photosynthesis the sun the ultimate energy source for life on planet earth
TRANSCRIPT
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Photosynthesis
The SunThe Ultimate Energy Source
For Life on Planet Earth
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Overview: The Process That Feeds the Biosphere
Photosynthesis is the process that converts solar energy into chemical energy
Directly or indirectly, photosynthesis nourishes almost the entire living world
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Pyramid of Biomass
The Sun
Photosynthesis (P/S)
Producers
Consumers
Autotrophs
Heterotrophs
Trophic Levels
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Autotrophs sustain themselves without eating anything derived from other organisms
Autotrophs are the producers of the biosphere, producing organic molecules from CO2 and other inorganic molecules
Almost all plants are photoautotrophs, using the energy of sunlight to make organic molecules
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Autotrophs
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Photosynthesis occurs in plants, algae, certain other protists, and some prokaryotes
These organisms feed not only themselves but also most of the living world
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(a) Plants(b)Multicellular
alga
(c)Unicellularprotists
(d) Cyanobacteria
(e)Purple sulfurbacteria
10 m
1 m
40 m
Figure 10.2
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Heterotrophs obtain their organic material from other organisms
Heterotrophs are the consumers of the biosphere
Almost all heterotrophs, including humans, depend on photoautotrophs for food and O2
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Heterotrophs
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The Earth’s supply of fossil fuels was formed from the remains of organisms that died hundreds of millions of years ago
In a sense, fossil fuels represent stores of solar energy from the distant past (“Ancient Sunlight”)
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Fossil Fuels
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Concept 10.1:
Photosynthesis converts light energy to the chemical energy of food
Chloroplasts are structurally similar to and likely evolved from photosynthetic bacteria
The structural organization of these cells allows for the chemical reactions of photosynthesis
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Photosynthesis (net consumption of water)
6CO2 + 6H2O C6H12O6 + 6O2Light
Chlorophyll
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Photosynthesis (tracking atoms)
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How Does CO2 Get In The Plant?
CO2 enters the ____________
What is the source of CO2? ______Air
78.08 %N2
20.95 %O2
0.03 %CO2argon, helium, hydrogen, ozone, methane
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How Does Water Get In The Plant?
Water enters through the __________
Most water is lost from the plant through the _________
To reduce water loss leaves are covered with a waxy cuticle (plant “chapstick”)
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Chloroplasts: The Sites of Photosynthesis in Plants
Leaves are the major locations of photosynthesis
Their green color is from chlorophyll, the green pigment within chloroplasts
Chloroplasts are found mainly in cells of the mesophyll, the interior tissue of the leaf
Each mesophyll cell contains 30–40 chloroplasts
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CO2 Enters
The Leaf is Covered with a Waxy Cuticle: “Plant Chapstick”
H2O Exits
If water cannot get out of the leaf through the waxycuticle what cannot getinto the leaf for P/S?
Mesophyll cell or photosynthetic cell: Note the chloroplasts
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CO2 enters and O2 exits the leaf through microscopic pores called stomata
The chlorophyll is in the membranes of thylakoids (connected sacs in the chloroplast); thylakoids may be stacked in columns called grana
Chloroplasts also contain stroma, a dense interior fluid
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H2OExits
CO2 Enters
Open Stomate
Closed Stomate
2 Guard Cellssurrounding apore
Transpiration=Evaporation of water from plant stomate
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Stomata On A Pine Needle
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Transpiration The evaporative water loss by a
plant, primarily through stomata
Degree of Stomatal Opening
Closed Partially FullyOpen Open
TranspirationRate
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Photosynthesis and Transpiration
High Rates of P/S are associated with high transpiration rates
Degree of Stomatal Opening
Closed Partially FullyOpen Open
Rate ofP/S
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Tracking Atoms Through Photosynthesis: Scientific Inquiry Photosynthesis is a complex series of
reactions that can be summarized as the following equation:
6 CO2 + 12 H2O + Light energy C6H12O6 + 6 O2 + 6 H2O
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The Splitting of Water
Chloroplasts split H2O into hydrogen and oxygen, incorporating the electrons of hydrogen into sugar molecules and releasing oxygen as a by-product
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Photosynthesis as a Redox Process
Photosynthesis reverses the direction of electron flow compared to respiration
Photosynthesis is a redox process in which H2O is oxidized and CO2 is reduced
Photosynthesis is an endergonic process; the energy boost is provided by light
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Figure 10.UN01
Energy 6 CO2 6 H2O C6 H12 O6 6 O2
becomes reduced
becomes oxidized
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The Two Stages of Photosynthesis: A Preview
Photosynthesis consists of the light reactions (the photo part) and Calvin cycle (the synthesis part)
The light reactions (in the thylakoids) Split H2O Release O2
Reduce NADP+ to NADPH Generate ATP from ADP by
photophosphorylation
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The Calvin cycle (in the stroma) forms sugar from CO2, using ATP and NADPH
The Calvin cycle begins with carbon fixation, incorporating CO2 into organic molecules
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Concept 10.2: The light reactions convert solar energy to the chemical energy of ATP and NADPH
Chloroplasts are solar-powered chemical factories
Their thylakoids transform light energy into the chemical energy of ATP and NADPH
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Structure of a Chloroplast
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Chlorophyll
Chlorophyll is a green pigment Pigments absorb light energy The color you see is the color that is
reflected White versus Black
Why is chlorophyll green? Would you expect green light to be an
effective color of light for P/S?
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The electromagnetic spectrum is the entire range of electromagnetic energy, or radiation
Visible light consists of wavelengths (including those that drive photosynthesis) that produce colors we can see
Light also behaves as though it consists of discrete particles, called photons
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Light: Energy is Inversely Proportional to Wavelength
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Photosynthetic Pigments: The Light Receptors
Pigments are substances that absorb visible light
Different pigments absorb different wavelengths Wavelengths that are not absorbed are reflected
or transmitted Leaves appear green because chlorophyll
reflects and transmits green light
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Chlorophyll Reflects and Transmits Green Light
Chlorophyll absorbs blue and red light
Accessory pigments such as carotenoids absorb green light and pass the energy to chlorophyll
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A spectrophotometer measures a pigment’s ability to absorb various wavelengths
This machine sends light through pigments and measures the fraction of light transmitted at each wavelength
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Absorption Spectra
An absorption spectrum is a graph plotting a pigment’s light absorption versus wavelength
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The absorption spectrum of chlorophyll a suggests that violet-blue and red
light work best for photosynthesis
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Photosynthesis is a Two Step Process
6CO2 + 6H2O C6H12O6 + 6O2
The Light Reactions: Light Dependant Photochemical
Convert light energy into chemical energy
The Light Independent Temperature Dependant Use the chemical energy created in the
light reactions to convert CO2 to glucose
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Photosynthesis Overview(2 step process)
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Locations of Reactions The Light
Dependent Reactions occur on the grana
The Light Independent Reactions take place in the stroma
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High Potential Energy Molecules
Is this an example of oxidation or reduction?(oxidation = loss of e-, reduction = gain of e-)
Does the formation of NADPH require or release energy?
NADP+ + 2e- (2H) NADPH + H+
P Pe-e-
NADPH + H+ NADP+ + 2e-
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High Potential Energy Molecules
Does the formation of NADPH require or release energy?
NADP+ + 2e- (2H) NADPH + H+
NADPH + H+ NADP+ + 2e-
Reduction (endergonic)
Oxidation (exergonic)
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High Potential Energy Molecules
ATP: Adenosine Triphosphate - Universal Energy Molecule
Adenosine P P P ---------- Adenosine P P + P + E
ATP ---------- ADP + P + Energy
ADP + P + Energy ------------------- ATP
Adenosine Triphosphate Adenosine Diphosphate + P + Energy
High P.E. Low P.E.
Low P.E. High P.E.
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The Light Reactions Photochemical: Light energy is
converted to chemical energy in the form of two high potential energy molecules.
ADP + P ATP
H2O ------------------------------------- ½ O2
NADP + 2e- NADPH
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The Light Reactions The Two High Potential Energy
Molecules Produced are: __________ __________
The Electron Source is?__________ When water gives up electrons
what waste product is produced?
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Excitation of Chlorophyll by Light
When a pigment absorbs light, it goes from a ground state to an excited state, which is unstable
When excited electrons fall back to the ground state, photons are given off, an afterglow called fluorescence
If illuminated, an isolated solution of chlorophyll will fluoresce, giving off light and heat
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Figure 10.12
Excitedstate
Heat
e
Photon(fluorescence)
Groundstate
PhotonChlorophyllmolecule
En
erg
y o
f ele
ctr
on
(a) Excitation of isolated chlorophyll molecule(b) Fluorescence
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A Photosystem: A Reaction-Center Complex Associated with Light-Harvesting Complexes
A photosystem consists of a reaction-center complex (a type of protein complex) surrounded by light-harvesting complexes
The light-harvesting complexes (pigment molecules bound to proteins) transfer the energy of photons to the reaction center
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Figure 10.13a
(a) How a photosystem harvests light
Th
yla
koid
mem
bra
ne
PhotonPhotosystem STROMA
Light-harvestingcomplexes
Reaction-centercomplex
Primaryelectronacceptor
Transferof energy
Special pair ofchlorophyll amolecules
Pigmentmolecules
THYLAKOID SPACE(INTERIOR OF THYLAKOID)
e
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• A primary electron acceptor in the reaction center accepts excited electrons and is reduced as a result
Solar-powered transfer of an electron from a chlorophyll a molecule to the primary electron acceptor is the first step of the light reactions
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• There are two types of photosystems in the thylakoid membrane
• Photosystem II (PS II) functions first (the numbers reflect order of discovery) and is best at absorbing a wavelength of 680 nm
• The reaction-center chlorophyll a of PS II is called P680
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• Photosystem I (PS I) is best at absorbing a wavelength of 700 nm
• The reaction-center chlorophyll a of PS I is called P700
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Noncyclic (Linear) Electron Flow
During the light reactions, there are two possible routes for electron flow: cyclic and linear
Noncyclic (linear) electron flow, the primary pathway, involves both photosystems and produces ATP and NADPH using light energy
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The Model For The Light ReactionsNoncyclic Electron Flow
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A photon hits a pigment and its energy is passed among pigment molecules until it excites P680
An excited electron from P680 is transferred to the primary electron acceptor (we now call it P680+)
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Each electron “falls” down an electron transport chain from the primary electron acceptor of PS II to PS I
Energy released by the fall drives the creation of a proton gradient across the thylakoid membrane
Diffusion of H+ (protons) across the membrane drives ATP synthesis
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In PS I (like PS II), transferred light energy excites P700, which loses an electron to an electron acceptor
P700+ (P700 that is missing an electron) accepts an electron passed down from PS II via the electron transport chain
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Each electron “falls” down an electron transport chain from the primary electron acceptor of PS I to the protein ferredoxin (Fd)
The electrons are then transferred to NADP+ and reduce it to NADPH
The electrons of NADPH are available for the reactions of the Calvin cycle
This process also removes an H+ from the stroma
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The Model For The Light ReactionsCyclic Electron Flow
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Cyclic Electron Flow
• Cyclic electron flow uses only photosystem I and produces ATP, but not NADPH
• No oxygen is released Cyclic electron flow generates surplus ATP,
satisfying the higher demand in the Calvin cycle
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The Light Reactions are Light Dependent
Light Intensity
Rate ofP/S
Light Saturation
Low Med High
Can you think of a habitator ecosystem where P/S might be limited?
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The Light Independent Reactions
The Light Independent Reactions or the “Dark Reactions”
ATP ADP + P
6CO2 ---------------------------------------------------- C6H12O6
NADPH2 NADP + 2e-
What is the “fuel” that drives the light independent reactions?__________
Where was this “fuel” produced?_______________________
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ATP and NADPH are produced on the side facing the stroma, where the Calvin cycle takes place
In summary, light reactions generate ATP and increase the potential energy of electrons by moving them from H2O to NADPH
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Figure 10.18
STROMA(low H concentration)
STROMA(low H concentration)
THYLAKOID SPACE(high H concentration)
Light
Photosystem II
Cytochromecomplex Photosystem I
Light
NADP
reductase
NADP + H
ToCalvinCycle
ATPsynthase
Thylakoidmembrane
2
1
3
NADPH
Fd
Pc
Pq
4 H+
4 H++2 H+
H+
ADP+P i
ATP
1/2
H2OO2
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Concept 10.3: The Calvin cycle uses the chemical energy of ATP and NADPH to reduce CO2 to sugar
The Calvin cycle regenerates its starting material after molecules enter and leave the cycle
The cycle builds sugar from smaller molecules by using ATP and the reducing power of electrons carried by NADPH
Carbon enters the cycle as CO2 and leaves as a sugar named glyceraldehyde 3-phospate (G3P)
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Calvin Cycle (Step One)
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Input
3 (Entering oneat a time)
CO2
Phase 1: Carbon fixation
Rubisco
3 P P
P6
Short-livedintermediate
3-Phosphoglycerate6
6 ADP
ATP
6 P P1,3-Bisphosphoglycerate
CalvinCycle
6 NADPH
6 NADP
6 P i
6 PPhase 2: Reduction
Glyceraldehyde 3-phosphate(G3P)
3P PRibulose bisphosphate
(RuBP)
1 PG3P
(a sugar)Output
Glucose andother organiccompounds
Figure 10.19-2
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Concept 10.4: Alternative mechanisms of carbon fixation have evolved in hot, arid climates
Dehydration is a problem for plants, sometimes requiring trade-offs with other metabolic processes, especially photosynthesis
On hot, dry days, plants close stomata, which conserves H2O but also limits photosynthesis
The closing of stomata reduces access to CO2
and causes O2 to build up These conditions favor an apparently wasteful
process called photorespiration© 2011 Pearson Education, Inc.
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Photorespiration: An Evolutionary Relic?
• In most plants (C3 plants), initial fixation of CO2, via rubisco, forms a three-carbon compound (3-phosphoglycerate)
• In photorespiration, rubisco adds O2 instead of CO2 in the Calvin cycle, producing a two-carbon compound
Photorespiration consumes O2 and organic fuel and releases CO2 without producing ATP or sugar
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• Photorespiration may be an evolutionary relic because rubisco first evolved at a time when the atmosphere had far less O2 and more CO2
• Photorespiration limits damaging products of light reactions that build up in the absence of the Calvin cycle
In many plants, photorespiration is a problem because on a hot, dry day it can drain as much as 50% of the carbon fixed by the Calvin cycle
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C4 Plants
C4 plants minimize the cost of photorespiration by incorporating CO2 into four-carbon compounds in mesophyll cells
This step requires the enzyme PEP carboxylase PEP carboxylase has a higher affinity for CO2 than
rubisco does; it can fix CO2 even when CO2 concentrations are low
These four-carbon compounds are exported to bundle-sheath cells, where they release CO2 that is then used in the Calvin cycle
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Saltbush
Saltbushcan live in salty soils sinceit has salt glands
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Saltbush – A C4 Plant
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Kranz Anatomy
Note that thechloroplastsare in thecenter of theleaf
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CAM Plants
Some plants, including succulents, use crassulacean acid metabolism (CAM) to fix carbon
CAM plants open their stomata at night, incorporating CO2 into organic acids
Stomata close during the day, and CO2 is released from organic acids and used in the Calvin cycle
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Cactus – A Stem Succulent
GreenStem
CAM P/s
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The Importance of Photosynthesis: A Review
The energy entering chloroplasts as sunlight gets stored as chemical energy in organic compounds
Sugar made in the chloroplasts supplies chemical energy and carbon skeletons to synthesize the organic molecules of cells
Plants store excess sugar as starch in structures such as roots, tubers, seeds, and fruits
In addition to food production, photosynthesis produces the O2 in our atmosphere
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Mitochondrion Chloroplast
MITOCHONDRIONSTRUCTURE
CHLOROPLASTSTRUCTURE
Intermembranespace
Innermembrane
Matrix
Thylakoidspace
Thylakoidmembrane
Stroma
Electrontransport
chain
H Diffusion
ATPsynthase
H
ADP P iKey Higher [H ]
Lower [H ]
ATP
Figure 10.17
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A Comparison of Chemiosmosis in Chloroplasts and Mitochondria
Chloroplasts and mitochondria generate ATP by chemiosmosis, but use different sources of energy
Mitochondria transfer chemical energy from food to ATP; chloroplasts transform light energy into the chemical energy of ATP
Spatial organization of chemiosmosis differs between chloroplasts and mitochondria but also shows similarities
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In mitochondria, protons are pumped to the intermembrane space and drive ATP synthesis as they diffuse back into the mitochondrial matrix
In chloroplasts, protons are pumped into the thylakoid space and drive ATP synthesis as they diffuse back into the stroma
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In the last 150 years since the Industrial Revolution, CO2 levels have risen greatly
Increasing levels of CO2 may affect C3 and C4 plants differently, perhaps changing the relative abundance of these species
The effects of such changes are unpredictable and a cause for concern
350.org, davidsuzuki.org
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Climate Change