botany lecture - chapter 7

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    Root system

    1. Anchoring the plant firmly to a substrate

    (soil)

    2. Absorbing water and minerals

    3. production of hormones

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    Three types of root system

    1. Tap root

    from radicle

    Anchorage

    In dicot andgymnosperm

    Secondary

    growth

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    2. Fibrous

    After the death ofradicle

    Delicate and hair-like

    Absorption

    No prominentenlarged primary root

    In monocot

    http://www.cactus-art.biz/note-book/Dictionary/aaa_Dictionary_pictures/Fibrous_root_system.jpg
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    a. brace root-arise

    from main trunk stem

    b. prop rootarise

    from the lateral branches ofthe main stem.

    Brace roots of corn

    which arise from themain trunk or stem of

    Zea mays

    3. Adventitious do not arise from pre-existing roots

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    Germinating seed showing the emergence of primary

    root from the radicle.

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    1. Taproot single prominent

    root

    2. Lateral or branch roots-

    smaller and arise from the

    taproot

    3. Root hairs

    arise fromepidermis, single celled

    extension which increase

    absorptive area, no line of

    demarcation in the epidermal

    cells.

    4. Root tip- the growing portion

    protected by root cap.

    structure of roots

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    L-section of root shows 3 zones

    Zone of maturation

    Zone of elongation

    Zone of division/apical

    meristem

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    peripheral

    columella

    Rootcap

    ROOT TIP IS PROTECTED BY A ROOT CAP

    procambium

    Ground meristem

    protoderm

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    Root cap

    Slime or mucigelsecreted by the

    dictyosomes of the root cap

    which helps to lubricate the

    passage of root through the soil.

    Quiescent centeris the mitotically

    inactive region of the root apical

    meristem, act as reserve of

    healthy cells, resistant to toxic

    substances and radiations.The three zones of root apical

    meristem:

    1. protoderm

    2. ground meristem

    3. procambium

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    Cross section of

    Ranunculus root (Buttercup)

    Internal anatomy of young dicot root

    shows three general sections:

    1. Dermal region epidermis with

    root hairs

    2. Cortex outer collenchyma

    middle parenchyma

    inner endodermis

    3. Stele or vascular cyclinder

    consists of the following:

    1. pericycle outermost layer

    2. primary xylem

    3. primary phloem

    4. vascular cambium

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    B

    C

    D

    E

    b.Pericycle

    c.Primary phloem

    d. Vascular cambium

    e. Primary xylem

    Cross section of young dicot root

    Beside is the enlarged view of

    Vascular cylinder or stele region.

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    Variations in the number of xylem strands in roots

    Triarch tetrarch pentarch polyarch

    Three xylem poles four xylem poles five xylem poles many xylem poles

    eudicot eudicot eudicot monocot

    Protostele protostele protostele siphonostele

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    Endodermis- controls the

    passage of minerals

    across the vascular

    tissues

    Consist of thick-walled cells

    (encrusted with

    suberin and lignin)

    and passage cells

    (thin-walled)

    Casparian stripsare bandsof lignin and suberin on the radial

    walls (top, bottom and side walls)

    causing the cell walls to be waterproof.

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    Diffusion paths in rootsApoplastic

    Water and mineral

    diffuses only through

    walls in intercellular

    spaces

    Symplastic

    The material has passed

    through a plasma membrane

    and enters the protoplasm

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    Internal anatomy of monocot root Internalanatomy of monocot rootshows 3 regions:

    1. Dermal region epidermis

    2. Cortex which shows:

    a. outer zone of collenchyma

    b. middle zone of parenchyma

    c. innermost endodermid

    3. Vascular Cylinder

    a. pericycle

    b. xylem

    c. phloem

    d. pith parenchyma

    Siphonostele

    a band of vascular

    tissues surrounding a

    parenchymatous pith.

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    Monocot root

    Epidermis

    Outer collenchyma

    Middle parenchyma

    Endodermis

    PithPericycle

    Primary xylem

    Primary phloem

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    Vascular cylinder of monocot root

    siphonostele strands of xylem and phloem

    surrounding a parenchymatous pith

    pithpith

    O i i d D l t f L t l t

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    Origin and Development of Lateral roots

    Lateral root

    Vascular tissues

    Expaded epidermis

    And cortex of parental

    root

    Lateral rootis iniatiated by

    cell divisions in the

    pericycle. As it pushes

    outward, it destroys the

    cells of the cortex and

    epidermis that lie in its

    path, ultimately destroying

    endodermis.

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    Old dicot root

    Old dicot root shows two distinct

    regions:

    1. Region of secondar vascular

    tissues which include the

    a. secondary phloem

    b. vascular cambium

    c. secondary xylem

    2. Region of periderm layers:

    a. phellem or cork

    b. phellogen or cork cambium

    c. phelloderm or cork parenchyma

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    Brace roots in corn plantfor additional supportand absorption

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    Water retention 3. velamen in aerial roots of

    orchids

    photosynthesis 4. chlorophyllous roots

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    Movement - Contractile roots of bulbs like onions,

    gladiolus, garlic

    http://www.cactus-art.biz/schede/HAWORTHIA/Haworthia_emelyae/Haworthia_emelyae_comptoniana/Haworthia_emeliae_v_comptoniana_contractile_roots_540.jpg
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    Parasitic (haustorial roots) for

    absorption

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    Buttresses- expanded roots for

    great support

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    Pneumatophores- breathing roots

    of mangove for aeration

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    enlarged fleshy tap root- carrots, ube,

    tugue, radish, turnips,

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    Mycorrhizae- association between

    a soil fungus and roots

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    Other types and root modifications

    8. Protection 8. spiny roots of tugue

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    Other types and root modifications

    5. root nodules associations between

    nitrogen fixing bacteria

    and roots of legumes

    such as makahiya.

    6. Parasitism 6. haustorial roots of

    parasitic flowering

    plants like mistle toe