new stiliderus motschulsky species from the indo-ausíralia...
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ANNALES HISTORICO-NATURALES MUSEI NATION ALI S HUNGARIC1 Tomus 78. Budapest, 1986 p. 79-82.
New Stiliderus Motschulsky species from the Indo-Ausíralian Region
(Coleoptera, Staphylinidae)*
by G . M . D E R O U G E M O N T , L o n d o n
Abstract— A list is given o f material studied, including new records. The aedeagus of S. kiloloensis ( L A S T ) is figured for the first t ime; S. loksai sp. n. . S. wauensis sp. n . , S. maritimus sp. n . and 5. madangensis sp. n. are described, and a plate o f figures is given o f the aedeagi o f the nine species now known belonging to the New Guinea hieroglyphicus group. W i t h 10 figures.
T h a n k s t o the kindness o f D R . O . M E R K L I have been able t o examine m a t e r i a l o f the genus Stiliderus (sensu l a t o : B L A C K W E L D E R 1939, & R O U G E M O N T ) i n the col lec t ions o f the H u n g a r i a n N a t u r a l H i s t o r y M u s e u m . T h e study has revealed three new species, a n d a f o u r t h was p r o v i d e d by m y f r i end M R . H . L A S T , w h o recognised i t ' s o r i g i n a l i t y a n d i n v i t e d m e t o describe i t . I a m gra te fu l t o b o t h these colleagues, as also t o the staff o f the B r i t i s h M u s e u m , N a t u r a l H i s t o r y , f o r a l l o w i n g frequent access t o the col lec t ions .
S Y S T E M A T I C S
Stiliderus kiloloensis ( L A S T )
Stilicoderus kiloloensis L A S T , 1984, Annls hist.-nat. Mus. natn. hung. 76: 137. 1 was unaware of the existence of this species which was described after the final draft o f the
manuscript o f my revision, part 1. I n accordance wi th the criteria used in my revision & key, a short supplement to the author's description and a figure of the aedeagus are given below.
The th i rd known member o f the funebris group, to which i t runs at the beginning o f my key. I t differs f rom bo th the other species, S. funebris L A S T and S. nubilus L A S T at first sight by the b i -colorous elytra and indistinctly paler terminal segments o f the abdomen. I t differs furthermore by the slightly asperate micro-puncturation o f the elytra (simple in the other two species). The median tooth of the labrum is equal in length but stouter than the lateral pair. The aedeagus is characteristic, bu t typical i n structure, of the group, w i th a very large wish-bone shaped dorsal plate o f the median lobe.
Proportions o f holotype: length: ca. 9.5 m m . ; length o f head: 128; breadth o f head: 130; length o f antennae: 268; length o f p ronotum: 132; breadth of p rono tum: 125; length o f elytra: 146; breadth o f elytra: 150.
Stiliderus aerosus ( L A S T )
Stilicoderus aerosus L A S T , 1984, Fo l . ent. hung. 45 (2): 114. Stiliderus aerosus: R O U G E M O N (1986) Ent. A b h . Mus . Tierk. Dresden 49 (8): 154.
2 cfcf & 1 $ : New Guinea / N E , Wau, M c A d a m Park I X . 1968 / no. N G - W - B 38 / leg. D r . J.. Balogh; 1 9 : New Guinea / N E , Wau 22. I X . - 3 0 . I X . 1969 / N o . N G W - R 9 : leg. D r . J . Balogh.
These exx. agree wi th the S. aerosus holotype which I have seen, including the long recumbent pubescence and the aedeagus, but lack the aeneous lustre of the fore-body. The new material comes f rom a locality far to the SE o f that o f the holotype material.
* 22nd contribution to the knowledge of Staphylinidae.
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Stiliderus discalis ( F A U V E L )
Stilicoderus discalis F A U V E L , 1895, Rev. d 'Ent 14: 225. .Stiliderus discalis: R O U G E M O N T (1986) Ent. A b h . Mus . Tierk. Dresden 49(8): 163.
1 Ç : Vietnam, Cuc-phù6ng B ô n g jungle, extracted f rom li t ter 12-15. X I I . 1965 leg. T . P ó c s ; 1 rf & 1 9 : ib id . , extracted f rom roots o f Asplenium nidus 15. X I I . 1965, leg. T . P ó c s .
This is the first member o f the genus recorded f rom Vietnam. The species is k n o w n f rom Burma •and Thailand.
Stiliderus nepalensis R O U G E M O N T
Stiliderus nepalensis R O U G E M O N T (1986) Ent. A b h . Mus . Tierk . Dresden 49(8): 172. Stiliderus nepalensis R O U G E M O N T (1986) Rev. Suisse de Zool . 93(1): 233
1 $ : India, W. Bengal, Darjeeling Dis t r . on road to Lebong. leg. Gy. T o p á i . N o . 311 sifted l i t ter , mosses 11. I V . 1967.
This species is known wi th certainty f rom two localities i n Nepal , and possibly (1 9 ) from Sikkim. Only the eventual discovery o f males w i l l prove that this ex. belongs to S. nepalensis and n o t to the variable S. kambaitiensis SCHEERPELZ.
* * *
The fol lowing five taxa belong to the New Guinea hieroglyphicus-group, comprising species which are so similar that they may only be determined by examination o f the aedeagus. A t least some o f these species are sympatric, so that females among material f rom the same local i ty containing more than one species are for the present indeterminable, and are labelled by me "Stiliderus sp. hieroglyphicus gr.". Because I can find no other characters for the seperation o f these species, I have, as i n a previous paper (ROUGEMONT, i n press Reichenbachia, Mus . Tierk. Dresden) l imi ted descriptions to the proportions of holotypes; i n a l l other respects the characters may be assumed to be those o f S. hieroglyphicus F A U V E L as redescribed i n my revision ( R O U G E M O N T 1986). T o facilitate determinations wi thout recourse to other papers, I give below a plate o f figures o f the aedeagi of a l l the species known so far.
Stiliderus loksai sp. n .
rf holotype & 4 çfçf paratypes : New Guinea / N E , Wau, K i l o l o Creek, 26. V I I I . 68 / N o . N G - W :R.7 / leg. D r . I . Loksa; 2 çfçf Paratypes: New Guinea / N E , Wau 8. I X . 68 / N o . N G - W R.19 / leg. D r . I . Loksa.
Proportions o f holotype: length: 4.6 m m ; length o f head: 63; breadth o f head: 76; length o f antennae: 125; length o f p rono tum: 66; breadth o f p rono tum: 60; length o f e lytra: 77; breadth o f elytra: 74.
This species is best characterised by the shape of the parameroid lobes, which resemble those o f 5. hornabrooki R O U G E M O N T , but are different i n outline (see Figs. 6 & 9). The paramere is also similar to that species, but is not quite so broad ( in ventral view), and is longer (not reaching beyond the apices of parameroid lobes i n S. hornabrooki), it 's apex recurved as i n S. hieroglyphicus F A U V E L , 5. leai R O U G E M O N T and others.
Stiliderus wauensis sp. n .
çf holotype: New Guinea / N E / Wau, K i l o l o Creek, 26. V I I I . 1968 / N o . N G - W R. 7 / leg. D r . I . Loksa.
Proportions of holotype: length: 5 m m ; length o f head: 67; breadth o f head: 72; length o f antennae: 126; length o f p rono tum: 66; breadth o f p rono tum: 62; length of e lytra: 82; breadth o f elytra: 80.
The broad subtriangular ( in lateral view) shape of the parameroid lobes is unl ike that o f any other member o f the hieroglyphicus group except the fol lowing species. The paramere is fairly broad, laterally compressed, the apex scarcely reaching beyond the parameroid lobes, not dilated or recurved (fig. 4).
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Stiliderus maritimus sp. n .
rf holotype: New Guinea / SE / Port Moresby, R igo , 18. V I I I . 1968 / N o . NG-P R 2 / leg. D r . I . Loksa.
Proportions o f holotype: length: 4.8 m m . ; length of head: 68; breadth o f head: 73; length o f antennae: 126; length o f p rono tum: 64; breadth o f p rono tum: 60; length o f e ly t ra : 78; breadth o f e ly t ra : 73.
The aedeagus of this species is proportionately smaller and more elongate as i t is also in S. sharpi R O U G E M O N T . A further resemblance of these two species lies i n the internal structures of the median lobe, but the paramere o f the new species is longer, it 's apex laterally compressed, and the parameroid lobes are o f quite another shape, resembling those of the preceeding new species, Aedeagus: F ig . 5.
Stiliderus madangensis sp. n .
cf holotype: T 779. Madang N . G . 22.12.69 (in coll . Last) This new species is the only one o f the hieroglyphicus group to be clearly differentiated on external
characters as wel l as by those o f the aedeagus. Proportions o f holotype: length: 5.6 m m . ; length of head: 73; breadth o f head: 83; length o f
antennae: 154; length o f p rono tum: 72; breadth o f p rono tum: 68; length o f e lyt ra : 86; breadth o f elytra: 82.
This species differs f rom a l l other members o f the hieroglyphicus group by i t ' s greater size, black elytra (brown in the other spp.) and by the puncturation o f the elytra, which is finer and denser;
Figs 1-9 Aedeagi in right lateral and ventral views: 1 = Stiliderus hieroglyphicus F A U V E L , 2 = 5 madangensis sp. n . , 3 = 5 . leai R O U G E M O N T , 4 = 5 . wauensis sp. n . , 5 = 5 maritimus sp. n , 6 = S,loksaisp.n.,7 = 5. sharpi R O U G E M O N T , 8 = 5 wallacei R O U G E M O N T , 9 = 5 hornabrooki^ R O U G E M O N T . — Fig . 10. Aedeagus, i n right lateral view of 5. kiloloensis L A S T . — Scale lines represent 1 m m .
Smaller scale line for 5. hornabrooki ROUGEMONT, larger l ine for a l l other species.
6 Term. Tud. Múz. Évk. 1986.
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although the granules are smaller, they are more prominent, i n sharper contrast to the shiny interstices.
The insect's greater size wou ld make i t run i n m y key (Revision) to members of the aerosus group, bu t the strongly transverse head and the conformation o f the aedeagus (Fig . 2) clearly show it 's true affinity. The very long parameroid lobes are only comparable wi th those o f S. hieroglyphicus F A U V E L (Fig . 1) but are o f a different shape. I n ventral view the laterally compressed blades o f the parameroid lobes appear together wi th the acicular paramere to fo rm a single long, pointed structure.
Stiliderus sp. hieroglyphicus group
1 cf: New Guinea / N E / Baiyer River Sanctuary, 1-5. I X . 1969 / N o . N G - B 85 / leg. Dr . J. Balogh.
This is apparently also a new species, but cannot be described because the specimen is immature, and the characters o f the aedeagus are deformed and unclear.
Stiliderus spp. $ $ (hieroglyphicus group)
1 ex.: New Guinea / N E / Wau , K i l o l o Creek, valley 900 m., 23. V I I I . 1968 / N o . N G - W - B 15 / leg. D r . J. Balogh; 4 exx.: Ib id , (wi thou t "valley 900 m . " ) ; 26. V I I I . 1968 / N o . N G - W . ?.7 / leg. D r . I . Loksa; 1 ex. : New Guinea / N E Wau, M t . K a i n d i , 10, I X . 1968 / N o . N G - W - B 94, leg. D r . J. Balogh; 1 ex. : New Guinea / N E / W a u , 8. I X . 1968 / N o . N G - W . ? 19, leg. D r . I . Loksa.
Many of these are probably S. loksai sp. n. , bu t must remain for the present undetermined.
The last species represented i n the material belongs to the cicatricosus group, to be studied in the second part o f m y revision.
Stiliderus crassus ( K R A A T Z )
Psilotrachellus crassus K R A A T Z , 1859, Arch . Naturg . 25: 124. 1 9 : Indo-China. 5. crassus was described f r o m Ceylon, and is cited from several countries in eastern Asia by
C A M E R O N , B E R N H A U E R and by K R A A T Z himself I have yet to confirm, by examination o f the holotype, whether the Far Eastern exx. belong to the same taxon as the Ceylanese holotype. I know the taxon to which the above specimen belongs w i t h certainty f rom Thai land, Singapore and China. I t is new to any o f the three countries which formerly constituted French Tndochina'.
References
L A S T , H . (1984) : Recorded and new species o f Coleoptera (Staphylinidae, Paederinae) i n Papua New Guinea. — Fol. ent. hung. 4 2 : 109-125.
L A S T , H . (1984): Further new species o f New Guinea Staphylinidae (Coleoptera) i n the collections of Hungarian and British Museums. — Annls hist.-nat. Mus. nat. hung. 7 6 : 133-138.
R O U G E M O N T , G . D E (1986): Revision o f the genus Stiliderus Motschulsky 1958. Part I ( = Stilicoderus Sharp). (Coleoptera, Staphylinidae). 14th contr ibut ion to the knowledge o f Staphylinidae. — Ent. Abh. Mus. Tierk. Dresden 4 9 (8): 139-187.
R O U G E M O N T , G . D E (1986): N e w data on the genus Stiliderus (Coleoptera, Staphylinidae). 18th contr ibut ion to the knowledge o f Staphylinidae. — Reichenbachia, Mus. Tierk. Dresden
R O U G E M O N T , G . D E (in press) : N e w records o f Stiliderus f rom Nepal (Coleoptera, Staphylinidae). 21st contr ibut ion to the knowledge o f Staphylinidae. — Rev. Suisse de Zool. 9 3 (1) : 233-236.
Author's address: GUILLAUME M . DE ROUGEMONT
27 Walcot Square London SE11 Great Britain