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Timber and Wildlife Ecology Research Update from the Jones Center Kevin McIntyre Education Coordinator & Steve Jack Conservation Ecologist Forest Landowner’s Association 2014 Southeast Regional Forest Resource Owner and Manager Conference Valdosta, GA October 28, 2014

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Longleaf Pine Ecosystems Productivity and biodiversity patterns of a longleaf pine ecosystem. Ecological forestry and restoration of longleaf pine ecosystems. Ecological role of mesopredators, effects of control, and habitat approaches. Aquatic Ecology and Water Resources Hydrologic variation and human development in the lower Flint River Basin Depressional wetlands on the coastal plain landscape: maintenance of regional biodiversity

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Page 1: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Timber and Wildlife Ecology Research Update from the Jones

Center

Kevin McIntyreEducation Coordinator

&Steve Jack

Conservation Ecologist

Forest Landowner’s Association 2014 Southeast Regional Forest Resource Owner and Manager Conference

Valdosta, GA October 28, 2014

Page 2: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Joseph W. Jones Ecological Research Center at Ichauway

Supported by the Robert W. Woodruff Foundation

Page 3: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Ichauway, circa 1929

Robert Woodruff

Joseph W. Jones

Joseph W. Jones Ecological Research Center at Ichauway

Page 4: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA
Page 5: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Long-Term Research

Longleaf Pine Ecosystems1) Productivity and biodiversity patterns of a

longleaf pine ecosystem.2) Ecological forestry and restoration of longleaf

pine ecosystems.3) Ecological role of mesopredators, effects of

control, and habitat approaches.

Aquatic Ecology and Water Resources4) Hydrologic variation and human development in

the lower Flint River Basin5) Depressional wetlands on the coastal plain

landscape: maintenance of regional biodiversity

Joseph W. Jones Ecological Research Center at Ichauway

Page 6: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Conservation & Natural ResourceManagement

• Stewardship of Ichauway & its natural resources

• Management of forests, wildlife & wetlands, including prescribed fire

• Monitoring of key environmental variables, plant & animal populations

• New approaches for restoration & adaptive management

• Key role in participation with education & research programs

Page 7: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Education and Outreach

• Natural Resource Professionals

• Policymakers

• Land Owners/Managers

• Undergraduate University Classes

• Graduate Students

Page 8: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

• Overall goal of doubling acreage of LLP

from 4m to 8m by 2025

• Supported by diverse coalition of

Federal, State, NGO and private sector

• Identified Significant Geographic Areas

for longleaf restoration focus

• Significantly increased funding for LLP

• Decline of LLP reversed from low of

3.84m to 4.28m as of 2010

• 157,000 acres LLP established in 2013

• 50% incentive funded

• 50% funded by private landowners

Regional Longleaf Pine Conservation

and Restoration

8

Page 9: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Forest Management Limitations

Timber

Page 10: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Forest Management Limitations Tradeoffs

Timber

Timber

Page 11: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Wildlife Research

Quail

Page 12: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Is Hard Mast a Limiting Factor?

Page 13: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Ichauway Turkey Track Count Data

Page 14: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Ichauway WTD Track Counts

Page 15: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Ichauway WTD Spotlight Data

Page 16: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA
Page 17: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Season of Burn & Wildlife Impacts

Page 18: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Growing-Season Fire Impacts

Nest Success:

• Growing-season fires had an effect (p-value=0.03)– 11% of nests burned

– Majority burned in April

– 80% of hens renested

Poult Survival:

• One newly hatched brood lost to fire

Page 19: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Poult Survival Data

Year Site n

Day 0-9

(%lost)

Day 10-16

(%lost)

Day 17-30

(%lost)

Day 30+

(%lost)

2011 JC 6 67 17 17 N/A

SL 3 33 0 33 0

Pooled 9 56 11 22 0

2012 JC 9 78 11

SL 5 40 0 20 40

Pooled 14 64 7 7 21

Both Years/Sites 23 61 17 26 22

• Average # of poults per hen at hatch: 8.6

• 5 of the 23 brood hens killed by predators

Page 20: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Management Implications

Predation much stronger driver of turkey population dynamics than season of burn

Use rotating small-scale growing-season burns

More to Wild Turkey Population Dynamics than Burns…

Manage for habitat quality and population vs. individual nests

Page 21: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Predator Exclusion Experiment

Page 22: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Do Exclosures Exclude?

Page 23: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Annual Coyote Diet

Page 24: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Indirect Effects?

Photo credit: Austin Baggarley

Page 25: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Deer use of predator exclosures

0

10

20

30

40

50

60

70

Gestation Fawning Breeding

Wh

ite-

tail

ed d

eer

det

ecti

on

s

Control

Exclosure

Page 26: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

22 fawns/year0.41 vs. 0.77 fawns/doe

Page 27: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Longleaf Management and Restoration Research

Page 28: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Timber Research

Timber

Page 29: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Conventional Wisdom

USDA Silvics Manual – Volume 1: Conifers (Agriculture Handbook 654, 1990)

Pinus palustris (Mill.)(from chapter by Boyer)

“Reaction to Competition-Longleaf pine is intolerant of competition, whether for light or for moisture and nutrients. The species will grow best in the complete absence of all competition, including that from other members of the species.”

Thus, general recommendation was to manage using even-aged systems (Boyer and Peterson, 1983, Agriculture Handbook 445)

Page 30: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Natural Stands – Multi-aged

Page 31: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Regeneration Dynamics Questions

• Below-canopy light environment

• Competition (both inter-and intra-specific)

• Role of fire

• Management implications

Page 32: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

CompetitionPine Seedling Response

to Canopy Gap Size

Seedling growthT

ota

l b

iom

as

s (

g)

0

5

10

15

20

25

30

Seedling survival

Overstory treatment

Uncut

Single

-tree

Small

group

Large g

roup

Me

an

su

rviv

al

(%)

60

65

70

75

80

85

a

ab

cbc

a

a a

b

Pecot et al. 2007. CJFR

Page 33: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Three Stage Model for

Seedling Survival and Growth

Page 34: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Canopy Gap Size & Advanced Regeneration

Think in terms of releasing advanced regeneration rather than creating opening to secure regeneration

Page 35: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Canopy Gap Size & Advanced Regeneration

O’Brien et al. 2008. Ambio

Page 36: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Diverse Ground Cover Community

Page 37: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Functionally important ground cover species

Hendricks et al. 2002

Days in Field

0 200 400 600 800 1000 1200

Ma

ss R

em

ain

ing

(%

)

0

10

20

30

40

50

60

70

80

90

100

Wiregrass (soil)

Wiregrass (elevated)

Longleaf pine (soil)

Longleaf pine (elevated)

Lower decomposition rate of elevated fine fuels(wiregrass & pine needles)

Page 38: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Legume Species

Fo

liar

Nd

fa (

%)

0

10

20

30

40

50

60

70

80

90

100

Wet-Mesic Site

Intermediate Site

Cassianictitans

Crotalariarotundifolia

Centrosemavirginiana

Lespedezavirginiana

Legumes

No. of species identified: 43

Average density: 115,000stems/ha

Potential N2-fixation by legumes

Functionally important species:

Hainds et al. 1999

Hiers et al. 2001

Cathey et al. 2010

Page 39: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

How Do We Best Manage to Maintain and Restore This Forest Type While Utilizing Resources?

Page 40: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Financial Comparison

Harvest Scenario Low Medium High

20-year IRR w/ land value (%) 3.21 3.29 3.48

Total accumulated net cash flow ($) 1,796,027 3,732,401 4,986,258

Ending Total Value (Cash flow + ending value) ($) 6,703,877 7,445,119 8,104,815

Conversion Scenario Moderate Rapid

20-year IRR w/ land value (%) 4.21 5.77

Total accumulated net cash flow ($) 5,171,745 4,928,824

Ending Total Value (Cash flow + ending value) ($) 6,612,212 6,569,429

Page 41: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Managing Longleaf Forests

Using natural disturbance regime as guide to management uneven-aged approaches

Page 42: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Which Silvicultural System?

• Disturbance regime and regeneration dynamics point toward uneven-aged systems to meet objectives

• Group or individual tree selection are feasible approaches for restoring toward reference objective

Page 43: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Long-Term Management and Restoration Experiments

Page 44: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Fuels, Fire & Silviculture

Attentuation of Needle Cast with Distance from Gap Edge

Distance from Gap Edge

-2 0 2 4 6 8 10 12 14 16

Needle

Cast

(gra

ms p

er

1/4

m2 t

rap)

0

10

20

30

40

grams of needles/ trap

Page 45: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Regeneration Dynamics

Page 46: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Ground Cover Recovery

Page 47: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Harvest Treatments

Uncut Control

Group Selection

Single-tree Selection

Group Selection

with Retention

Page 48: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Stem Maps

Page 49: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Skidder Traffic – Operational Example

* Also have “experimental” skidder treatments

Page 50: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Soil and Ground Cover Disturbance

4 Passes 8 Passes

1 Pass 2 Passes

Page 51: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Soil Compaction Operational Skid

Trails

Page 52: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Ground Cover ChangesWiregrass Dominated

Pre-harvest, 2009 Post-harvest, 2010

Page 53: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Ground Cover ChangesOld-Field Dominated

Pre-harvest, 2009 Post-harvest, 2010

Page 54: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Planted Seedling Biomass

Page 55: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Management Project Summary

• Still preliminary results, early in study

• Short-term weather patterns can affect results

• Few statistically significant results due to high variability, though are some evident trends

• Harvesting operations have impacts but do not appear severe, recovery occurs over long time span

Page 56: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Restoration Research

Restoration – timing and pathways

– Species conversion

– When to introduce ground cover

– Wildlife habitat and which species present at different stages of development

Page 57: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Plantations to “Natural” Stands

Page 58: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Plantation Species Conversion

Underplanting longleaf seedlings in gaps

Thinning planted slash pine with gap creation

Reintroduction of ground cover?- when and what species

Page 59: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Old Field Conversion

Page 60: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Ground Cover Restoration

Page 61: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Hardwood Encroachment in Uplands

Legacy of past management

How to reconnect the landscape?

Page 62: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Hardwood Encroachment in Uplands

• Silvicultural treatment is species conversion – removal followed by planting

• Focused on trees, other things not immediately restored

Page 63: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

• Explicitly factor time into considerations –thinking of time as an ecological factor that cannot be purchased or replaced

• In restoration, make use results of time already “invested”

Page 64: “Timber and Wildlife Ecology Research Update from the Jones Center”  Kevin McIntyre & Steve Jack, Joseph W. Jones Ecological Research Center, Newton, GA

Summary

• Integration between research and operational management and between research subject areas

• Research is focused on long-term projects with operational scale

• Attempting to examine multiple resource responses to manipulations – trade-offs from maintaining all ecosystem characteristics