static-content.springer.com10.1007... · web viewdesignated 7350-6, selected from the piping rock...

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Online Resource 5. Breeding history/origin summaries for 78 Agrostis, Apera, and Polypogon cultivars, accessions, and collections in the current study. Entry order follows the order presented in the UPGMA diagram (Fig. 1), with the exception of the outgroup species of Apera and Polypogon.

Entry Breeding History or Origin

007 A. stolonifera Developed using 23 parent plants – 19 plants originating from Rutgers University

breeding program and 4 plants originating from Blue Moon Farms (Lebanon, Oregon).

Rutgers clones: selections from ‘L-93’ and ‘Southshore’; collections tracing back to

putting greens from the Piping Rock Club (Locust Valley, NY), Spring Lake GC (Spring

Lake Heights, NJ), Northshore CC (Glen Head, NY), Rumson CC (Rumson, NJ), and

Harkers Hollow Golf and CC (Harmony, NJ); and collection from a golf course fairway

in Chicago, Illinois. Blue Moon Farms clones: trace to germplasm from L-93 and

collection from old putting green in Atlanta, GA. (Rutgers, the State University of New

Jersey and R.H. Hurley 2009)

Pinup A. stolonifera Maternal progenies of 13 clones collected from old golf course putting greens and

fairways – 46% of the germplasm tracing to collections from the Piping Rock club

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(Locust Valley, NY); 31% of the germplasm tracing to a plant collected from Harkers

Hollow Golf and CC (Harmony, NJ); and 23% of the germplasm tracing to a plant

collected from National Golf Links of America (Southampton, NY). The creeping

bentgrass plants selected from old golf courses were of unknown origin, however, it is

likely that these plants originated from South German mixed bentgrass as this was one of

the few commercially available sources for bentgrass seed in the early 1900’s. (Stacy

Bonos – personal communication)

AP18 A. stolonifera Maternal progenies of 10 clones collected from old golf course putting greens and

fairways – 10% tracing to a plant selected from Harkers Hollow Golf and CC (Harmony,

NJ); 20% tracing to a plant selected from Rumson CC (Rumson, NJ); 20% tracing to a

plant selected from the Piping Rock Club (Locust Valley, NY); 10% tracing to a plant

selected from Spring Lake GC (Spring Lake Heights, NJ); and 40% tracing to L93-10 (a

dollar spot resistant clone selected from L93). (Stacy Bonos – personal communication)

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13M A. stolonifera Maternal progenies of 21 clones collected from old golf course putting greens and

fairways - 32% of the germplasm tracing to several plants collected from putting greens

and fairways from golf courses in Arizona; 19% of the germplasm tracing to a plant

selected from or related to ‘L93’; 19% of the germplasm tracing to a plant selected from

the Piping Rock Club in Locust Valley, NY; 10% tracing to a plant selected from Lake

Success Golf Club in Great Neck, NY; 10% tracing to a plant selected from Hollywood

Golf Club in Deal, NJ; and another 10% tracing to a plant selected from Barrington Hills

CC in Barrington, IL. The creeping bentgrass plants selected from old golf courses were

of unknown origin, however, it is likely that these plants originated from South German

mixed bentgrass as this was one of the few commercially available sources for bentgrass

seed in the early 1900’s. (Rutgers, the State University of New Jersey 2009a)

SRP1WM A. stolonifera Traces to clones selected from long-term research plots at Michigan State University

(Leah Brilman – personal communication)

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Barracuda A. stolonifera Maternal progenies of 9 clones collected from old golf course putting greens and

fairways – 44.4% of the germplasm tracing to a plant collected from Spring Lake Golf

Club in Spring Lake Heights, NJ; 33.3% of the germplasm tracing to several plants

selected from the Piping Rock Club in Locust Valley, NY; 11.1% tracing to a plant

selected from Lake Success Golf Club in Great Neck, NY; and 11.1% tracing to a plant

collected from a golf course in Portugal. The creeping bentgrass plants selected from old

golf courses were of unknown origin, however, it is likely that these plants originated

from South German mixed bentgrass as this was one of the few commercially available

sources for bentgrass seed in the early 1900’s. (Peak Plant Genetics LLC and Rutgers, the

State University of New Jersey 2013)

Independence A. stolonifera Maternal progenies of 14 clones collected from old golf course putting greens and

fairways – Attractive plants were collected from Harkers Hollow Golf and CC (Harmony,

NJ), Old Westbury Golf and CC (Old Westbury, NY) Suburban Golf Club (Union, NJ),

Canoe Brook Country Club (Summit, NJ), Shackamaxon Country Club (Scotch Plains,

NJ), the Piping Rock Club (Locust Valley, NY), and Lake Success Golf Club (Great

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Neck, NY). The creeping bentgrass plants selected from old golf courses were of

unknown origin, however, it is likely that these plants originated from South German


seed in the early 1900’s. Additionally, approximately 7% of the parental germplasm

traces to plants selected from or related to ‘L93’. (Lebanon Seaboard Corporation and

Rutgers, the State University of New Jersey 2005)

King Pin A. stolonifera Maternal progenies of 18 clones collected from old golf course putting greens and

fairways – 89% of the maternal germplasm traces to a plant designated ‘AZ-8’, selected

from an Arizona golf course. 11% of the maternal germplasm traces to a plant designated

‘AZ-144’, selected from an Arizona golf course. The creeping bentgrass plants selected

from old golf courses were of unknown origin, however, it is likely that these plants

originated from South German mixed bentgrass as this was one of the few commercially

available sources for bentgrass seed in the early 1900’s. (Stacy Bonos – personal

communication)

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Luminary A. stolonifera Maternal progenies of 6 clones with improved dollar spot resistance – 50% of the

germplasm tracing to collections made in 1998 from Lake Waramaug Country Club in

New Preston, CT; 33% of the germplasm tracing to outstanding clones from an L-93

research trial border planted at Rutgers Horticultural Research Farm II, in New

Brunswick, NJ in 1996; 17% of the germplasm tracing to collections made in 1996 from

the Piping Rock Club in Locust Valley, NY (Stacy Bonos – personal communication).

Focus A. stolonifera Maternal progenies of 9 clones collected from old golf course putting greens and

fairways – 56% of the germplasm tracing to a plant collected from the Piping Rock Club

in Locust Valley, NY; 11% tracing to a plant designated ‘AZ-144’ from an Arizona golf

course; and 33% tracing to a segregating clone of a Penncross seeded research trial

border area in North Brunswick, NJ. The creeping bentgrass plants selected from old

golf courses were of unknown origin, however, it is likely that these plants originated

from South German mixed bentgrass as this was one of the few commercially available

sources for bentgrass seed in the early 1900’s. (Stacy Bonos – personal communication)

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Alpha A. stolonifera The origin of ‘Alpha’ traces to 15 lines: 3 of the 15 lines traced to a Massachusetts golf

course; 3 lines traced to a North Carolina golf green; and the remaining lines traced to

selections of low growing surviving plants from a herbicide damaged Foundation field of

‘Putter’ creeping bentgrass in the Willamette Valley of Oregon. Selections of segregated

patches from old putting greens were 30 to 100 years old and were likely sown to South

German Mixed Bentgrass or ‘Penncross’. (Brede 2007)

Authority A. stolonifera Traces to 2 plants designated C952 and C953 selected from an experimental turf trial

established in 1992 at the Pinehurst Resort (Pinehurst, NC) (Novel Ag Inc 2009)

Shark A. stolonifera Fifty percent of ‘Shark’ traces to 2 plants designated C952 and C953 selected from an

experimental turf trial established in 1992 at the Pinehurst Resort (Pinehurst, NC). An

additional 50% of Shark traces to clones collected from putting greens of old golf courses

– golf course collection sites include Gulph Mills Golf Club (Conshohocken, PA), Brae

Burn Country Club (West Newton, MA), Richmond County Country Club (Staten Island,

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NY), and Green Brook Country Club (North Caldwell, NJ). The creeping bentgrass

plants selected from old golf courses were of unknown origin, however, it is likely that

these plants originated from South German mixed bentgrass as this was one of the few

commercially available sources for bentgrass seed in the early 1900’s. (Novel Ag Inc and

Rutgers, the State University of New Jersey 2009)

V8 A. stolonifera V8 traces to plants from a collection that were tested in a turf trial at the Pinehurst Resort

in Pinehurst, NC in 1992. Three selected clones (PH-N, PH-S, PH-TC) were turf tested at

Rutgers NJAES in North Brunswick, NJ, along with twelve attractive progeny from

them, labeled R1-R12. Remnant seed saved from the PH-N and R2 plants were used to

establish a spaced plant breeder block of V8 in June 2010 near Connell, WA. (Tom

Brentano – personal communication)

Benchmark DSR A. stolonifera Maternal progenies of 13 clones collected from old golf course putting greens and

fairways – 76% of the maternal germplasm traces to plants designated AZ-8 and AZ144,

selected from an Arizona golf course; 14% of the maternal germplasm traces to a plant

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designated 7350-6, selected from the Piping Rock Club in Locust Valley, NY; and 8% of

the germplasm traces to a plant collected from a golf course fairway at Barrington Hills

CC in Barrington, IL. The creeping bentgrass plants selected from old golf courses were

of unknown origin, however, it is likely that these plants originated from South German


seed in the early 1900’s. (Rutgers, the State University of New Jersey 2009b).

Memorial A. stolonifera Maternal progenies of 25 clones from old golf course putting greens and fairways – 27%

of the germplasm traces to plants selected from the Piping Rock Club in Locust Valley,

NY; 14% tracing to plants selected from a fairway on Lake Success Golf Club in Great

Neck, NY; 13% tracing to a plant designated AZ-8 from an Arizona golf course; 13%

tracing to a plant designated AZ-64 from an Arizona golf course; 13% tracing to a plant

designated AZ-144 from an Arizona golf course; 13% tracing to a plant selected from or

related to L-93; 7% tracing to a plant designated 7356-5 collected from Hollywood Golf

Club in Deal, NJ. The creeping bentgrass plants selected from old golf courses were of

unknown origin, however, it is likely that these plants originated from South German

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seed in the early 1900’s. (Stacy Bonos – personal communication)

Century A. stolonifera Polycross of 6 clones: clone TAES 2831 (collected from green #9 Brookhollow CC,

Dallas, TX in 1984); clone TAES 3153 (collected from Rome Acquasanta GC Italy in

1986); clone TAES 3250 (from Nimes Campagne GC France 1986); clone TAES 3307

(also identified as U6 from Firenze GC France 1986); clone TAES 3794 (individual plant

selected from Syn4 (Cato)); and TAES 3799 (individual plant selected from Syn4

(Cato)). (Texas Agricultural Experiment Station 2004a).

Imperial A. stolonifera Seven clone polycross synthetic: clone TAES 2833 (collected from green #9

Brookhollow CC, Dallas, TX in 1984); clone TAES 2845 (collected from green #9

Brookhollow CC, Dallas, TX in 1984); clone TAES 2916 (collected from green #14

Walnut CC, East Lansing, MI in 1984); clone TAES 2922 (collected from green #5

Platteville CC, Platteville, WI in 1984); clone TAES 3106 (also identified as AGBP-1/#4

from Kevin McVeigh, Oregon in 1986); TAES 3293 (from Nimes Campagne GC France

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1986); and TAES 3307 (also identified as U6 from Firenze GC France 1986). (Texas

Agricultural Experiment Station 2004b).

Unknown A. stolonifera Unknown contaminate creeping bentgrass plants in ‘Greenwich’ velvet bentgrass seed lot

(M50-9-70351)

Southshore A. stolonifera Broad based cultivar derived from the progenies of 203 selected clones. Clones were

collected from old golf course putting greens primarily in New Jersey, New York,

California, Arizona, and Pennsylvania. (Lofts Seed Inc 1994)

L93 A. stolonifera Progeny of 19 clones. Plant material originally traces to 15 maternal clones of the 203

clone ‘Southshore’ nursery (see above). The 15 clones were from plants collected in

New Jersey, New York, and Arizona. Seed harvested from 15 clones was subjected to

additional cycles of selection: approximately 1000 seedlings from original 15 clones

panted in a nursery at Rutgers Plant Science Research Farm, Adelphia, NJ in 1987; 34 of

the most promising plants subsequently vegetatively established at Lofts Research Farm,

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Martinsville, NJ in 1989; vegetative propagules of selected clones established in an

isolated polycross nursery at Lofts Great Western Research Farm in Lebanon, OR in

1990, with progeny subsequently planted at Martinsville NJ in 1991. L93 breeders seed

nursery established using 12 clones from the 1989 Martinsville NJ trial, 1 clone from a

‘Southshore’ production field in Oregon, and six of the progenies of the 1991

Martinsville NJ trial. (Richard Hurley – personal communication)

Penncross A. stolonifera First generation synthetic produced by random crossing of three vegetatively propagated

clonal strains. Developed by H.B. Musser in 1954 (Hein, 1958). As one of the first

improved seeded varieties of creeping bentgrass, Penncross was used to seed or overseed

many golf courses throughout North America (Musser 1959; Bonos and Huff 2013).

Penneagle II A. stolonifera Population improvement project from ‘Penneagle’. Plants that produced breeder seed

trace to the following sources: 53% to PST-OPN, of which 44% trace maternally to

Penneagle; 23.5% to ‘Pennlinks’ pollinated by Penneagle; 8.5% to Penneagle; 8.5% to

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Pennlinks; 3.5% to PennA1 pollinated by Penneagle; and 3% to ‘Seaside II’ pollinated by

Penneagle. (Pure Seed Testing Inc 2006)

SR 1119 A. stolonifera Five clone synthetic. Clones trace back to breeder block of ‘Providence’ (SR 1019)

creeping bentgrass. (Seed Research of Oregon Inc 2004)

Pennlinks A. stolonifera Original parental clone selected from a putting green at The Country Club, Shaker

Heights, Ohio, in 1964, with The Country Club putting greens originally established to

South German Mixed Bentgrass in 1930. Original clone outcrossed with 9 Penn State

experimental bentgrass selections in 1966. Progeny subjected to two cycles of selection

and then backcrossed to original parental clone. Backcrossed progeny subjected to

additional selection, followed by first generation breeder seed of Pennlinks produced

utilizing approximately 100 selected parental clones. (Pennsylvania Agricultural

Experiment Station 1987)

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PennA4 A. stolonifera Polycross procedure where single parent selection A4 was crossed with experimental

selections of Penn G-series followed by two generations of crossing selected sib plants of

PennA4. Penn A-series collected from four putting greens of the main course at Augusta

National Golf Club where putting greens were originally seeded to Penncross creeping

bentgrass. Penn G-series collected from two putting greens of the Par 3 course at

Augusta National Golf Club where putting greens were originally seeded to Penneagle

creeping bentgrass. (Pennsylvania Agricultural Experiment Station 2004)

Proclamation A. stolonifera Maternal progenies of 12 different clones with improved dollar spot (Sclerotinia

homoeocarpa F.T. Bennet.) resistance. Sixty-five percent of the germplasm used in the

development of Proclamation creeping bentgrass traces to collections made in 1996 from

the Piping Rock Club in Locust Valley, NY. Seventeen percent trace to plants collected

from several golf courses in Northern Italy. Ten percent trace to plants collected from

several golf courses in Eastern France. Eight percent trace to a plant collected from

Harker’s Hollow Golf course in Harmony, NJ in 1996. (Stacy Bonos – personal

communication)

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T1 A. stolonifera The origin of ‘T1’ traces to 11 clonal lines: one clone traced to a selection from the

cultivar ‘Providence’; one clone traced to a plant selected from a South Carolina putting

green; one clone traced to a selection from ‘Putter’; 3 clones traced to two Massachusetts

putting greens; and the remaining clones traced to Oregon polycross 94-8283 (originating

from a Massachusetts putting green) and ‘Putter’. Selections of segregated patches from

old putting greens were 30 to 100 years old and were likely sown to South German

Mixed Bentgrass or ‘Penncross’. (Brede 2007)

Carmen A. stolonifera European cultivar

Penneagle A. stolonifera Four clone synthetic with the following origin of parental clones: 1) 3rd generation

seedling selection from ‘Washington’ bent (1938 source); 2) 3rd generation seedling

selection from ‘Seaside’ (1930 source); 3) 3rd generation seedling selection from Cocoos

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bent (1932 source); 3rd generation seedling selection from ‘Washington’ bent (1933

source). (Pennsylvania Agricultural Experiment Station 1980)

Crenshaw A. stolonifera Six clone synthetic: 5 clones obtained from the University of Arizona and 1 clone

collected from Brookhollow CC (Dallas, TX). (Texas Agricultural Experiment Station

1996).

Providence A. stolonifera 5 clone synthetic: 1) clone P19 – 1965 selection of a creeping bentgrass plant from plot of

‘Piper’ velvet bentgrass in a putting green trial at the University of Rhode Island; 2) clone

P-7 - 1965 selection of a creeping bentgrass plant from plot of ‘Arlington’ creeping

bentgrass in a putting green trial at the University of Rhode Island; 3) clone P-27 - 1965

selection of a creeping bentgrass plant from a plot of ‘Highland’ bentgrass; 4) clone P-43

- 1976 selection from polycross progeny from University of Rhode Island collections and

selections; and 5) clone P-87 – 1972 selection from old putting green. (University of

Rhode Island 1991)

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Putter A. stolonifera Hybrid of two lines: AP-10 and pollen parent BPA-163. AP-10 is an improved line

developed through consecutive cycles of intercrossing 26 bentgrass clones collected from

around the US Northwest and the United States. BPA-163 was a single plant selection

from the Puyallup Research and Extension Center (Washington State University) that

exhibited high seed yield potential. (Brauen et al. 1993; Jacklin Seed Company 1991)

Emerald A. stolonifera Single clone selection from ‘Congressional’ bentgrass (Alderson and Sharp 1994)

PI 531251 Hungary A. stolonifera ‘Keszthelyi 4’ – cultivar from Hungary

Seaside A. stolonifera Landrace originally harvested for seed from naturalized stands in the Coos Bay, Umpqua,

and Siuslaw regions of Oregon (Schoth 1930; Warnke 2003).

PI 235440 Switzerland A. stolonifera Collection from near Schuls, Switzerland

PI 494118 Ukraine A. stolonifera ‘Klonobaja’ – cultivar from Ukraine

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PI 302902 Spain A. stolonifera Collection from Spain

PI 318934 Spain A. stolonifera Collection from Zaragoza, Spain

PI 608809 US A. stolonifera Maternal progeny of MSCB-3 from Mississippi Agriculture and Forestry Experiment

Station, polycrossed with MSCB-4, MSCB-10, MSCB-11, MSCB-19, MSCB-37,

MSCB-52, MSCB-73, MSCB-81, MSCB-304, MSCB-330, and MSCB-388; where

MSCB-3 and MSCB-4 were derived from tissue culture regenerated callus of Penncross

on culture media with high salt concentration; MSCB-10, MSCB-19, MSCB-37, MSCB-

52, MSCB-81, MSCB-304, and MSCB-388 were developed from tissue culture

regenerated callus of Penncross exposed to high temperature; and MSCB-11, MSCB-73,

and MSCB-304 were ecotypic selections from Mississippi. (Krans et al. 2000).


Station, polycrossed with MSCB-3, MSCB-10, MSCB-11, MSCB-19, MSCB-37,

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MSCB-52, MSCB-73, MSCB-81, MSCB-304, MSCB-330, and MSCB-388; where

MSCB-3 and MSCB-4 were derived from tissue culture regenerated callus of Penncross

on culture media with high salt concentration; MSCB-10, MSCB-19, MSCB-37, MSCB-

52, MSCB-81, MSCB-304, and MSCB-388 were developed from tissue culture

regenerated callus of Penncross exposed to high temperature; and MSCB-11, MSCB-73,

and MSCB-304 were ecotypic selections from Mississippi. (Krans et al. 2000).

Declaration A. stolonifera Maternal progenies of 8 clones with improved dollar spot (Sclerotinia homoeocarpa F. T.

Bennet.) resistance. 72% of the maternal germplasm traces to several plants selected

from the Piping Rock Club in Locust Valley, NY in 1996. 14% of the maternal

germplasm traces to a putting green collection from Spring Lake Golf Club in Spring

Lake Heights, NJ in 1996, and another 14% traces to a plant selected from Harkers

Hollow Golf and CC in Harmony, NJ in 1996. The creeping bentgrass plants selected

from old golf courses were of unknown origin, however, it is likely that these plants

originated from South German mixed bentgrass as this was one of the few commercially

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available sources for bentgrass seed in the early 1900’s. (Lebanon Seaboard Corporation

and Rutgers, the State University of New Jersey 2009a).

National A. stolonifera Originated from plant selections from old turf areas made by A. C. Ferguson in 1963,

1967, and 1973 near Winnepeg, Manitoba. (Pickseed West Inc. 1991)

PI 221906 Afghanistan A. stolonifera Collection from Afghanistan


Station, polycrossed with MSCB-3, MSCB-4, MSCB-10, MSCB-19, MSCB-37, MSCB-

52, MSCB-73, MSCB-81, MSCB-304, MSCB-330, and MSCB-388; where MSCB-3 and

MSCB-4 were derived from tissue culture regenerated callus of Penncross on culture

media with high salt concentration; MSCB-10, MSCB-19, MSCB-37, MSCB-52, MSCB-

81, MSCB-304, and MSCB-388 were developed from tissue culture regenerated callus of

Penncross exposed to high temperature; and MSCB-11, MSCB-73, and MSCB-304 were

ecotypic selections from Mississippi. (Krans et al. 2000).

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PI 269838 Germany A. stolonifera Collected material from Germany

PI 659907 Kyrgyzstan A. stolonifera Collection from Ysyk-Kol, Kyrgyzstan

PI 204390 Turkey A. stolonifera Collection from near Kayseri, Turkey

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PI 423036 Spain A. stolonifera Collection from Rio Guadarrama at Route 6, west of Madrid

PI 235541 Sweden A. stolonifera Collection from natural pasture near Landskrona, Sweden

PI 283174 A. transcaspica Collection from former Soviet Union

PI 255869 A. gigantea Collected material from Poznan, Poland

PI 619558 A. gigantea Collection from Xinjiang, China

PI 439027 Soviet Union A. stoloniferaa Collection from former Soviet Union

PI 608813 US A. stoloniferaa Maternal progeny of MSCB-19 from Mississippi Agriculture and Forestry Experiment




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PI 222073 Afghanistan A. stoloniferaa Collection from east base of Sibar Pass, Kabul, Afghanistan

PI 230235 Iran A. stoloniferaa Collection from Dastana, Iran

W6 21826 Ukraine A. stoloniferaa Collection from Krym, Ukraine

Alister A. capillaris Origin traces to 10 colonial bentgrass plants collected from bluegrass roughs at the Piping

Rock Club, Locust Valley, NY. (Pure Seed Testing Inc 2004)

FT12 A. capillaris FT12 colonial bentgrass was selected from the maternal progenies of 11 clones – 32%

tracing to a few plants collected from Hebo Cemetery in Hebo, Oregon in 1996; 32%

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tracing to East border of creeping bentgrass research trial at Horticultural Research Farm

II, in New Brunswick, NJ in 1996. 23% tracing to a plant collected from a mountain

meadow in Eastern Europe in 1997; and 13% tracing to a few plants collected from Brae

Burn Country Club in West Newton, MA in 1997 (Stacy Bonos – personal

communication).

Capri A. capillaris Maternal progenies of 24 clones – 64% tracing to several plants selected from the

Connecticut Agricultural Experiment Station; 26% of the germplasm tracing to

segregating plants from the border area of a putting green research trial in North

Brunswick, NJ; and 10% tracing to plants collected from Hebo Cemetery in Hebo,

Oregon. (Stacy Bonos – personal communication)

EBM A. capillaris EBM colonial bentgrass was selected from the maternal progenies of 51 clones – 42%

tracing to a few plants received from the Connecticut Experiment Station in 1996; 34%

tracing to a few plants collected from Hebo Cemetery in Hebo, Oregon in 1996; 13%

tracing to a plant collected from Century CC in Purchase, NY in 1996; and 11% tracing

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to several plants collected from mountain meadows and old golf courses in Eastern

Europe in 1997 (Stacy Bonos – personal communication).

SR7100 A. capillaris Origin traces to 90 plants from the European colonial bentgrass ‘Bardot’ selected for

improved seed yield. (Seed Research of Oregon Inc 1994)

SR7150 A. capillaris Progeny of four selected plants tracing to plants selected from a PI collection planted at

USDA Farm, Beltsville, MD. (Leah Brilman – personal communication)

Tiger II A. capillaris Traces maternally to: 33.3% from the cultivar Bardot, and 66.7% from the cultivar Denso

(Steve Reid – personal communication)

CDE composite A. canina CDE velvet bentgrass was selected from the maternal progenies of 29 clones. Sixty-

seven percent of the maternal sources trace to several plants collected from Lake Success

Golf Club in Great Neck, NY in 1996. Thirty-three percent of the maternal sources trace

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to plants selected out of the 1985 velvet bentgrass test located at Hort Farm II, acquired

by Jerry Pepin (Stacy Bonos – personal communication).

SMM composite A. canina SMM velvet bentgrass was selected from the maternal progenies of 25 clones. Sixty-four

percent of the maternal sources trace to several plants collected from Lake Success Golf

Club in Great Neck, NY in 1996. Thirty-six percent of the maternal sources trace to

plants selected out of the 1985 velvet bentgrass test located at Hort Farm II, acquired by

Jerry Pepin (Stacy Bonos – personal communication).

SME composite A. canina SME velvet bentgrass was selected from the maternal progenies of 21 clones. Seventy-

one percent of the maternal sources trace to several plants collected from Lake Success

Golf Club in Great Neck, NY in 1996. Twenty-four percent of the maternal sources

traces to plants selected out of the 1985 velvet bentgrass test located at Hort Farm II,

acquired by Jerry Pepin. Five percent trace to plants collected from Lake Sunapee CC in

New London, NH in 2004 (Stacy Bonos – personal communication).

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VTP composite A. canina VTP velvet bentgrass was selected from the maternal progenies of 18 clones. Seventy-

eight percent trace to plants collected from Lake Sunapee CC in New London, NH in

2004. Eleven percent trace to plants collected from Winchester CC in Winchester, MA in

2004. Another 11% trace to plants collected from Pound Ridge Golf Club in Pound

Ridge, NY in 2004 (Stacy Bonos – personal communication).

Legendary A. canina Selected from the maternal progenies of nine clones. 20% of the germplasm traces to

selections from ‘Emanuel Francis’ velvet bentgrass acquired from Dr. Jerry Pepin; 50%

of the maternal sources trace to several plants collected from Lake Success Golf Club

(Great Neck, NY); 20% traces to plants collected from the National Golf Links of

America in Southhampton, NY; and 10% traces to a plant collected from Rock Ridge CC

in Newtown, CT. (Lebanon Seaboard Corporation & Rutgers, The State University of

New Jersey 2009b)

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Villa A. canina Material traces to germplasm acquired from Dr. Jerry Pepin, plants collected from Lake

Success Golf Club (Great Neck, NY), and plants collected from the National Golf Links

of America in Southhampton, NY. (DLF International Seeds 2012)

SR 7200 A. canina Traces back to 4 clones collected from New England area putting greens by Dr. Richard

Skogley (University of Rhode Island). (Seed Research of Oregon Inc 1998)

PI 203444 Apera interrupta Collection from Turkey

PI 204389 Apera interrupta Collection from Turkey

PI 372227 Polypogon chilensis Collection from Uruguay

PI 204395 Polypogon viridis Collection from Malatya, Turkey

a indicates putative Agrostis gigantea PI accessions originally labeled as Agrostis stolonifera by the USDA NPGS.

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References

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Brauen SE, Goss RL, Brede AD (1993) Registration of ‘Putter’ Creeping Bentgrass. Crop Sci 33:1100

Brede AD (2007) ‘Alpha’ and ‘T-1’ creeping bentgrass, new cultivars for golf. HortScience 42:1301-1302

Bonos SA, Huff DR (2013) Cool-Season Grasses: Biology and Breeding. In: Stier JC, Horgan BP, Bonos SA (eds) Turfgrass:

Biology, Use and Management. Agron Monogr 56. ASA, CSSA, and SSSA, Madison, WI. pp 591-660

DLF International Seeds (2012) Velvet bentgrass ‘Villa’. U.S. Plant Variety Protection Certificate 200800167. Date Issued: 12 March

2012

Hein MA (1958) Registration of varieties of strains of grasses. Bentgrasses: Penncross creeping (Reg No. 1). Agron J 50:399–401

Jacklin Seed Company (1991) Creeping bentgrass ‘Putter’. U.S. Plant Variety Protection Certificate 9000231. Date Issued: 30

September 1991

Krans JV, Philley HW, Goatley JM, Maddox VL (2000) Registration of twelve creeping bentgrass germplasms selected in

Mississippi. Crop Sci 40:582

Lebanon Seaboard Corporation, Rutgers the State University of New Jersey (2005) Creeping bentgrass ‘Independence’. U.S. Plant

Variety Protection Certificate 200300300. Date Issued: 26 July 2005

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Lebanon Seaboard Corporation, Rutgers the State University of New Jersey (2009a) Creeping bentgrass ‘Declaration’. U.S. Plant

Variety Protection Certificate 200500333. Date Issued: 16 July 2009

Lebanon Seaboard Corporation, Rutgers the State University of New Jersey (2009b) Velvet bentgrass ‘Legendary’. U.S. Plant Variety

Protection Certificate 200500332. Date Issued: 1 May 2009

Lofts Seed, Inc (1994) Creeping bentgrass ‘Southshore’. U.S. Plant Variety Protection Certificate 9200256. Date Issued: 31 October

1994

Musser HB (1959) Turf management: Grasses. USGA Journal and Turf Management 12:31-32

Novel Ag, Inc (2009) Creeping bentgrass ‘Authority’. U.S. Plant Variety Protection Certificate 200600044. Date Issued: 12 August

2009

Novel Ag, Inc, Rutgers the State University of New Jersey (2009) Creeping bentgrass ‘Shark’. U.S. Plant Variety Protection

Certificate 200600164. Date Issued: 12 August 2009

Peak Plant Genetics, LLC, Rutgers the State University of New Jersey (2013) Creeping bentgrass ‘Barracuda’. U.S. Plant Variety

Protection Certificate 201200052. Date Issued: 20 February 2013

Pennsylvania Agricultural Experiment Station (1980) Creeping bentgrass ‘Penneagle’. U.S. Plant Variety Protection Certificate

7900009. Date Issued: 20 November 1980

31

Pennsylvania Agricultural Experiment Station (1987) Creeping bentgrass ‘Pennlinks’. U.S. Plant Variety Protection Certificate

8700030. Date Issued: 30 June 1987

Pennsylvania Agricultural Experiment Station (2004) Creeping bentgrass ‘A-4’. U.S. Plant Variety Protection Certificate 9700060.

Date Issued: 22 November 2004

Pickseed West, Inc (1991) Creeping bentgrass ‘National’. U.S. Plant Variety Protection Certificate 8800225. Date Issued: 29

November 1991

Pure Seed Testing, Inc (2004) Colonial bentgrass ‘Alister’. U.S. Plant Variety Protection Certificate 200200090. Date Issued: 24

November 2004

Pure Seed Testing, Inc (2006) Creeping bentgrass ‘Penneagle II’. U.S. Plant Variety Protection Certificate 200500060. Date Issued:

14 February 2006

Rutgers the State University of New Jersey (2009a) Creeping bentgrass ‘13M’. U.S. Plant Variety Protection Certificate 200600280.

Date Issued: 16 July 2009

Rutgers the State University of New Jersey (2009b) Creeping bentgrass ‘Benchmark DSR’. U.S. Plant Variety Protection Certificate

200600166. Date Issued: 12 August 2009

Rutgers the State University of New Jersey, R.H. Hurley (2009) Creeping bentgrass ‘007’. U.S. Plant Variety Protection Certificate

200600263. Date Issued: 17 February 2009

32

Schoth HA (1930) Bentgrass seed production in the Pacific Northwest. USGA Green Section Record 10(11):206-211

Seed Research of Oregon, Inc (1994) Colonial bentgrass ‘SR 7100’. U.S. Plant Variety Protection Certificate 9400121. Date Issued:

30 November 1994

Seed Research of Oregon, Inc (1998) Velvet bentgrass ‘SR 7200’. U.S. Plant Variety Protection Certificate 9500303. Date Issued: 30

January 1998

Seed Research of Oregon, Inc (2004) Creeping bentgrass ‘SR 1119’. U.S. Plant Variety Protection Certificate 9700390. Date Issued:

10 September 2004

Texas Agricultural Experiment Station (1996) Creeping bentgrass ‘Crenshaw’. U.S. Plant Variety Protection Certificate 9400015.

Date Issued: 29 February 1996

Texas Agricultural Experiment Station (2004a) Creeping bentgrass ‘Century’. U.S. Plant Variety Protection Certificate 9800331.

Date Issued: 27 September 2004

Texas Agricultural Experiment Station (2004b) Creeping bentgrass ‘Imperial’. U.S. Plant Variety Protection Certificate 9800332.

Date Issued: 27 September 2004

University of Rhode Island (1991) Creeping bentgrass ‘Providence’. U.S. Plant Variety Protection Certificate 8900287. Date Issued:

28 June 1991

33

Warnke SE (2003) Creeping bentgrass (Agrostis stolonifera L.). In: Cassler MD, Duncan RR (eds) Turfgrass Biology, Genetics, and

Breeding. John Wiley and Sons, Hoboken, NJ pp 175-185

Classification of bentgrass (Agrostis) cultivars and accessions based on microsatellite (SSR) markers. Genetic Resources and Crop Evolution. Josh A. Honig*, Christine Kubik, Vincenzo Averello, Jennifer Vaiciunas, William A. Meyer, and Stacy A. Bonos. *Dep. of Plant Biology and Pathology, School of Environmental and Biological Sciences, Rutgers, The State University of New Jersey, 59 Dudley Rd., New Brunswick, NJ 08901-8520, [email protected]

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