western Pomerania: 1447/1, Isle of Rügen, Bobbin, churchyard, 13°31'38''E/54°33'13''N, alt. 31 m, church wall, on Caloplaca ruderum over mortar, 26 October 2013, leg. & det. U. de Bruyn, C. Dolnik & U. Schiefelbein (Schi 3752); 2246/3, Kleines Landgrabental, Binsenberg, c. 2.3 km SSE of Siedenbollenthin, 13°23'86''E, concrete piles at the edge of a meadow, on Lecanora albescens, 02 November 2012, leg. & det. U. Schiefelbein, conf. A. Flakus (Schi 3382); Schleswig-Holstein: 1223/1, Schleswig, Husby, churchyard, 09°34'38''E/54°45'56''N, alt. 48 m, church wall, 22 January 2013, leg. & det. C. Dolnik, conf. U. Schiefelbein (KIEL, Do 2575); 1423/4, Moldenit, churchyard, 09°37'06''E/54°32'38''N, alt. 24 m, north-exposed church wall, on Caloplaca pusilla (Calogaya p.), 11 October 2013, leg. & det. C. Dolnik (Do 2645).

New to Schleswig-Holstein. According to Wirth et al. (2011) this lichenicolous fungus is moderately common in Germany. However, only Giersberg et al. (1992) and Hawksworth (1979) have hitherto published records from northern Germany.

*Intralichen lichenicola (M.S.Christ. & D.Hawksw.) D.Hawksw. & M.S.ColeSchleswig-Holstein: 1813/1, Helgoland, Mittelland southern part of the island, 07°53'11''E/54°10'51''N, alt. 12 m, sandstone boulders and artificial stones on the south-facing inactive cliff, on Candelariella aurella over sandstone, 23 September 2011, leg. & det. C. Dolnik, conf. U. Schiefelbein (KIEL, Do 2551); 1819/3, Dithmarschen, Büsum, wastewater treatment plant S of Büsum, 08°52'43''E/54°07'38''N, alt. 2 m, coastal protection wall, on Candelariella aurella, 22 September 2011, leg. & det. C. Dolnik & U. Schiefelbein (Do 2586).

New to northern Germany. A rather common (Wirth et al. 2011) but probably overlooked species in Germany.

Lecania rabenhorstii (Hepp) ArnoldMecklenburg-Western Pomerania: 2634/4, Ludwigslust, castle park, artificial grotto N of the castle, 11º29'19''E/53º19'48''N, alt. 40 m, ruins, on bog iron, 24 May 2008, leg. U. Schiefelbein, erroneously published as Lecania turicensis (Schiefelbein 2009), det. P. P. G. van den Boom (Schi 2209); Schleswig-Holstein: 1819/3, Dithmarschen, Büsum, wastewater treatment plant S of Büsum, 08°52'43''E/54°07'38''N, alt. 2 m, coastal protection wall, 22 September 2011, leg. C. Dolnik & U. Schiefelbein, det. P. P. G. van den Boom (Schi 3512).

Schiefelbein et al.: Records of lichenized, lichenicolous and saprophytic fungi from northern Germany 245

New to Mecklenburg-Western Pomerania. A rare species in Germany (Wirth et al. 2013) and more common only in the lowland of Lower Saxony (Hauck & de Bruyn 2010). Lecania rabenhorstii has never been reported from north-eastern Germany before.

Lecanora hybocarpa (Tuck.) BrodoMecklenburg-Western Pomerania: 1540/2, Darß peninsula, dunes near the Tiefenstück-Weg, 12º29'08''E/54º27'04''N, alt. 5 m, old beeches on dunes, on Fagus sylvatica, 7 September 2013, leg. R. Thiemann, R. Stordeur & U. Schiefelbein, det. H. J. M. Sipman & U. Schiefelbein (Schi 3552, 3557; TLC: atranorin, roccellic acid).

New to Mecklenburg-Western Pomerania. Member of the L. subfusca-group, similar to L. chlarotera, which differs in the epipsamma and chemistry. Lecanora chlarotera has a layer of crystals that dissolve in K and N lying above the hymenium and contains atranorin, gangaleoidin with or without californin (Edwards et al. 2009). In Germany, it has been recognized only in the Prignitz, Brandenburg (Rätzel et al. 2002) and in the Eifel, Rhineland-Palatinate (Aptroot 2005).

Lecanora intumescens (Rebent.) Rabenh.Mecklenburg-Western Pomerania: 1749/3, Rostocker Heide, Wiedort, c. 1,7 km W of Torfbrücke, 12°11'54''E/54°14'33''N, alt. 4 m, mixed deciduous coastal forest, on Quercus sp., 30 December 2012, leg. & det. U. Schiefelbein, conf. H. J. M. Sipman (Schi 3656; TLC: atranorin, psoromic acid).

The species is still fairly common in southern Germany (Wirth et al. 2011), but disappeared from many regions of the northern German Lowland (Dolnik et al. 2010, Hauck & de Bruyn 2010, Litterski & Schiefelbein 2007, Otte & Rätzel 2004). The last record in Mecklenburg-Western Pomerania provided B. Litterski 1998 from the Stubnitz on the Isle of Rügen.

Lecanora sambuci (Pers.) Nyl.Mecklenburg-Western Pomerania: 1747/2, Isle of Rügen, peninsula Groß Zicker, cliff on the western side, 13°39'17''E/54°17'23''N, alt. 1 m, active coastal cliff, on Artemisia maritima, 27 October 2013, leg. U. de Bruyn, C. Dolnik & U. Schiefelbein, det. C. Dolnik (Do 2766).

Similar to L. persimilis and L. hagenii, the species is actually quite common in the southern and rare or absent in the northern part of Germany (Dolnik et al. 2010, Hauck & de Bruyn 2010, Litterski & Schiefelbein 2007, Otte & Rätzel 2004, Wirth et al. 2013). Litterski (1993) found the species in Mecklenburg-Western Pomerania for the last time. The species is characterized by more than 8 spores per ascus. Records with only 8 spores per ascus refer to L. persimilis.

Lecanora sarcopidoides (A.Massal.) A.L.Sm.Mecklenburg-Western Pomerania: 2451/2, Uckermärkisches Hügelland, forest complex Gottesheide, bog at the northern edge of the Gottesheide, 14°18'15''E/53°34'01''N, alt. 20 m, mesotrophic acidic open bog, on wood, 16 May 2012, leg. & det. U. Schiefelbein (Schi 3272; TLC: pseudoplacodiolic, placodiolic and usnic acids).

New to Mecklenburg-Western Pomerania and rediscovered in northern Germany after more than 50 years. Lecanora sarcopidioides is a member of the L. saligna group and differs from L. saligna in the absence of the distinctive broadly falcate macroconidia and the different chemistry. Whereas L. saligna has isousnic and neousnic acids, atranorin and zeorin, L. sarcopidioides contains pseudoplacodiolic, placodiolic and usnic acids (Edwards et al. 2009). In northern Germany it was collected rarely by Sandstede (1912) and Erichsen (1957) (see also van den Boom & Brand 2008).

Lecanora stenotropa Nyl.Mecklenburg-Western Pomerania: 2650/4, Randow valley, Radewitzer Heide, c. 0,8 km N Radewitz, 14º09'14''E/53º18'09''N, alt. 40 m, xerothermic grasslands, on sandstone, 31 August 2013, leg. & det. U. Schiefelbein (Schi 3602).

New to Mecklenburg-Western Pomerania. The species usually has not been distinguished from the very similar L. polytropa. Lecanora stenotropa is characterized by the brown-green to green-grey thal-lus consisting of dispersed to somewhat coalescing rounded granules, the pale brown to grey-brown apothecial discs and the much smaller ascospores. In addition, the thalline exciple is entire to flexuose, white-grey, scarcely raised and becoming almost excluded (Edwards et al. 2009).

246 Herzogia 27 (2), 2014

In northern Germany it is hitherto reported from Schleswig-Holstein (Dolnik & Neumann 2014), where it seems to be widely distributed but not as common as the similiar Lecanora polytropa.

Lecidea nylanderi (Anzi) Th.Fr.Mecklenburg-Western Pomerania: 2051/3, Isle of Usedom, nature reserve Zerninsee-Senke, Swinemoor, c. 2 km N of Kamminke, 14º12'31''E/53º40'01''N, alt. 1 m, drained raised bog, bog woodland, on wood, 10 February 2011, leg. & det. U. Schiefelbein (Schi 3465; TLC: divaricatic acid).

New to northern Germany. Lecidea nylanderi is characterized by the distinct, bluish black prothallus, the bluish grey soredia and divaricatic acid in the medulla (Aptroot et al. 2009), and is mostly to be found in sterile condition. The species occurs in coniferous forests of the boreal zone and in high-mountain areas of the temperate zone (Wirth et al. 2013) and also in old pine and alder forests of the Central European lowland (Schiefelbein et al. 2012).

Lepraria umbricola TønsbergBrandenburg: Märkische Schweiz, trail along Sophienfließ N of Buckow, 14°03'51''E/52°35'12''N, alt. 80 m, slope along little stream in forest valley, on Acer platanoides trunk, 30 April 2006, leg. H. J. M. Sipman & P. P. G. van den Boom, det. H. J. M. Sipman (Bo 54599, B 60 0130783; TLC: atranorin, tr. zeorin, thamnolic acid); Mecklenburg-Western Pomerania: 1540/2, Darß peninsula, beech forest along the Tiefenstück-Weg, 12º29'29''E/54º27'07''N, alt. 10 m, acidophilous beech forest, on Fagus sylvatica, 7 September 2013, leg. R. Stordeur, R. Thiemann & U. Schiefelbein, det. H. J. M. Sipman & U. Schiefelbein, conf. M. Kukwa (Schi 3533; TLC: thamnolic acid).

New to Brandenburg and Mecklenburg-Western Pomerania. This sterile lichen is only rarely reported from very few regions in south- and north-western Germany, but most likely overlooked (Wirth et al. 2013).

+Leptorhaphis atomaria (Ach.) SzatalaMecklenburg-Western Pomerania: 1747/2, Isle of Rügen, peninsula Groß Zicker, western side, Nonnenloch, 13°39'10''E/54°17'39''N, alt. 34 m, patch of poplar trees, on Populus tremula, 27 October 2013, C. Dolnik, U. de Bruyn & U. Schiefelbein (Schi 3725); 2034/1, Wismar bay, Eggers Wiek, Hohen Wieschendorf Huk, N of the ma-rina, 11°20'37'E/53°56'59''N, alt. 2 m, shrubberies and trees along the coast, on Populus tremula, 19 October 2012, leg. & det. C. Dolnik, U. de Bruyn, M. Schultz & U. Schiefelbein (Br 4956).

Leptorhaphis atomaria seems to be a rare species but is probably overlooked in Germany. It is known from only few regions, mainly in north-western Germany (Wirth et al. 2013). In the coastal region of Lower Saxony L. atomaria occurs quite frequently on smooth bark of young Populus tremula. In contrast, in Mecklenburg-Western Pomerania hitherto it has been found only once in the Nossenthiner Heide (de Bruyn et al. 1999).

Megalaria laureri (Hepp ex Th.Fr.) HafellnerMecklenburg-Western Pomerania: 1447/4, Isle of Rügen, Jasmund National Park, Waldhalle, 13º40'38''E/54º32'12''N, alt. 65 m, old coastal beech forest, on Fagus sylvatica, 2 September 2010, leg. & det. P. P. G. van den Boom, (Bo 44966, Schi 3514); 1447/2 Isle of Rügen, Jasmund National Park, N of Stubbenkammer, 13°40'07''E/54°33'36''N, alt. 96 m, old coastal beech forest, on old Fagus sylvatica, 9 June 2013, leg. R. Thiemann, det. H. J. M. Sipman (HAL 4893).

Rediscovered in northern Germany after more than 50 years and in Mecklenburg-Western Pomerania after more than 100 years. Megalaria laureri is an extremely rare species in Germany and extinct in many regions (Wirth et al. 2013). It is presently reported only from the Kellerwald in Hesse (Wirth et al. 2013) and from the Eifel in Rhineland-Palatinate (John 1990). The last records from northern Germany were published by Erichsen (1957). In Mecklenburg-Western Pomerania Sandstede (1903) found the species in the Stubnitz on the Isle of Rügen for the last time.

Micarea viridileprosa Coppins & P.BoomMecklenburg-Western Pomerania: 2351/1, Ueckermünder Heide, Ahlbecker Teerofen, former peat-ditches c. 0.5 km S of Ahlbeck, 14°10'35''E/53°39'47''N, alt. 20 m, mesotrophic acidic wooded bog, on wood, 30 April 2012, leg. & det. U. Schiefelbein, conf. P. Czarnota (Schi 3207); 2645/1, Müritz National Park, 7 km E of Neustrelitz, W of Carpin, SE of Zinow, north side, marked path in forest, to Serrahn, near bird-spotting place, 13°11'47''E/53°21'26''N,

Schiefelbein et al.: Records of lichenized, lichenicolous and saprophytic fungi from northern Germany 247

alt. 70 m, mixed trees such as Betula, Pinus, Quercus, Fagus and mature Vaccinium shrubs, on Betula, 21 October 2009, leg. & det. P. P. G. van den Boom (Bo 43585).

New to Mecklenburg-Western Pomerania. Micarea viridileprosa is most likely an overlooked lichen in Mecklenburg-Western Pomerania because it is known from many surrounding regions (Dolnik et al. 2010, Hauck & de Bruyn 2010, Otte & Rätzel 2004).

*Muellerella erratica (A.Massal.) Hafellner & Volk.JohnMecklenburg-Western Pomerania: 1447/1, Isle of Rügen, coast between Kluckow and Nardewitz, c. 2.8 km W of Nardewitz, 13°31'36''E/54°34'36''N, alt. 1 m, boulder beach, on Tephromela atra over granite, 25 October 2013, leg. C. Dolnik, U. de Bruyn & U. Schiefelbein, det. U. Schiefelbein (Schi 3776); Schleswig-Holstein: 1223/1, Anglia, Rüllschau, churchyard, 09°32'00''E/54°46'58''N, alt. 57 m, churchyard wall, on Caloplaca flavescens (Variospora f.), 16 May 2013, leg. & det. C. Dolnik, conf. U. Schiefelbein (Do 2607).

New to Mecklenburg-Western Pomerania, but known from Schleswig-Holstein (Triebel 1989). According to Wirth et al. (2011) M. pygmaea, a taxon previously considered as a variety, is moderate-ly common in Germany. However, since the varieties of M. pygmaea often have not been distinguished in the past the distribution is unsatisfactorily known.

*Muellerella hospitans Stizenb.Schleswig-Holstein: 1533/4, Isle of Fehmarn, Katharinenhof, Baltic Sea coast near the camping site, 11°16'45''E/54°26'41''N, alt. 13 m, coastal woodland, on Acer campestre, on Bacidia rubella, 26 April 2011, leg. C. Dolnik, det. U. Schiefelbein (KIEL, Do 2919).

New to Schleswig-Holstein, rediscovered in Germany after more than 55 years. The only known findings in Germany are from Baden-Württemberg by Lettau (1958) and Mecklenburg-Western Pomerania by H. G. Flörke (Schiefelbein et al. 2014).

*Muellerella lichenicola (Sommerf.) D.Hawksw.Mecklenburg-Western Pomerania: 1747/2, Isle of Rügen, nature reserve “Zicker”, western bank of the penin-sula Groß Zicker, 13º39'07''E/54º18'03''N, alt. 1 m, boulder beach, on sandstone, 22 September 2009, leg. & det. U. Schiefelbein (Schi 2781); Schleswig-Holstein: 1819/3, Dithmarschen, Büsum, wastewater treatment plant S of Büsum, 08°52'43''E/54°07'38''N, alt. 2 m, coastal protection wall, 22 September 2011, leg. C. Dolnik & U. Schiefelbein, det. U. Schiefelbein (Do 2586).

This Muellerella species seems to be rarer in northern Germany than M. erratica. P. Jacobsen found it already 1985 in Glücksburg, Schleswig-Holstein (Triebel 1989) and Weber (2001) in Central Mecklenburg.

*Muellerella pygmaea (Körb.) D.Hawksw.Mecklenburg-Western Pomerania: 2248/3, Schwerinsburg, former castle, 13º40'59''E/53º44'02''N, alt. 12 m, castle wall, on Lecidea fuscoatra, 12 October 1996, leg. & det. U. Schiefelbein, conf. A. Flakus (Schi 3460).

First verified record from Mecklenburg-Western Pomerania. The record of Bachmann (1913) on Lecanora sambuci from Drewoldke on the Isle of Rügen is probably another species as M. pygmaea occurs almost exclusively on epilithic lichens. Thiel & Spribille (2007) present the only reliable record from northern Germany from sandstone outcrops in the vicinity of Göttingen.

*Nectriopsis micareae Diederich, P.Boom & G.ErnstMecklenburg-Western Pomerania: 2645/1, Müritz National Park, 7 km E of Neustrelitz, W of Carpin, SE of Zinow, north side, marked path in forest, to Serrahn, near bird-spotting place, 13°11'47''E/53°21'26''N, alt. 70 m, mixed trees such as Betula, Pinus, Quercus, Fagus and mature Vaccinium shrubs, on Betula, on Micarea viridile-prosa, 21 October 2009, leg. & det. P. P. G. van den Boom (Bo 43584).

New to Mecklenburg-Western Pomerania. Nectriopsis micareae already has been found in Bavaria (Brackel 2009, 2014), Lower Saxony (Sérusiaux et al. 1999) and Hesse (Cezanne et al. 2002).

Ochrolechia subviridis (Høeg) ErichsenMecklenburg-Western Pomerania: 1936/1, Biendorf, churchyard, 11°41'55''E/54°04'29''N, alt. 48 m, trees in the churchyard, on Tilia sp., 20 October 2012, leg. C. Dolnik, U. de Bruyn, M. Schultz & U. Schiefelbein, det. H. J. M. Sipman & U. Schiefelbein (Schi 3621, 3622; TLC: gyrophoric acid); Schleswig-Holstein: 1122/3,

248 Herzogia 27 (2), 2014

Niehuus (Harrislee), village centre, 09°22'53''E/54°49'36''N, alt. 24 m, on bark of old oak tree, 01 March 2013, leg. & det. C. Dolnik (KIEL, Do 2860; TLC: gyrophoric acid); 1324/2, Anglia, Norderbrarup, churchyard, 09°46'05''E/54°39'23.7''N, alt. 29 m, on Tilia sp., 9 July 2013, leg. & det. C. Dolnik (KIEL, Do 2850; TLC: gyro-phoric acid); 1422/3, Silberstedt, forest Rumbrand, 09°21'04''E/54°31'11''N, alt. 18 m, bark of old oak, 31 August 2013, leg. & det. C. Dolnik (KIEL, Do 2706; TLC: gyrophoric acid).

Ochrolechia subviridis is widespread but not very common in atlantic and subatlantic regions (Kukwa 2011, Litterski 1999, Wirth et al. 2013). In Mecklenburg-Western Pomerania it had been found by U. de Bruyn in the Granitz on the Isle of Rügen ultimately (Litterski 1999).

Opegrapha gyrocarpa Flot.Mecklenburg-Western Pomerania: 1647/4, Isle of Rügen, Lanken-Granitz, churchyard, 13°37'56''E/54°22'03''N, church wall, 27 October 2013, leg. & det. C. Dolnik, U. de Bruyn & U. Schiefelbein (Schi 3755; TLC: gyrophoric acid); 1945/3, Sassen-Trantow, Sassen, churchyard, 13 October 2013, 13°11'15''E/54°01'22''N, alt. 37 m, on brick of the north-facing church wall, leg. & det. C. Dolnik, (Do 2894; TLC: gyrophoric acid); 2033/4, Hohenkirchen, churchyard, 11°18'58''E/53°55'09''N, alt. 33 m, on brick of the north-facing church wall, 19 October 2012, leg. C. Dolnik, U. de Bruyn & U. Schiefelbein, det. H. J. M. Sipman & U. Schiefelbein (Do 2896; TLC: gyropho-ric acid); Schleswig-Holstein: 1225/1, Anglia, Gelting, churchyard, 09°53'53''E/54°44'56''N, alt. 3 m, on brick of north-facing church wall, 03 September 2013, vid. C. Dolnik; 1324/2, Anglia, Norderbrarup, churchyard, 09°46'03,7''E/54°39'23''N, alt. 28 m, on brick and granite rocks of the north-facing church wall, 09 July 2013, vid. C. Dolnik; 1325/4, Schwansen, Karby, churchyard, 09°58'33''E/54°37'23''N, alt. 28 m, on brick of the north-facing church wall, 30 April 2013, vid. C. Dolnik; 1423/4, Kahleby, churchyard, 09°39'26.4''E/54°32'47.5''N, alt. 10 m, on brick of the north-facing church wall, 03 January 2013, leg. & det. C. Dolnik, (KIEL, Do 2648; TLC: gyro-phoric acid); 1423/4, Moldenit, 09°36'58''E/54°32'37.6''N, alt. 23 m, on brick of the north-facing church wall, 11 June 2013, vid. C. Dolnik; 1424, Kosel, hunting area stone mark (granite) from king Christian VII. (1766 –1808), 09°46'47.6''E/54°30'51''N, alt. 16 m, 27 March 2014, vid. C. Dolnik.

New to Mecklenburg-Western Pomerania. The core distribution area of O. gyrocarpa is in moun-tains of oceanic regions. Therefore, the species is much more present in southern and south-western Germany than in northern Germany. In the lowland it inhabits mainly old churches or churchyard walls (Hauck & de Bruyn 2010), but has probably been overlooked.

Opegrapha lithyrga Ach.Lower Saxony: 3210/2, Ems-Hunte-Geest, megalithic monument “Deymanns Mühle” 2 km W Klein-Berßen, 07°25'47.43''E/52°46'23.91''N, alt. 19 m, shaded underside of top stone, 19 September 2012, leg. A. Willenborg, det. U. de Bruyn (BREM).

New to the northern German lowland. Rather rare in the montane and high montane belt of southern Germany (Wirth et al. 2013), but extinct in upland areas of Lower Saxony (Hauck & de Bruyn 2010). Megalithic monuments are the only habitats for the species in northern Germany.

Opegrapha viridipruinosa B.J.Coppins & R.Yahr Lower Saxony (only selected records): 2510/2, Ostfriesisch-Oldenburgische Geest, Georgsheil 10 km W Aurich, trees in village, 07°19'18''E/53°28'28''N, alt. 0 m, bark of Aesculus, Fraxinus, Salix, 07 März 2014, leg. & det. U. de Bruyn (Br 4955); 2510/2, Ostfriesisch-Oldenburgische Geest, city of Aurich, hedge bank area W Extum 07°26'11''E/53°27'39''N, bark of oak in hedge bank, 20 Februar 2012, leg. & det. U. de Bruyn (Br 4952); 2511/4, Ostfriesisch-Oldenburgische Geest, city of Aurich, 07°30'48''E/53°28'06''N, alt. 1 m, abandoned landscape garden next to industrial area, shaded Carpinus betulus and Acer pseudoplatanus, 15 März 2014, leg. & det. U. de Bruyn (Br 4953).

New to Germany. Opegrapha viridipruinosa was described in 2011 from Great Britain (Lumbsch et al. 2011). It is also known from the Netherlands (Timmermann & Aptroot 2012). Opegrapha viridi-pruinosa is closely related to Alyxoria varia. It is distinguished by a conspicuous bright yellow thallus and pruina, smaller ascospores and smaller apothecia as demonstrated in Timmermann & Aptroot (2012). It seems to be a rapidly spreading species together with other lichen species with Trentepohlia photobionts (Aptroot & van Herk 2007, van den Broeck 2010, Hauck et al. 2013). Records are mainly from younger Acer pseudoplatanus, Fraxinus excelsior or Populus tremula covering larger are-as in most cases associated with Opegrapha rufescens. Opegrapha viridipruinosa also occurs on rough bark of Populus canadensis, Salix alba-hybrids and slightly eutrophicated bark of Quercus robur. It is

Schiefelbein et al.: Records of lichenized, lichenicolous and saprophytic fungi from northern Germany 249

very unlikely that this species has been overlooked in the past due to its conspicuous yellow thallus. The occurrence on younger trees also indicates its ability to spread.

Oxneria ulophyllodes (Räsänen) S.Y.Kondr. & KärnefeltLower Saxony: 2311/4, Ostfriesisch-Oldenburgische Geest, city of Esens, 07°37'E/53°38'51''N, alt. 7 m, roadside trees near main churchyard, 17 March 1997, leg. T. Homm & U. de Bruyn, det. M. Hauck & U. de Bruyn, October 2012 (Br).

New to the lowland area of Lower Saxony, extinct in upland areas of Lower Saxony (Hauck & de Bruyn 2010). A very rare species in northern Germany with no tendency to spread (Dolnik et al. 2010, Litterski & Schiefelbein 2007).

Parmotrema perlatum (Huds.) M.ChoisyMecklenburg-Western Pomerania: 1837/2, Nienhagen, Nienhäger Holz, 11°56'18''E/54°09'45''N, alt. 15 m, me-sophytic beech forest, on Fagus sylvatica, 21 October 2012, leg. & det. C. Dolnik, U. de Bruyn, M. Schultz & U. Schiefelbein (Schi 3642).

Rediscovered in Mecklenburg-Western Pomerania after more than 140 years. Parmotrema perlatum belongs to the currently recovering species (e.g. Dolnik et al. 2010, Hauck & de Bruyn 2010, Wirth et al. 2013). A. Müller provided the only record so far from Mecklenburg-Western Pomerania near Güstrow (Brockmüller 1863).

Peltigera extenuata (Nyl.) Vain.Lower Saxony: 3012/4, Ems-Hunte-Geest, NSG Markatal, c. 2 km SW Markhausen, 07°49'26.65''E/52°56'01.93''N, alt. 20 m, on sandy ground along path, 9 October 2010, leg. A. Willenborg, det. U. de Bruyn (BREM).

New to Lower Saxony. In the past rarely distinguished from P. didactyla, but in northern Germany alrea-dy known from Brandenburg (Rätzel et al. 2004), Mecklenburg-Western Pomerania (Schiefelbein 2013) and Schleswig-Holstein (Neumann & Dolnik 2012).

Phaeographis inusta (Ach.) Müll.Arg.Mecklenburg-Western Pomerania: 1447/2, Isle of Rügen, Jasmund National Park, close to Brisnitzer Bach, 13°39'52''E/54°33'16''N, alt. 96 m, old coastal beech forest, on Fagus sylvatica, 7 June 2013, leg. R. Thiemann, det. H. J. M. Sipman (HAL 4891).

New to Mecklenburg-Western Pomerania. An overview on the distribution of Phaeographis species in Germany is given by Ernst & Hauck (1994). Phaeographis inusta occurs only at very few localities in north-western Lower Saxony (de Bruyn 2000, de Bruyn et al. 2008, Homm & de Bruyn 2000) and in North Rhine-Westphalia in the Wolbecker Tiergarten near Münster (Ernst & Hauck 1994). The record on the Isle of Rügen represents in the temperate floristic zone the easternmost edge of this Atlantic species.

Physcia tribacioides Nyl.Lower Saxony: 2912/3, Ostfriesisch-Oldenburgische Geest, Küstenkanal 5 km NE Esterwegen 07°41'57.37''E/53°01'01.21''N, alt. 8 m, free standing tree in uncultivated land, one well developed thallus on hori-zontal branch of Salix spec., with Flavoparmelia soredians and Parmotrema perlatum, 2010, leg. A. Willenborg & U. de Bruyn., det. U. de Bruyn (Br 4951).

New to Lower Saxony. Physcia tribacioides seems to spread in Belgium and the Netherlands (Aptroot et al. 2011, Frahm et al. 2010, van den Broek et al. 2008), and was recorded also in North Rhine-Westfalia several times in the last decade (Frahm et al. 2010).

Punctelia jeckeri (Roum.) KalbMecklenburg-Western Pomerania: 2034/1, Wismar bay, Hohen Wieschendorf Huk, coast from the marina to the cape, 11°20'25''E/53°57'16''N, alt. 2 m, old trees along the coast, on Quercus sp., 20 October 2012, leg. & det. C. Dolnik, U. de Bruyn, M. Schultz & U. Schiefelbein (Schi 3662).

Punctelia jeckeri has not been distinguished from P. subrudecta in many regions of Germany as well as in Mecklenburg-Western Pomerania for a long time. Thus, the record of H. J. M. Sipman from Groß Belitz in Central Mecklenburg near Schwaan has also been published by Litterski (1999) under P.

250 Herzogia 27 (2), 2014

subrudecta. Hauck & de Bryun (2010) and Dolnik et al. (2010) recognized that the species was spreading in north-western Germany in the recent past.

Pycnora sorophora (Vain.) HafellnerMecklenburg-Western Pomerania: 2451/2, Uckermärkisches Hügelland, pine stand SW of Vorwerk Lenzen, along the overland route to Pampow, 14°16'39''E/53°33'38''N, on Pinus sylvestris, 16 May 2012, leg. & det. U. Schiefelbein, conf. M. Kukwa (Schi 3233; TLC: alectorialic acid).

New to northern Germany. The distribution of P. sorophora is similar to that of Lecidea nylanderi. It is widely distributed in the boreal floristic zone and in mountain regions of the temperate zone and extremely rare in the Central European lowland (Fałtynowicz 2003, Santesson et al. 2004, Wirth et al. 2013).

Ramalina baltica LettauMecklenburg-Western Pomerania: 2343/1, Oberes Peenegebiet, Jürgensdorf S of Stavenhagen, Rottmanshagen, western edge of the village, road side trees to a small chapel, 12º51'06''E/53º38'54''N, alt. 40 m, old lime trees, on areas sheltered from rain, 12 October 2007, leg. U. de Bruyn, det. H. J. M. Sipman & U. Schiefelbein (Schi 3492, TLC: evernic acid).

New to Mecklenburg-Western Pomerania. An extremely rare species in Germany collected recently only in Anglia (Schleswig-Holstein), Hesse highlands and Odenwald-Spessart (Wirth et al. 2013).

Rinodina teichophila (Nyl.) ArnoldMecklenburg-Western Pomerania: 1447/1, Isle of Rügen, Bobbin, churchyard, 13°31'38''E/54°33'13''N, alt. 31 m, church wall, on mortar, 26 October 2013, leg. C. Dolnik, U. de Bruyn & U. Schiefelbein, det. U. Schiefelbein (Schi 3754).

The very rare R. teichophila occurs in northern Germany exclusively on churches and churchyard walls (e.g. de Bruyn 2007b, de Bruyn et al. 2005, Weber 2001). Weber (2001) presented the only other record from Mecklenburg-Western Pomerania.

*Sclerococcum tephromelarum Etayo & Calat.Mecklenburg-Western Pomerania: 1346/1, Isle of Rügen, coast between forest “Schwarbe” and “Höllen-Liet”, c. 1 km NE of Varnkewitz, 13º22'45''E/54º41'01''N, boulder beach, alt. c. 3 m, on Tephromela atra, 23 September 2009, leg. U. Schiefelbein, det. M. Kukwa (Schi 3461).

New to Germany. Sclerococcum tephromelarum was described by Etayo & Calatayud (2003) from the Valencia Province in Spain. In addition to the type locality and two other localities in Spain it is known from the Netherlands (van den Boom 2003), France (Hafellner 2007, Roux 2012) and Madeira (Hafellner 2002, 2007).

Strigula jamesii (Swinscow) R.C.HarrisMecklenburg-Western Pomerania: 1447/2, 1448/1, Isle of Rügen, Jasmund National Park, 13°39'43''E/54°33'10''N, 13°40'07''E/54°33'36''N, alt. 96 m, old coastal beech forest, loose stock of trees, on Fagus sylvatica, 6 June 2013 and 9 June 2013, leg. & det. R. Thiemann, conf. H. J. M. Sipman, R. Stordeur & U. Schiefelbein (HAL 4892).

New to Mecklenburg-Western Pomerania. Strigula jamesii is an extremely rare species of the tem-perate and submediterranean zone with a preference for subatlantic regions (Roux & Sérusiaux 2004, Wirth et al. 2013). In northern Germany it was collected already in the Solling, Lower Saxony (Gerken et al. 2008) and in the Uckermark, Brandenburg (Otte et al. 2001, Roux & Sérusiaux 2004).

Thelidium minutulum Körb.Mecklenburg-Western Pomerania: 1836/3, Bad Doberan, Baltic Sea coast W of Heilgendamm, 11°49'52''E/54°08'41''N, alt. 2 m, active coastal cliff, loamy soil, 21 October 2012, leg. & det. C. Dolnik (Do 2815); 1935/3, Salzhaff, Boiensdorfer Werder, camping site, 11°31'07''E/54°01'42''N, alt. 4 m, edge of a path, on pleistoce-ne sand, 19 October 2012, leg. & det. C. Dolnik, U. de Bruyn, M. Schultz & U. Schiefelbein (Schi 3679).

The species is easily recognized by the prominent perithecia without involucrellum and the 1-septate ascospores. It is widespread in the whole of Germany, also in its northern part (Dolnik et al. 2010,

Schiefelbein et al.: Records of lichenized, lichenicolous and saprophytic fungi from northern Germany 251

Hauck & de Bruyn 2010, Wirth et al. 2013). Litterski (1993) provided the first and hitherto only record from Meckenburg-Western Pomerania.

Trapelia obtegens (Th.Fr.) HertelMecklenburg-Western Pomerania: 1547/1, Isle of Rügen, Feuersteinfelder SW of Mukran, 13°33'31''E/54°28'08''N, alt. 3 m, large natural deposits of flint, on flint, 25 October 2013, leg. & det. C. Dolnik, U. de Bruyn & U. Schiefelbein (Schi 3797; TLC: gyrophoric acid).

New to Mecklenburg-Western Pomerania. This Trapelia species seems to be overlooked in Mecklenburg-Western Pomerania because it occurs in many surrounding regions (Dolnik et al. 2010, Hauck & de Bruyn 2010, Otte & Rätzel 2004, Wirth et al. 2013).

*Tremella pertusariae DiederichMecklenburg-Western Pomerania: 1540/2, Darß peninsula, beech forest along the Tiefenstück-Weg, 12º29'29''E/54º27'07''N, alt. 10 m, acidophilous beech forest, on Fagus sylvatica, on Pertusaria hymenea, 7 September 2013, leg. R. Stordeur, R. Thiemann & U. Schiefelbein, det. U. Schiefelbein, conf. M. Kukwa (Schi 3653); 1640/2, Vordarß, Ahrenshooper Holz, northern part of the forest, 12º26'34''E/54º23'11''N, alt. 2 m, acidophi-lous beech forest, on Fagus sylvatica, on Pertusaria hymenea, 08 September 2013, leg. R. Stordeur, R. Thiemann & U. Schiefelbein, det. U. Schiefelbein (Schi 3391).

New to Mecklenburg-Western Pomerania. The hitherto only records in Germany originate from Flensburg in Schleswig-Holstein by C. F. E. Erichsen (Diederich 1996) and from Bavaria (Brackel 2014).

Verrucaria maculiformis Kremp.Mecklenburg-Western Pomerania: 1747/2, Isle of Rügen, peninsula Groß Zicker, cliff on the western site, 13°39'17''E/54°17'23''N, alt. 1 m, boulder beach, on limestone, 27 October 2013, leg. & det. C. Dolnik, U. de Bruyn & U. Schiefelbein (Schi 3711); Schleswig-Holstein: 1123/2, Flensburger Förde, Holnis peninsula, eastern cape, 09°36'51''E/54°52'32''N, alt. 2 m, limestone bolder, 16 May 2013, leg. & det. C. Dolnik (KIEL, Do 2613); 1326/3, Schwansen, Baltic Sea coast, 10°01'55''E/54°36'59''N, alt. 2 m, shingle beach, on limestone gravel, 13 April 2011, leg. & det. C. Dolnik (Do 2634); 1630/3, Howacht, Baltic Sea coast, 10°36'E/54°18'N, alt. 2 m, active coastal cliff, on pebble in loamy soil, 17 March 2007, leg. & det. C. Dolnik (KIEL, Do 405); 1723/4, Schachtholm, sand bank close to Kiel Canal, 09°36'19''E/54°13'31''N, alt. 7 m, stone pebbles on the sand bank, 27 October 2006, leg. & det. C. Dolnik (Do 343); 1732/4, Dameshöved, Baltic Sea coast, 11°05'33''E/54°12'00''N, alt. 2 m, coastal cliff, brick in loamy soil, 31 October 2010, leg. C. Dolnik & M. Schultz as V. dolosa in Dolnik et al. 2011, rev. Dolnik (KIEL, Do 1745, Schu 07199); 1813/1, Helgoland, cliff E of northern harbour, 07°53'15''E/54°11'01''N, alt. 10 m, inactive cliff, on sandstone, 23 September 2011, leg. C. Dolnik & U. Schiefelbein, det. B. Krzewicka (Do 2739).

New to Mecklenburg-Western Pomerania and Schleswig-Holstein, but probably overlooked or not distinguished from other species of the Verrucaria acrotella group.

Verrucaria tristis (A.Massal.) Kremp.Schleswig-Holstein: 1813/1, Helgoland, northern part of the island, cliff S of the sports field, 07º52'46''E/54º11'10''N, alt. 18 m, inactive cliff, on sandstone, 23 September 2011, leg. & det. C. Dolnik & U. Schiefelbein, conf. B. Krzewicka (Schi 3399).

New to northern Germany. The record on the Isle of Helgoland is very remarkable because in Central Europe the species usually inhabits the high-mountain and alpine belt of calcareous mountains (Krzewicka 2012, Hafellner & Türk 2001, Wirth et al. 2013).

*Xanthoriicola physciae (Kalchbr.) D.Hawksw.Mecklenburg-Western Pomerania (only selected records): 1540/2, Darß peninsula, dunes near the Tiefenstück-Weg, 12º29'08''E/54º27'04''N, alt. 5 m, old beeches on dunes, on Fagus sylvatica, on Xanthoria parietina, 7 September 2013, leg. R. Stordeur, R. Thiemann & U. Schiefelbein, det. U. Schiefelbein (Schi 3549, 3556); 1949/3, Isle of Usedom, peninsula Gnitz, nature reserve Südspitze Gnitz, southern cape, 13º52'00''E/54º00'29''N, alt. 2 m, row of poplar trees, on Xanthoria parietina, 13 March 2004, leg. & det. U. Schiefelbein (Schi 1903); 2034/1, Wismar bay, Hohen Wieschendorf Huk, coast from the marina to the cape, 11°20'25''E/53°57'16''N, alt. 2 m, trees along the coast, on Populus sp., on Xanthoria parietina, 20 October 2012, leg. & det. U. de Bruyn, C. Dolnik, M. Schultz & U. Schiefelbein (Schi 3645); Schleswig-Holstein: 1422/2, Idstedtwesterfeld, meadow near grave mound, 09º29'40''E/54º35'08,4''N, alt. 23 m, on Xanthoria parietina over twigs of Sarothamnus sco-

252 Herzogia 27 (2), 2014

parius, 26 September 2013, leg. & det. C. Dolnik, (KIEL, Do 2516); 1813/1, Helgoland, Oberland, churchy-ard St. Nikolai, 07º53'05''E/54º10'59''N, alt. 39 m, on Xanthoria parietina over Sambucus nigra, leg. & det. C. Dolnik & U. Schiefelbein, 22 September 2011 (Schi 3416); 1813/1, Helgoland, Düne, pond in the centre of the dune, 07º54'56''E/54º11'04''N, alt. 2 m, shrubland, on Salix sp., 24 September 2013, leg. & det. C. Dolnik & U. Schiefelbein (Schi 3420).

New to Schleswig-Holstein. One of the most common and rapidly spreading lichenicolous fungi in Germany (Wirth et al. 2011).

*Zwackhiomyces lecanorae (Stein) Nik.Hoffm. & HafellnerMecklenburg-Western Pomerania: 1336/1, Isle of Rügen, Nordufer Wittow, between Varnkewitz and Schwarbe, 13°21'57''E/54°40'53''N, alt. 1 m, boulder beach, on Lecanora crenulata over limestone, 26 October 2013, leg. U. Schiefelbein, det. U. Schiefelbein (Schi 3775).

New to northern Germany. A rare lichenicolous fungus according to Wirth et al. (2011) but probably overlooked and much more common.

AcknowledgementsWe would like to thank U. Arup (Lund), P. Czarnota (Niedźwiedź), A. Flakus (Krakow), B. Krzewicka (Krakow) and M. Kukwa (Gdańsk) for the revision of specimens. Our warm thanks are also due to C. Skarbek (Minnesota) for linguistic revision.

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Manuscript accepted: 29 September 2014.

Addresses of the authorsPieter P. G. van den Boom, Arafura 16, 5691JA Son, The Netherlands. E-mail: pvdboom@zonnet.nlUwe de Bruyn, von-Müller-Straße 30, 26123 Oldenburg, Germany. E-mail: udebruyn@web.deChristian Dolnik, Ökologie-Zentrum, Institut für Natur- und Ressourcenschutz, Universität Kiel, Olshausenstraße 40, 24098 Kiel, Germany. E-mail: cdolnik@ecology.uni-kiel.deBirgit Litterski, DUENE e. V., Soldmannstraße 15, 17489 Greifswald, Germany. E-mail: litterski@t-online.deRegine Stordeur & René Thiemann, Martin-Luther-University, Institute of Biology, Geobotany and Botanical Garden, Neuwerk 21, 06108 Halle/S., Germany. E-mails: regine.stordeur@botanik.uni-halle.de, sphyrna_zyg@yahoo.de Ulf Schiefelbein, Blücherstraße 71, 18055 Rostock, Germany. E-mail: ulf.schiefelbein@gmx.de Matthias Schultz, Biozentrum Klein Flottbek und Botanischer Garten der Universität Hamburg, Ohnhorststraße 18, 22609 Hamburg, Germany. E-mail: matthias.schultz@uni-hamburg.deHarrie J. M. Sipman, Freie Universität Berlin, Botanischer Garten und Botanisches Museum, Königin-Luise-Straße 6 – 8, 14195 Berlin, Deutschland. E-mail: h.sipman@bgbm.org