HUNDRED OFAYERS

HUNDRED OFSTRANGWAYS

HUNDREDOF BAGOT

HUNDREDOF GUY

HUNDRED OFHUTCHISON

HUNDREDOF PATON

H o lm e sJun g le

N a tu re P a rk

B lack Jungle /Lam bells Lagoon

ConservationReserve

Territory W ildlifePark / Be rry

Springs Nature Park

Vernon Is landsC onserva tion

R eserve

D jukb in jN ational

Park

C asuarinaC oasta lR eserve M elacca Sw am p

C onserva tion A rea

Fogg DamConservation

Reserve

H ow ard SpringsH unting R eserve

H o w a rd S p r ing sN a tu re P a rk

C har les D arw inN ational Park

Shoa l BayC oasta lR eserve

H arrison D amC onserva tion A rea

1

2a 2b

4

5

2

3

6

2c

Marrakai C reek

B lackmoreRiv erDarw in River

Berry Cr e ek

Adelaide R iv er

H o ward R iv er

Eliza b e th River

Strangways Rd

Stuart

Hw

y

Kentish Rd

Girraween Rd

Lowther Rd Produ

ce Rd

Gulnare RdFin

n Rd

Townend Rd

Freds P assRd

Hillier

Rd

Howard Springs R

d

Whitewood Rd

Morga

nRd

Goode Rd

Arnhem Hwy

Hicks Rd

Pioneer Dr

Redcl

iffe Rd

Lambe

lls Lag

oon R

d

Wand

errie R

dCox Peninsula Rd

Virgin

ia Rd

Freds Pass Rd

Old Bynoe Rd

Alverly Rd

Cox Peninsula Rd

Thorng

ate Rd

Livingstone Rd

Powe

r Rd

Elizabeth Valley Rd

B roo

king C

ctHo

pewe

ll Rd

Alpha

tonia R

d

Jenkins Rd

Ewart Rd Anzac

Pde

Fog Bay R d

Litchfie

ldPar

k Rd

Bees

Creek

Rd

Tr ipp

eRd

Channel Island Rd

Mocat

to Rd

Wickham

Point Rd

Gunn Point Rd

Gunn Point Rd

NT Por 2012LIMILNGAN-WULNA (LAND HOLDING)

ABORIGINAL CORPORATION

NT Por 2017LIMILNGAN-WULNA (LAND HOLDING)

ABORIGINAL CORPORATION

NT Por 4477KOOLPINYAH

NT Por 1299WOOLNER

NT Por 4476KOOLPINYAH

Palmerston

Darwin

Sec. 149

Sec. 8

Sec. 1624

5182

NT Por 7224

Sec. 628

Sec. 1529

Sec. 150

Sec. 1559

Sec. 4570

NT Por 2008

NT Por 2301

NT Por 3601

NT Por 6166

NT Por 5911

Por. 1263

Sec. 62 Sec. 63

Sec. 93

Sec. 104Sec. 107

Sec. 109

Sec. 64

Sec. 91Sec. 92

Sec. 4226

Sec. 1821

Por. 2270

Sec. 1823Sec. 1826

Sec. 2530Sec. 631

Sec. 633Sec. 655

Por. 2727

Sec. 146Sec. 147Sec. 151

Sec. 233

Sec. 239 Sec. 241

5775

Sec. 2746

Sec. 10

Sec. 1554

Sec. 1538

Sec. 1563

Sec. 1580Sec. 1580

Sec. 1580

Sec. 2743

Sec. 1597

Sec. 1603Sec. 1603

9375

4251

Sec. 1

Sec. 3

Sec. 4572

Sec. 4578

Sec. 4131

NT Por 4419

Sec. 5730

NT Por 2009

Sec. 1877

NT Por 6134

Sec. 1820

Sec. 105

Sec. 237

Sec. 5623Por. 2821

Sec. 113

NT Por 7045

Sec. 1892

11482

Sec. 1902

Sec. 5642

NT Por 7263

Sec. 5758

Sec. 7219

3952

Sec. 145

Sec. 1510

Sec. 1786Sec. 1561

Sec. 4076

Sec. 643

NT Por 2626

Sec. 238

Sec. 4225

Sec. 1828

Sec. 1840

NT Por 2010

NT Por 5998

Por. 2260

Sec. 1574

Sec. 144

Sec. 234

Sec. 90

Por. 1409

4254

7003

1851

4

Sec. 4048 Sec. 5761

Sec. 5543

Sec. 642

Sec. 1539

Sec. 1897

700000

700000

710000

710000

720000

720000

730000

730000

740000

740000

750000

750000

85900

00

85900

00

86000

00

86000

00

86100

00

86100

00

86200

00

86200

00

86300

00

86300

00

86400

00

86400

00

86500

00

86500

00

86600

00

86600

00

Proje ction: Unive rsal Transverse Me r cator (UTM)Map Grid of Australia (MGA), Z one 52

Horizontal Datum : GDA94(Inset only - V e rtical Datum: AHD66)

0 2 4 6 81km km

Threatened Species Distributionin the Greater Darwin Region

HUNDREDOF GUY

HUNDRED OFHUTCHISON

Black Jung le /Lam be lls Lagoon

C onserva tion R eserve

2a

2b

2a

2a

2

2a

2

2 2

2c

3530

20

20

2015

20

15

2015

10

5

10

30

30

30

5

25

25

25

25

Sec. 8

Sec. 1611

Sec. 568

Sec. 1683

Sec. 569

Sec. 1724

Sec. 1688

Sec. 1673Sec. 1580

Sec. 1660

Sec. 1672Sec. 1610

Sec. 1609

Sec. 1767

Sec. 1764

Sec. 1765

Sec. 572

Sec. 1659

Sec. 1661

NT Por 4476

738000

738000

739000

739000

740000

740000

741000

741000

742000

742000

743000

743000

744000

744000

86100

00

86100

00

86110

00

86110

00

86120

00

86120

00

86130

00

86130

00

86140

00

86140

00

86150

00

86150

00

86160

00

86160

00

86170

00

86170

00

86180

00

86180

00

HUNDRED OFHUTCHISON

HUNDREDOF BAGOT

H ow ard SpringsN atu re Park

33

3

Ho ward Rive r

35

15

10

40

3530

20

30

40

35

30

20

20

1510

30

3030

30

15

35

20

20

15 15

15

35

25

25

25

25

25

25

25

25

25

25

2525

25

Bronzewing Av

Mcgill Rd

Whitewood Rd

Azure Ct

Hicks Rd

Cornel

ius Cc

t

Currawong Dr

Deborah Rd

Wetherby RdAplin Rd

Rogers

Rd

Hillier

Rd

Sandpiper Gr

Littlejohn Rd

Mcintyre Rd

Goy Rd

Sophi a Rd

Barker Rd

Finlay Rd

Callist

emon

Rd

Makhara Rd

Gunn

Point Rd

Dougall Ct

Wagtail Ct

Ninnis Ct

Tobin Rd

Corella Av

Jason

Rd

Bastin

Rd

Price Rd

Melale

uca Rd

Parakeet Pl

Macleod Rd

Stanley Rd

Mahaf

fey Rd

Madsen Rd

Dicho

ndra

Rd

Watling Rd

Sittel la

Rd

Comp

igne R

d

G allac

herR

d

5453

76

Sec. 265

NT Por 3601Sec. 4296

Sec. 266Sec. 267

Sec. 269

65

Sec. 5349

Sec. 5585

Sec. 4116

Sec. 4116

Sec. 4449

56

Sec. 246Sec. 247

75

NT Por 6134

55

90

Por. 2821

Sec. 5077

Sec. 6791

Sec. 6791

Sec. 7085Sec. 7092

49

Sec. 4287Por. 2822

66

Sec. 5588

Sec. 268

Sec. 244

Por. 1409

Sec. 6790

Sec. 5586

Sec. 5019

Sec. 245

724000

724000

726000

726000

728000

728000

730000

730000

86160

00

86160

00

86180

00

86180

00

86200

00

86200

00

86220

00

86220

00

Inset - Intensive Survey Area: Howard River Catchment

Inset - Intensive Survey Area: Adelaide River Catchment

0 10.5km km

0 21km km

Ptychosperma macarthurii

Known Extentof Occurrence

Map com p ile d : 6/07/2016Caroline Gre e n and N icholas Cuff, De p artm ent of Land Resour c e Managem e nt

0 20 40km kmBroad Mapping Reliability

C O N T E N T S Page 1 of this document contains an Interactive PDF Map.

Use Ad ob e Read e r and op en the left p anel to r eview ind ivid ual map laye rs. Use rs ar e encourage d to hid e/show laye rs to find out m or e ab out this sp e cies d istrib ution,

p articularly in the inset map s, whe r e som e laye rs may mask laye rs und e r neath. Scroll to Page 2 to read the Guidelines for Map Use.

U S E O F M A P

Map s of Thr eatene d Sp e cies Distrib utions in the Gr eate r Darwin Ar ea should b e inte r p r ete d with the attache d Guidelines for Map Use. Scroll to page 2.

This map p r ovid es the m ost up to d ate availab le inform ation r egar d ing the known d istrib ution and extent of the sp e cies at the tim e of p ub lication.

Refe r to the N T GOV we b site to view Thr eatene d Sp e cies Inform ation She ets. Refe r to http ://eflora.nt.gov.au our online r esour ce for N orthe r n Te r ritory's flora.

For furthe r infor m ation, p lease contact;Flora and Fauna DivisionDepartment of Land Resource Management (DLRM)CSIRO Com p le x, 564 V and e r lin Drive, Be r rim ah N T 0828 W e b : http s://land r esour c es.nt.gov.au/

Cr eative Com m ons 4.0 Inte r national Lice nse http s://c r eativecom m ons.or g/lic e nses/b y-nc-sa/4.0/

Attrib ution-N onCom m e r cial-Shar eAlike

This p r od uct and all m ate rial form ing p art of it is cop yright b e longing to the N orthe r nTe r ritor y of Australia. Y ou m ay use thismate rial for your p e rsonal, non-com m e r cial use or use it within your or ganisation for non-com m e r cial p ur p oses, p r ovid e d thatan ap p r op riate acknowle d ge m e nt is mad e and the mate rial is not alte r e d in any way. Sub je ct to the fair d ealing p r ovisions of theCop yright Act1968, you must not m ake any othe r use of this p r od uct (includ ing cop ying or r e p rod ucing it or p art of it in any way)

unless you have the writte n p e r m ission of the N orthe r n Ter ritory of Australia to d o so.The N orthe r n Ter ritor y of Australia d oes not war rant that the p r od uct, or any p art of it, is cor r e ct or com p lete and will not b e

liab le for any loss, d am age or injur y suffe r e d b y any p e rson as a r esult of its inac curacy or incom p lete ness.

© N orthe r n Te r ritory of Australia

Stuart Hwy

Batchelor

Pine Creek

Palmerston

Adelaide River

DundeeBeach

Darwin

Wadeye

Nauiyu

(unsuitable habitat)

Confidence RatingHighMod e rateLow

N orthe r nTe r ritor y

* Subpopulation location reference number Details ab out sp e cies p op ulation d ensity are d escrib e d in the tab le, Population Status of Ptychosperma macarthurii, July 2016 in the attache d Guid elines for Map Use. * * Highest Likelihood of Occurrence Re p r esent the m ost suitab le ar eas for Ptychosperma macarthurii within the Top End and ar e thought to r e p r esent the largest p ossib le ar ea within which the sp e cies could conceivab ly b e encounte r e d (b ase d on cur r e nt colle ction d ata and e cological knowle d ge). These suitab le ar eas have b e en d elineate d p rimarily on the b asis on the d istrib ution of ‘Sp ring Fe d ’ for ests in the Monsoon Vine-Forests of the Northern Territory map p ing d ata. Data Source Flora Data and Surveys: - Flora and Fauna Division, De p artm ent of Land Resour ce Managem e nt - EcOz Environm ental Consultants Pty Ltd 2015 Potential Hab itat: - Flora and Fauna Division, De p artm ent of Land Resour c e Managem e nt Monsoon V ine-For ests in the N orthe r n Te r ritory (scale 1:100 000 to 1:250 000) Major, Minor Str eams: (scale 1:250 000) - © Com m onwealth of Australia (Bureau of Meteorology) 2012 N T Parks and Rese rves: - Parks and W ild life Com mission of the N orthe r n Te r ritory Cad astr e/Road s/Placenam es/Contours/Image ry: - De p artm ent of Land s, Planning and Environm ent Ae rial Photograp hy - N TLIS W MS Image Se rve r, DLPE

LEGENDConfirm e d Ab se nc e

Re cor d e d sp e cies location (p oint)Confirm e dUnc e rtain

Extent of Occur r e nc e (EOO)Confirm e dUnc e rtain

Highest Like lihood of Occur r e nc e **

Pote ntial Hab itat

N T Park/Rese rve

Cad astre (hund r e d s)

Cad astre (p ar c e ls)Main map: land parcels >1km2, Inset: all land parcels shownContours (Inset) scale 1:25 000

Ind ex (25m inte rvals)Inte r m e d iate (5m inte rvals)

- Sub p op ulation location r efe r e nc e num b e r *1

Minimum Convex Polygon

Guidelines for Map Use, July 2016 - Stylidium ensatum 1

Department of Land Resource Management Flora and Fauna Division July 2016, Version 1.0

Threatened Species of the Greater Darwin Region – Ptychosperma macarthurii

Guidelines for Map Use

Data used to compile this map product is current at the date of publication. Users are encouraged to check the project Metadata record for more recent versions of this map product.

1. Potential habitat mapping is derived from mapping of Northern Territory rainforest vegetation between approximately 1978 and 1989, produced from aerial photograph interpretation of black and white hardcopy prints. Vegetation types were delineated at scales between 1:100 000 and 1:250 000 and subsequently transferred to digital media; consequently the accuracy of this data at fine scales may be limited by the resolution of the original data. For the purposes of this map product, only ‘Spring Fed’ and ‘Riparian’ forest types were considered to represent potential habitat for the species. Site based assessment at an appropriate intensity should accompany use of this map data for all areas. More detail: Potential habitat mapping is derived from mapping produced to define the distribution of Monsoon Vine-Forests in the Northern Territory at scales ranging between 1:100 000 in the Darwin region and up to 1:250 000 elsewhere. These map products were derived using aerial photography interpretation and digitized on a range of mapping bases between the late-1970’s and 1980’s. It should be noted that the potential habitat mapping has an inherent level of spatial inaccuracy associated with the scale and methods of production used to derive the original mapping. These spatial inaccuracies are principally a function of the age of the surveys and the technology available at the time to produce the ORIGINAL maps, the mapping base (topographic or cadastral) on which the original hard-copy mapping was produced and the process of transferring these products to digital media at a later date. The scale of the original mapping was at best 1:100 000 and enlarging the mapping beyond this scale does not provide further detail.

2. The extent of potential habitat displayed on the map is current at the publication date of the latest available land use data for the region. More detail: Areas of remaining potential habitat were identified by intersecting the most up-to date land-use information to exclude areas of intensive land-use and/or cleared areas now unlikely to support viable potential habitat for the species. Data on the extent of remaining potential habitat is current to 2008 and it is likely that the area of remaining intact viable habitat differs from that indicated on the map. This is particularly true in the peri-urban region of Darwin where current land uses do not reflect the historic mapping of the area. This may result in areas of indicated potential habitat no longer existing or conversely transitional vegetation regrowing after historic land-use changes may provide additional habitat not accounted for by the mapping data.

3. The map should be used as a guide to identifying the probability that the species is present in any particular area and not a definitive assessment of distribution. The map can be used to assess the risk associated with a particular activity at a location and the

Guidelines for Map Use, July 2016 - Stylidium ensatum 2

likelihood that the activity may result in a significant impact upon a (sub-) population of a threatened species. More detail: The vegetation communities identified as potential habitat on the map represent those most likely to support populations of Ptychosperma macarthurii based on current ecological knowledge at the date of publication. They do not identify all areas where the species may occur and conversely P. macarthurii is highly unlikely to occur in all the areas identified on the map as potential habitat. The map should be used as a guide to identifying the probability that the species is present in any particular area and not a definitive assessment of distribution. The map can be used to assess the risk associated with a particular activity at a location and the likelihood that the activity may result in a significant impact upon a population of a threatened species.

4. Highest likelihood of occurrence of Ptychosperma macarthurii is known to be strongly associated with spring-fed rainforest patches on the margins of riverine floodplains with an ongoing water supply that results in free-water or saturated soil moisture conditions at/near the land surface for an extended part of the year. For mapping purposes, ‘Spring Fed’ forest identified in the potential habitat mapping was considered the vegetation community with the highest likelihood of supporting P. macarthurii in the greater Darwin Region. Inherent inaccuracies in the mapping data, discussed previously, may result in known locations of P. macarthurii falling outside of the potential and most suitable habitat areas. More detail: At a finer spatial resolution, current understanding of the site level distribution of P. macarthurii indicates that the occupancy envelope for the taxon appears to be centred on those parts of the landscape where saturated soil profiles (Hydrosols) or free surface water persist into the mid-late dry season. A buffer was not applied to the boundaries of the potential habitat polygons in this instance as the potential habitat is considered to significantly limit the likelihood that the species occurs beyond the boundaries of these polygons and is more likely to be restricted to core central areas within the potential habitat where water persists in the landscape for longer periods. Currently available spatial information regarding water body persistence does not correlate well with the hypothesised distribution of the species due to omission errors resulting from high tree canopy cover over the areas of spring fed water supply. Consequently, a precautionary approach was adopted in identifying areas of highest likelihood of P. macarthurii occurrence and all potential habitat was considered of uniform significance in his assessment. Consequently, occurrences of P. macarthurii, may fall outside the mapped areas inferred as most likely support the species as a result of issues associated with inaccuracies in the potential habitat mapping and site-scale variations in habitat conditions and/or error associated with the positional accuracy of the species record. As with any natural resource spatial product, it is recommended that this information be used as a guide to the most likely areas in which P. macarthurii may be encountered and should be accompanied by appropriately timed field survey to clarify the presence or absence of the species from a particular location more definitively.

5. Confirmed absence locations represent detailed floristic survey sites sampled at an appropriate time of year where Ptychosperma macarthurii was not recorded. More detail: Targeted search locations (absence) data are full-floristic sites sampled within the expected distribution of Ptychosperma macarthurii over a number of years between June and August. These sites represent locations sampled at an appropriate time of year where P. macarthurii was not recorded and collection data suggests the species should be present and most detectable. Surveyed sites were largely sampled using a standardised methodology (Brocklehurst et al. 2007) by experienced NTG botanical staff and represent the best available information on the known distribution of the. This does not definitively imply that the species was not present at the location given the cryptic life history of the taxon and the seasonal conditions at the time of

Guidelines for Map Use, July 2016 - Stylidium ensatum 3

sampling. However, it can be considered to represent the temporal ‘window’ of maximum-likelihood that the species would have been detected if it was present at the time of sampling.

6. ‘Point’ data on the occurrence of Ptychosperma macarthurii represent vouchered individual specimens or discrete locations at which the species has been recorded in the field. This data has been critically evaluated and refined for the purposes of this map product to represent locations at which the species presence is confirmed or otherwise.

7. Extent of Occurrence has been calculated using both the complete set of record data and the refined data with confirmed geocodes (see above). This is to allow a more thorough analysis of the species IUCN status based on the most reliable information available for the species. Both sets of results are presented on the following pages for comparison purposes.

8. Mapping reliability (Confidence Rating) has been assigned to areas in which varying intensities of field assessment have been undertaken or are considered to represent suitable habitat for the species. These generically range from ‘High’ being high-intensity, targeted, species specific surveys or areas in which confirmed sub-populations are known to occur through to ‘Low’ where reconnaissance level or incidental surveys have been undertaken or habitat is considered unsuitable. More detail: In the broader regional context (represented on the location map) areas within the greater Darwin region where appropriately timed general surveys have not encountered the species are considered a moderately reliable indication of species presence/absence based on the intensity of field data collection. No distinction is made between site-level reliability at this scale for display purposes and more detail is provided in the insets (see below) At the finer scale within the Extent of Occurrence (EoO – represented by the inset maps), the confidence levels have been combined with the potentially suitable areas (i.e. highest likelihood of occurrence) to give an indication of the confidence in not only the level of survey undertaken, and therefore, the presence/absence of the species, but also the presence of habitat suitable for the species. Areas where targeted surveys have been undertaken within highly suitable habitat are considered highly reliable, whereas areas of unsuitable habitat within the EoO are indicative of unsuitable habitat (e.g. wetlands or built up areas) and are also considered highly reliable (as absences) but not displayed on the inset maps. Areas of potentially suitable habitat with a general level of field survey are given a moderate reliability.

References

Name More Information

Mapped Distribution for Threatened Species of the Greater Darwin Region Project description, metadata record

• Metadata

Kerrigan, R., I. Cowie & B. Baker (2002d) Threatened species of the Northern Territory - Ptychosperma bleeseri (P. macarthurii) Burret, Darwin Palm Parks & Wildlife Commission, NT. (sourced from Flora NT website)

• Web Details • Information sheet

Brocklehurst, P., Lewis, D., Napier, D. and Lynch, D. (2007) Northern Territory Guidelines and Field Methodology for Vegetation Survey and Mapping. Technical Report No. 02/2007D, Department of Natural Resources, Environment and the Arts, Palmerston, Northern Territory.

• Report (NT Library)

Guidelines for Map Use, July 2016 - Stylidium ensatum 4

Liddle, D.T., Brook, B., Matthews, J., Taylor, S.M. and Caley, P. (2006). Threat and response: A decade of decline in a regionally endangered rainforest palm affected by fire and introduced animals. Biological Conservation 132, 362-375.

Anon. (1998). A Management Program for Ptychosperma bleeseri Burret in the Northern Territory of Australia. Parks and Wildlife Commission of the Northern Territory, Palmerston.

Guidelines for Map Use, July 2016 - Stylidium ensatum 5

Population status of Ptychosperma macarthurii (July 2016)

Sub-population Reference no Status Size* Trend Pressures Uncertainties Current Knowledge State

1 Bankers Jungle

Extant c. 50 plants Documented Decline

Introduced animal management Weed incursion

Ongoing subpopulation trend Effectiveness of management actions

Loss of significant numbers of adult individuals from sub-population between 1990 and 2001.

2

Extant >1000 plants

Stable? Hydrology Fire management Weed incursion

Location of specific records General area of records are known to be Black Jungle, however the specific location is uncertain.

2a Crocodile Creek, Black Jungle

Extant >1000 plants

Stable Hydrology Fire management Weed incursion

Ongoing subpopulation trend Effectiveness of management actions

Largest known population with relatively skewed size class distribution toward juveniles. Evidence suggests good levels of recruitment to adult size classes over time in the absence of major disturbance. Sub-population appeared relatively stable at time of last survey.

2b Black Creek, Black Jungle

Extant <100 plants Stable? Hydrology Fire management Weed incursion

Ongoing subpopulation trend Sub-population appeared relatively stable at time of last survey.

2c BJ3 Rainforest, Black Jungle

Extant <10 plants Stable? Hydrology Fire management Weed incursion

Ongoing subpopulation trend Sub-population appeared relatively stable at time of last survey.

3 Whitewood Road Jungle

Extant <250 plants Documented Decline

Hydrology Fire Management Weed incursion

Current status of population. Maintenance of hydrology with increasing development pressure

Only currently known subpopulation outside the Adelaide River catchment. Documented decline between 1990 and 2001 representing approximately 60% of the population.

4 7 and 8 km

Extant <50 plants Stable? Introduced animal

Ongoing subpopulation trend Maintenance of hydrology with

Considered as a single sub-population for the purposes of this analysis.

Guidelines for Map Use, July 2016 - Stylidium ensatum 6

Sub-population Reference no Status Size* Trend Pressures Uncertainties Current Knowledge State

Jungles, Sandy Creek

management Potential development Hydrology Fire Management Weed incursion

increasing development pressure, particularly sand and gravel extraction

Sub-population appeared relatively stable at time of last survey.

5 Daminmin Jungle

Extant >1000 plants

Stable Management Weed incursion

Ongoing subpopulation trend Drivers of sub-population demographics and structure

Sub-population appeared relatively stable in terms of overall numbers at time of last survey. Skewed demographics toward juvenile size classes although some evidence to suggest that recruitment to adult size classes is occurring albeit at low rates.

6 Shoal Bay Conservation Reserve

Unknown Unknown Unknown Geocode of collection appears incorrect

Population unlikely to be extant at location if the original geocode was correct. location.

*Includes juvenile plants as of 2000-2001.

Extent of Occurrence (excluding ocean) 200 km2 (281 km2)** Total EoO; 4.30 km2 (7.40 km2) Conservatively estimated potential habitat within currently known extent; & 4.00 km2 (6.95 km2) Potential habitat with highest likelihood of occurrence within the currently known extent.

Area of Occupancy 10 (13) Number of 2 km2 cells within which mapped records occur (brackets represent ALL historical records*)

Estimated Area of Occupancy at Reference Scale: 40 (52) km2

Estimated Area of Occupancy at Ecological Scale (200m grid cell): 17 cells /0.68 km2 (19 cells/0.76 km2) 1 International Union for Conservation of Nature (IUCN) Standards and Petitions Subcommittee. (2014). Guidelines for using the IUCN Red List categories and criteria. Version 11. Prepared by the Standards and Petitions Subcommittee. http://www.iucnredlist.org/documents/RedListGuidelines.pdf ** Figures in brackets represent the total EoO including all uncertain records.