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    LATEGLACIAL TO HOLOCENE ALLUVIATION AND LANDSCAPE DEVELOPMENT: HIGHSPEED 1 INVESTIGATIONS AT STRATFORD, LONDON BOROUGH OF NEWHAM

    Catherine Barnett, Rob Scaife and Chris J. Stevens

    SUMMARY

     A large hollow was observed cutting the underlying Pleistocene gravels during excavations in

    advance of construction of High Speed 1 (HS1; formerly Channel Tunnel Rail Link) at Stratford

    Box, NGR 538250 184650 (Figure 1; see Valler and Crockett 2011). This hollow (feature 4271;

    Figure 2) proved to be the edge of a substantial relict channel forming a tributary of the River Lea,

    that contained well-stratified waterlogged alluvial and peat deposits (Figure 3). Analysis of the

    sediments, pollen (Figure 4), waterlogged plant remains and wood, coupled with a series of three

     AMS radiocarbon dates, demonstrate this as an important, well-preserved Lateglacial to mid-

    Holocene sequence. A detailed picture of the evolution of the local fluvial, fen and terrace

    landscape with colonisation and succession of plant types from cold stage tundra types, to pine

    forest and ultimately deciduous woodland is presented. The peat itself formed in the early

    Holocene and early Mesolithic at 9460-9270 cal BC to 8450-8240 cal BC. Although its growth

    relates predominantly to local conditions, it may also have an indirect relationship with a more

    widespread regression that predates the Tilbury sequence described by Devoy (1979, 1980),

    Gibbard (1994) and Haggart (1995). The rich terrestrial fen environment represented by the peat

    would have offered plentiful food and other resources to any Mesolithic groups present in the area.

    There was a return to alluvial sedimentation and an increase in spring activity in later prehistory

    (Late Mesolithic/ Early Neolithic to Bronze Age), represented by deposits which may in part relate

    to the Thames III transgression observed elsewhere in the catchment (Devoy 1979, 1980; Gibbard

    1994; Haggart 1995). The environment was increasingly open due to human activity, which

    included the establishment of rough pasture and possible arable cultivation nearby. However,

    willow continued to grow along the wetlands and small stands of mixed deciduous types including

    ash, hornbeam and oak were available for management and use by humans. That they were

    exploited in this way is demonstrated by the worked wood assemblage of Middle Bronze Age,

    Middle Iron Age and Saxon dates at the site (Valler and Crockett 2011). Indeed stake alignments

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    of Bronze Age and Iron Age date which follow the eastern edge of the hollow/channel, pushed into

    the top of the earlier peat at a time of increased alluviation suggest a direct response to increasing

    water levels and minor flood events. This was perhaps an attempt to stabilise the wetland edge in

    order to protect agricultural land or to create a structure that enabled continued access to the area

    for water or grazing. These upper alluvial sediments therefore provide a useful record of the

    environment in part contemporary with human activity recorded from the archaeological remains.

    The very top of the sequence contains food plant remains (spelt wheat and fig) of likely Romano-

    British date, suggesting there may have been contemporaneous settlement nearby.

    INTRODUCTION

    High Speed 1 (HS1), formerly the Channel Tunnel Rail Link, is a new high speed railway linking

    London St Pancras to the Channel Tunnel, and crossing the River Lea at Stratford en route. Works

    undertaken at Stratford primarily comprised the construction of a substantial sub-surface structure,

    colloquially referred to as the ‘Stratford Box’, which carries the new track in a concrete-lined cutting

    across the east side of the Lea Valley.

    The site lies on the NW perimeter of Stratford town centre in the London Borough of Newham, East

    London (Figure 1). It covers most of the eastern side of the floodplain of the River Lea, and is

    bounded by the river to the west and higher ground overlooking the floodplain to the east. To the

    north is the A12 trunk road, and to the south formerly Carpenter’s Road, though much of this route

    has since been re-modelled as part of the London 2012 Olympic Park works. With specific

    reference to this more recent development, the Stratford Box passes centrally through the Olympic

    Village (within Olympic Delivery Authority Planning Delivery Zones 9 and 11; not illustrated).

    The land within the site is generally level, with modern ground surface at approximately 5m above

    Ordnance Datum (OD). Overall, the site comprises an approximate diamond-shaped parcel of land

    measuring at its extremities 1.2km east to west and 1km north to south, though the majority of

    archaeological investigations focussed on the central east to west corridor of Stratford Box itself.

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     As part of a comprehensive programme of archaeological investigations associated with the

    construction of HS1 (which also included desk-based assessment, boreholes, trial trenches and

    construction watching brief), an archaeological excavation (formed of two adjoining areas) was

    undertaken by Wessex Archaeology at the west end of the Stratford Box. The centre of the

    excavation was dominated by a large hollow (feature 4271; Figure 2), extending to the south

    beyond the limit of excavation.

    The hollow was filled with a sequence of alluvial channel edge and peat deposits (Figure 3) which,

    being waterlogged and in indirect association with excavated archaeological remains, were of

    raised palaeoenvironmental potential. Following assessment, selected samples were therefore

    analysed to elucidate the sedimentary history of the feature, and to build a picture of the past local

    environment using pollen, waterlogged plant remains and waterlogged wood. A series of

    radiocarbon dates was gained in order to place the sequence in its chronological and

    archaeological context.

    It should be noted that for consistency, this report will adopt the conventions Lateglacial and Post-

    glacial to describe the Palaeolithic – Mesolithic transition.

    RESULTS

    The Sediments

    Catherine Barnett

    Introduction

    The sedimentary sequence at Stratford Box is generally of made ground over stiff clays, sands and

    iron-rich gravels laid down under fluctuating fluvial conditions, most likely during the Devensian

    cold-stage, forming a gravel terrace. The local underlying solid geology was recorded as Tertiary

    London Clay, with facies of both the Woolwich and Reading Beds. The rafted organic deposits of

    the Lea Valley Arctic Beds which are found, albeit rarely, in the gravels in this area (see Allison  et

    al. 1952), were not found in any of the excavations.

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    One notable exception to this stratigraphy was recorded within the excavation area, where a large

    hollow or basin (feature 4271; Figure 2), filled with a complex of waterlogged fine sediments and

    peat of Holocene age, overlay the Pleistocene gravels. A monolith sequence some 1.67m deep

    was taken through the profile (Figure 3) and is described in detail in Table 1. The monoliths were

    cleaned prior to recording and standard geoarchaeological descriptions used, (following Hodgson

    1997) including Munsell colour, texture, structure and nature of boundaries.

    The feature was associated with a series of worked wooden stakes (see Valler and Crockett 2011),

    which were pushed into the top of the peat and through to the underlying gravels along one edge

    of the feature. These stakes have been dated, the main alignment being of Middle Bronze Age

    date with a solitary Middle Iron Age stake to the south. In addition to assessing the potential of

    these deposits for the preservation of environmental remains, of specific interest is the relationship

    between the changing depositional environment over time and the human response to these

    changes in the form of waterside structures.

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    Table 1: Sediment Descrip tions for Hollow 4271

    Site code  ARC SBX00 Notes Monolith 1084 and 1085 overlapping.Monolith 1084 0-1.05M and monolith 1085 0.88-1.67mMonolith 1084 0m = 0.953m OD

    Feature 4271 hollow

    Section no. 2059

    Context Sampleno.

    m OD Depth (m) Texture Uppercontact

    Description Interpretation

    From To

    4316 0.953- 0.703 0 0.25 Sandysilt

    - Mixed layer of slightly humicfine sand and silt (2.5Y 3/2 verydark greyish brown) with veryabundant shells and abundanttwig and wood fragments.Patches of greasy (2.5Y 4/1dark grey) silty clay. Largewood fragments in the darkgrey silty clay at 0.13-0.20m

    Edge depositwith alluvialinput

    4317 0.703- 0.513 0.25 0.44 Siltyloam

    Clear (2.5Y 3/2 very dark greyishbrown) Massive slightly humicsmooth silt with no inclusions

     Alluvium

    4317 0.513- 0.403 0.44 0.55 Sandysilt

    Clear - Abrupt

    2.5Y 3/2 very dark greyishbrown slightly humic silt andfine sand, no visible inclusions.

     Alluvium

    4318 0.403- 0.283 0.55 0.67 Sand Clear 2.5Y 4/3 Olive brown cleanmedium-coarse calcareoussands, common shells,common small stones, somewith calcareous silt coatingforming tufaceousnodules/concretions.

     Alluvium

    4319-4324

    0.283- 0.153 0.67 0.8 Siltysand

     Abrupt(erosional)

    Variable bands 1-5cm of 'dirty'humic medium sands withshells, organic silty sands and

    organic silty sands withreworked tufa/tufaceous marl -intercalation and inwash oferoded material. at 0.72 =organic silty sand

    Intercalatingedge andalluvial/spring

    deposits

    4325 0.153- 0.123 0.8 0.83 Peatysand

    Clear "twig layer" 70% twig and woodin medium coarse organiccalcareous sand (2.5Y 2.5/1black)

    humic edgedeposits

    4326 0.123- 0.083 0.83 0.87 Sand Sharp 2.5Y 4/2 dark greyish brownmedium dirty calcareous sand.Some finely comminutedorganics. Common shellincreasing fine sand and

    organic content to top of

     Alluvium

    4327 0.083- 0.053 0.87 0.89 Peat Sharpwavy?erosional

    7.5YR 2.5/2 very dark brownand black soft slightly humifiedslightly silty fibrous peat,numerous well-preservedrecognisable herbaceous plantremains and vertical rootlets totop of layer.

    Edge fen peat

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    Site code  ARC SBX00 Notes Monolith 1084 and 1085 overlapping.Monolith 1084 0-1.05M and monolith 1085 0.88-1.67mMonolith 1084 0m = 0.953m OD

    Feature 4271 hollow

    Section no. 2059

    Context Sampleno.

    m OD Depth (m) Texture Uppercontact

    Description Interpretation

    From To

    4327 0.053- 0.033 0.89 0.92 Siltypeat

    7.5YR 2.4/1 black humified siltypeat with common to abundantvery small - small roundedchalk fragments, rare mediumrounded chalk pieces anddegraded chalk matrix. Few finewood fragments, andrecognisable plant matterincluding fine woody stem/rootfragments.

    Peat and chalkyinwash

    4327 0.033- -0.357 0.92 1.31 Peat Abruptwavy/indurated

    7.5YR 2.4/1 Black humified(silty) Phragmites peat (finereed stems material common),very fine irregular bands (1mm)of fine silt, increasing at base.

    Peat with siltinwash

    4328 -0.357- -0.467 1.31 1.42 Silt Gradual Coarse banded very darkgreyish brown (7.5YR 3/2) siltsand humic silts with some veryfine lenses of freshwater shells(including Pisidium sp.) Mediumflint resting on contact

     Alluvium

    4329 -0.467- -0.717 1.42 1.67 Silt Sharp Zone of banded silts 1.42-1.47m dark greyish brown (2.5Y4/2) silt to silty clay. 1.47-1.59m very dark grey (2.5Y 2/1)humic fine banded silt with thinlaminae of freshwater shell(including Pisidium sp.) 1.59-1.62m massive very dark grey(2.5Y 3/1) humic silt, raremedium flint. 1.62-1.67mstrongly and distinctlybanded/laminated silt of olivegrey (5Y 5/2) and dark grey (5Y4/1) of between 2 and 4mmwidth.

     Alluvium (gravelinclusionsobservedimmediatelyunder this levelin excavation)

    Interpretation

    The hollow is suggested to represent the edge of a major Holocene age freshwater channel

    forming a tributary to the River Lea. This cut the Pleistocene gravels and perhaps exploited natural

    variations in their surface topography at the end of the Devensian period (the Lateglacial) or very

    early in the Holocene climatic amelioration. No flood couplets indicative of rhythmic tidal movement

    were found, however, the thin regular banding of the basal humic silts (context 4329) indicates

    repeated changes in energy of flow, perhaps seasonal, with fluctuation between fine alluvial

    sedimentation and semi-stabilisation with establishment of emergent edge fen vegetation providing

    the organic lenses.

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    The channel proceeded to migrate and meander, leaving a widened floodplain across which a

    variety of habitats could establish and where peat began to form away from submerged areas. This

    peat (context 4327) represents a stable, though waterlogged, terrestrial land surface on which

    vegetation established and which would have been accessible to grazing herbivores and for

    human exploitation. An erosional flood water event truncated its top and brought a return to fine

    alluvial sedimentation (context 4326) and the repeated provision of highly calcareous spring water

    is also indicated by the presence of tufaceous nodules (contexts 4319-4324). The hollow continued

    to receive periodic alluvial sediment input from the nearby channel later in its infill (contexts 4316-

    4318) as well as repeated periods of colonisation emergent vegetation and stabilisation.

    The dominant depositional environment from relatively early in the Holocene was therefore of

    channel edge activity with slow moving water and fen. The feature may ultimately have been cut off

    from the main channel, remaining as a waterlogged infilling oxbow although no evidence of a

    dividing bank was found in excavation.

    Dating the Sequence

    Catherine Barnett (neé Chisham)

    Discrete plant remains stratified in the top, middle and base of the peat (context 4327) were

    submitted for dating at the Rafter AMS radiocarbon dating facility as detailed in Table 2. Organic

    remains within the rest of the sequence were potentially reworked given their alluvial context and

    were not therefore chosen for dating. The AMS radiocarbon results were calibrated using Oxcal

    v.3.10 (Bronk Ramsey 1995), incorporating atmospheric data from Reimer et al (2004) and

    expressed at the 94.5% confidence level and to 2 sigma level.

    The results (Table 2) show that peat growth initiated within hollow 4271 during the early post-

    glacial period at 9460-9270 cal BC and it continued to accumulate into the Early Mesolithic period,

    its top at 8450-8240 cal BC eroded by inundation associated with deposition of fine alluvium.

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    Table 2: AMS Radiocarbon Dates from Hollow 4271

    Context Sampleno.

    Material dated,identification

    Comment Resultno.

    C13‰

    ResultBP

    Fractiondetails

    Cal dateBC (2sigma)

    Phase

    4327 1085

    0.88m

    Waterlogged

    wood: degradedtwigwood

    Top of peat. NZA

    27376

    -

    27.4

    909945 Treated

    wood-cellulose

    8450-8240

    cal BC

    early

    Mesolithic

    4327 10850.98 m

    Waterloggedwood: juvenileSalix/Populussp.

    Middle peat. NZA27358

    -27.9

    952240 Treatedwood-cellulose

    9140-8970BC(42.6%)8930-8730cal BC(52.8%)

    earlyPost-glacial

    4327 10851.31m

    Small fragmentsof woody stem

    Base of peat. Thelayer is true peat buthas alluvial siltelement. Providesdate for base of thispeat or the inception

    of peat in the localarea

    NZA27360

    -28.2

    991140 Plantmaterial

    9650-9610BC (2.0%)9460-9270cal BC(93.4%)

    earlyPost-glacial

    The fine humic sediments deposited prior to peat initiation were not suited to absolute dating but

    the analysis of plant remains and pollen (reported below) indicates they are of terminal Lateglacial

    to earliest post-glacial date (potentially pre-Early Mesolithic). Likewise the plant remains from the

    alluvial, spring and edge deposits above the peat give some indication of time span, with late

    prehistoric and Romano-British deposits at the top of the sequence (context 4316).

    Pollen

    Rob Scaife

    Introduction

    Pollen analysis was undertaken on eight 2ml samples taken from the monoliths through feature

    4271. These were processed using standard techniques for the extraction of the sub-fossil pollen

    and spores (Moore and Webb 1978; Moore et al.  1991) in combination with micromesh sieving

    (10u). Taxonomy follows Moore and Webb (1978) modified according to Bennett et al.  (1994) for

    pollen types and Stace (1997) for plant descriptions.

     All samples contained plentiful and well-preserved pollen, counts of 450-700 grains per level were

    achieved and five local pollen assemblage zones were defined as shown in Figure 4.

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    Results

    ZONE 1: -0.447- -0.267M OD (1.4M - 1.22M DEPTH), CONTEXTS 4329 - BASE OF 4327, BETULA-POACEAE .

    Herbs are dominant at c.80%, with Poaceae at 55% and a range of fen and aquatic types. Of note

    are the diverse assemblages of herbs that include diagnostic late-Devensian taxa including

    Polemonium caeruleum (Jacobs ladder), Helianthemum (rockrose), Sanguisorba officinalis (greater

    burnet), cf. Dryas octopetala  (mountain avens) and  Artemisia  (mugwort). Trees and shrubs are

    sparse, comprising Betula (birch) including B. nana  (dwarf birch) in the basal level, Pinus  (pine)

    and Juniperus communis  (juniper). The assemblage is diagnostic of the late-Devensian/earliest

    Holocene and includes taxa from a wide range of habitats including short turf grassland, tall herb

    communities, disturbed/ open ground, fen/ marsh, aquatic and dwarf shrub communities.

    ZONE 2: -0.267- -0.047M OD (1.22M - 1.00M DEPTH), CONTEXT 4327, PINUS-POACEAE.

    Herbs decline in diversity and quantity, especially Poaceae (to 8%) and aquatic Myriophyllum and

    Pediastrum.  There is a sharp expansion of Pinus to 78% at -0.087m OD (1.04m depth) and

    reduction in Betula and Juniperus. Of note is the arrival and expansion (i.e. rational limit) of

    Quercus from the base of this zone along with occasional shrubs including Rhamnus catharticus

    (alder buckthorn), Sorbus type (whitebeam or hawthorn) and Prunus/Malus (apple/plum).

    ZONE 3: -0.047- -0.083M OD (1.00M - 0.87M DEPTH), TOP OF CONTEXT 4327, PINUS-CORYLUS AVELLANA TYPE.

    Pinus declines to absence at the base of zone 4. Corylus avellana type  (hazel) expands to up to

    59%. Ulmus (elm) appears, with a single peak to 18% at 0.88m. Total herb percentages are at their

    lowest in this zone (5-10% maximum). Notably, no Taxus was found.

    ZONE 4: -0.083- 0.283 M OD (0.87-0.67M DEPTH), CONTEXT 4319-4325, ALNUS GLUTINOSA.

    Based on a single sample, this zone is characterised by a massive peak of Alnus glutinosa (alder;

    88%). Corylus avellana  remains present but with a reduced percentage (10%). Pinus  is absent

    while Quercus and Ulmus are also present at low levels. Herbs remain at low levels.

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    ZONE 5: 0.353 - 0.473 M OD (0.60M-0.48M DEPTH), CONTEXT 4317-4318, PLANTAGO LANCEOLATA-POACEAE-

    CYPERACEAE.

     Absolute pollen values are reduced with increased degradation noted.  Alnus is almost absent,

    replaced by Poaceae (50%), Plantago lanceolata (ribwort plantain; 6-7%) and a range of other

    herbs possibly of pastoral affinity. Marshy/fen taxa become more important with increases in

    Cyperaceae (12%), Potamogeton type (pondweed), Typha latifolia (greater reed-mace) and Typha

    angustifolia/Sparganium type (bur reed and lesser reedmace).

    Interpretation

    The Zone 1 pollen assemblage is typical of the Lateglacial cold stage (Younger Dryas/Zone III) of

    between 11600-12900 cal BC (10,000-11,000 BP). There was marked diversity of open

    herbaceous communities and habitats brought about by harsh climatic conditions of a Tundra

    environment in which plants and plant communities were under competition. The landscape was

    devoid of trees except, perhaps occasional birches. The presence of large numbers of algal

    Pediastrum cysts and of aquatic macrophytes suggests that the dominant depositional habitat was

    fen and open water.

    The depositional environment of Zone 2 was a drier grass sedge fen. This pre-Boreal/early Boreal

    period (Holocene Chronozone Ia and Ib) pollen assemblage shows the establishment and spread

    of coniferous woodland in the wider landscape and a consequent decline in herbs after climatic

    amelioration at the end of the Devensian cold stage. A number of warm-loving shrub and tree taxa

    including oak also appear, having migrated/ expanded out of their glacial refugia. Elm then first

    appears in Zone 3, expanding with oak and hazel. However, values of oak are smaller than

    encountered at many other sites for this period, indicating greater local dominance of hazel with

    remaining pine, comparable to the Boreal pine-hazel woodland of Godwin (1975). During this

    phase, the depositional habitat at Stratford Box remained fen but there is also evidence of willow,

    this tree/shrub is normally poorly represented in pollen assemblages and presence here may infer

    development of some floodplain carr woodland.

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    Zone 4 shows a significant change to complete dominance by alder, which colonised and

    expanded locally to form a floodplain alder carr woodland. That and the presence of lime/linden

    (Tilia cordata) suggest this assemblage zone is later in date, probably middle Holocene age

    (Atlantic/ Holocene II) from between ca. 6000-4300 cal BC, representing the late Mesolithic period.

    The change between zones is such as to suggest a hiatus (see sediments).

     A further large environmental change is evident in the uppermost pollen Zone 5, there was a return

    to grass sedge fen on the floodplain, with alder absent. This suggests that there may also be a

    further hiatus in the sedimentation between zone 4 and 5. A decline in woodland is also indicated,

    with only small values of oak and hazel (possibly related to managed stands). The local habitat of

    this phase was a floodplain of grass and sedge, possibly forming rough pasture, with pasture and

    probable cereal cultivation on better-drained soils as indicated by the presence of ribwort plantain

    and an expansion of other herbs, also small numbers of large pollen grains of Poaceae possibly

    from cereal cultivation. A late prehistoric date (Neolithic or post-Neolithic) is indicated for this upper

    sediment.

    The sequence shows the development of vegetation from the close of the last (Devensian) cold

    stage through the early Holocene, the middle Holocene and possibly into the start of the late

    prehistoric period. This profile therefore provides an important long vegetation history represented

    within a relatively short (compacted) sediment series. As such, closer interval pollen analysis,

    notably across the zone boundaries, may prove informative in the future.

    Waterlogged Plant Remains

    Chris J Stevens

    Introduction

     A sequence of fourteen bulk samples from the peat and alluvial deposits of feature 4271 was

    assessed for waterlogged remains, four of which were selected for full analysis (contexts 4316,

    4322, 4326 and 4328). Laboratory flotation was undertaken on 1L sub-samples with flots retained

    on a 0.25mm mesh and residues on a 0.5mm mesh. Residues were fractionated into 5.6mm and

    0.5mm groups and any coarse (>5.6mm) non-waterlogged fraction was sorted, weighed and

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    discarded. The flots and residues were sorted using a low-powered stereo-binocular microscope,

    waterlogged and charred plant material was extracted and identified using the nomenclature of

    Stace (1997). Waterlogged material was recorded in all samples, but was less abundant towards

    the base (contexts 4327 and 4329). Variations in the waterlogged assemblage have allowed

    differentiation into three broad phases.

    Results

    PHASE 1: -0.717 - -0.083 M OD (1.67-0.87M DEPTH), CONTEXTS 4329 TO 4327, BETULA-CHARA

    Tree and shrub species were sparse in the peat and humic silts, but represented by seeds and

    cone scales of birch (Betula sp), including one of dwarf birch (Betula nana). Several buds of

    probable willow (Salix sp.) were also recovered. The assemblage analysed in detail for context

    4328 was dominated by seeds and spores of aquatics including pondweed (Potamogeton  sp.),

    water-crowfoot (Ranunculus subgenus. Batrachium), rigid hornwort (Ceratophyllum demersum),

    water-cress (Rorippa nasturtium-aquaticum), and water-milfoil (Myriophyllum sp.). Also recovered

    were the ephippium or eggs of the water-flea (Daphnia sp.) indicative of still flowing or standing

    water.

    Water’s edge habitats were also represented, with willow-herb (Epilobium sp.), sedges (Carex sp.),

    bristle club rush (Isolepis setaria) and marsh cinquefoil (Potentilla palustris). Dog-violets were

    identified, Viola  subg. Viola, section Uncinatae, (cf. Viola odorata/hirta, attributed to these two

    species by virtue of their large size, Godwin 1984, 137). Given the nature of the rest of the

    assemblage, it is probable that hairy violet (Viola hirta), found within calcareous pastures and open

    scrub, is represented. Taxa of open and disturbed ground included nettle (Urtica dioica) and

    goosefoot type (Chenopodiaceae), also crowberry (Empetrum nigrum), a species today found only

    in the moorlands and uplands of northern Britain.

    PHASE 2: 0.083 - 0.283 M OD ( 0.87-0.67M DEPTH), CONTEXTS 4326 TO 4319, ALNUS GLUTINOSA

    These alluvial sediments were dominated by remains of alder ( Alnus glutinosa) with cones, fruits

    and catkins recovered in large numbers. The samples from context 4326 at the base and context

    4322 analysed in full contained other shrub types including hawthorn (Crataegus monogyna) fruit

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    and thorns, and seeds of dogwood (Cornus sanguinea) and elder (Sambucus niger ). Three hazel

    nut fragments (Corylus avellana) occurred in 4322. Herbaceous woodland or wooded-fen species

    were generally rare, but context 4322 yielded several seeds of woody nightshade (Solanum

    dulcamara). Local aquatic and emergent types were common, including seeds of yellow water-lily

    (Nuphar lutea), sedges (Carex sp.), common club-rush (Schoenoplectrus lacustris), and branched

    bur-reed (Sparganium erectum). Herbaceous water’s edge types were rare at the base but

    increased up the profile to include seeds of buttercup (Ranunculus repens, bulbosus or acris),

    greater spearwort (Ranunculus lingua), bog pimpernel ( Anagallis tenella), mint (Mentha sp.), and

    common nettle (Urtica dioica). Wild hop (Humulus lupulus), which tends to be found on the

    riverbank close to the edge of woodland fen-carr, was also identified.

    PHASE 3: 0.283-0.953M OD (0.67-0M DEPTH), CONTEXTS 4318 TO 4319, RANUNCULUS-RUMEX-

    SCHOENOPLECTRUS

     Alder ( Alnus glutinosa) was virtually absent in this phase, the assemblage dominated instead by

    species of rough grassland, wetlands and aquatic environs. Many species identified can be

    associated with rough-open wet grassland/pasture, such as buttercup (Ranunculus sp.), probably

    creeping buttercup (R. repens) and meadow-buttercup (R. acris), meadow-rue (Thalictrum flavum),

    ragged-robin (Lychnis flos-cuculi), common meadow-rue (Thalictrum flavum), thistle

    (Cirsium/Carduus  sp.) and dock (Rumex sp.), including both clustered dock (Rumex

    conglomeratus) and curled-leaved dock (Rumex crispus). Others are more directly associated with

    open ground, disturbance and nitrogen-rich soils, probably created by animals or human activity,

    such as orache ( Atriplex sp.), goosefoot (Chenopodium sp.), chickweed (Stellaria media), common

    nettle (Urtica dioica) and redshank/persicaria (Persicaria maculosa/lapathifolium).

    Occurrence of bramble (Rubus sp.), elder (Sambucus nigra), woody nightshade (Solanum

    dulcamara), bugle ( Ajuga reptans) and hazelnut (Corylus avellana) indicate presence of some

    wooded scrub or hedge. Herbaceous types representative of wetter muddier soils close to the

    water’s edge and in the shallows include winter cress, (Barbarea vulgaris), golden dock (Rumex

    maritimus), wild celery/fool's watercress ( Apium graveolens/ nodiflorum), yellow iris (Iris

    pseudacorus), bog-bean (Menyanthes trifoliata), narrow fruited water-cress (Rorippa nasturtium-

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    aquaticum), water-droplet (Oenanthe sp.). The vast number of seeds of common club-rush

    (Schoenoplectrus lacustris), sedge (Carex sp.), and branched bur-reed (Sparganium erectum)

    suggests a large wet marshland presence within these later deposits. Aquatics were also well

    represented, including yellow water-lily (Nuphar lutea), water-crowfoot (Ranunculus subgenus.

    Batrachium), as well as horned pondweed (Zannichellia palustris) and water-plantain ( Alisma

    plantago-aquatica).

    Two charred glume bases of spelt wheat (Triticum spelta) and a single seed of fig (Ficus carica)

    were also found in context 4316, demonstrating human activity in the catchment area during this

    phase.

    Interpretation

    PHASE 1: -0.717- -0.083 M OD (1.67-0.87M DEPTH), (EQUIVALENT TO POLLEN ZONES 1-3). LATEGLACIAL TO

    EARLY POST-GLACIAL AND EARLY MESOLITHIC.

    The radiocarbon dates reported for the peat demonstrate an early post-glacial date for the lower

    peat. The waterlogged remains from the humic silts underlying this peat (context 4328) at -0.357- --

    0.467m OD (1.31-1.42m depth), indicate a Lateglacial- early post-glacial date, with dwarf birch and

    crowberry particularly diagnostic of an early date. Deposits and associated waterlogged remains of

    this date are commonly encountered in the Lea Valley, having been recovered at Nazeing (Allison

    et al. 1952; Godwin 1984) and also at Innova Park (Ritchie et al. 2008) to the north. The aquatics

    suggest a large body of open neutral or calcareous water or an extensive open floodplain with

    large stretches of still or slow-flowing water.

    The data for the lower deposits complements the pollen data for Zone 1 (see Scaife above) well,

    both indicating a largely open landscape, with birch being the most significant tree taxa. Dwarf

    birch can grow on drier soils next to water and it is probable that birch, along with dwarf willows,

    grew up to the water’s edge. Seeds of grassland rarely survive in waterlogged deposits, but the

    pollen suggests that grassland (or perhaps common reed) dominated much of the immediate

    landscape. While bristle club-rush (Isolepis setacea) is found in marshy meadows it also grows on

    bare patches of sands and gravels. The upper context (4327), dated to the early Mesolithic,

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    contained relatively few plant macrofossils. The pollen evidence (Scaife’s Zone 2-3) suggests pine

    followed by pine and hazel forest during this phase. It was noticeable that wood and twig wood

    became more common suggesting a more forested environment, extending probably up to the

    water’s edge (see Barnett waterlogged wood below).

    PHASE 2: 0.083 - 0.283 M OD ( 0.87-0.67M DEPTH), (EQUIVALENT TO POLLEN ZONE 4). LATER MESOLITHIC TO

    NEOLITHIC/ BRONZE AGE.

    There appears to be a sedimentary hiatus at 0.083m OD (0.87m depth, between contexts 4327

    and 4326) and the precise date of the deposits lying between 0.083 - 0.283 m OD (0.87-0.67m

    depth) is unknown, however, the clear dominance of alder carr demonstrated by the plant remains

    and pollen suggests a late Mesolithic to Late Bronze Age date. The arrival of alder carr at Staines

    Moor was dated to after 8710±70BP/8460±65BP (OXA-6470/ OXA-6474; Keith-Lucas 2000), while

    the presence of lime in the pollen evidence suggests a Neolithic/Early Bronze Age date. The

    majority of dates for alder carr within the London region are for the Early Neolithic to Middle Bronze

     Age (Barnett et al. forthcoming; Crockett et al. 2002), as well as up-river at Runnymede (Greig

    1991) and the lower Colne Valley, Slough (Wessex Archaeology 2006), with clearance of this fen-

    landscape within the Middle to Late Bronze Age (Merriman 1992: 263, Scaife 2000; 2002; Allen et.

    al. 2005, cf. Greig 1991) associated with increasing human activity (Meddens 1996). While alder

    was recovered from Middle to Late Bronze Age deposits within a palaeochannel at Innova Park to

    the north it was noted that probably only small stands were present (Ritchie et al. 2008).

     Although alder would seem to have dominated the local vegetation during this period, within this

    densely wooded riverine habitat, hazel, hawthorn, elder and dogwood all seem to have found a

    niche, probably growing within drier patches on the more stable riverbanks. Within the more open,

    still-flowing areas of the river, permanent pools and ox-bows would be found yellow water-lily

    (Nuphar lutea), while rushes and reeds grew perhaps in narrow strips or occasional patches along

    the edges.

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    PHASE 3: 0.283-0.953M OD (0.67-0M DEPTH), (EQUIVALENT TO POLLEN ZONE 5). ?BRONZE AGE/ IRON AGE TO

    ROMANO-BRITISH.

    These uppermost layers are likely to be of a post-Middle Bronze Age, probable post-Late Iron Age

    date, as they overlie the dated wooden stakes reported by Valler and Crockett (2011). The

    presence of both fig and charred glumes of spelt wheat in the uppermost deposit (context 4316)

    suggests a Romano-British date for the top of this sequence, and indicates nearby settlement of

    this date. The earliest recorded spelt wheat in the general region is Middle Bronze Age from

    Princes Road, Dartford (Pelling 2003). However, it might be noted that Middle and Late Bronze

     Age assemblages both to the north from Innova Park (Ritchie et al. 2008) and to the south-west

    from West-Thurrock (Stevens 2009) have tended to be dominated more by emmer wheat (T.

    dicoccum) rather than spelt. The presence of spelt glumes is more in keeping with a Roman date

    and along with the seed of fig (Ficus carica) would tend to suggest that at least the upper contexts

    are likely to be Romano-British in date.

    The macrofossils and pollen again largely agree, indicating a more open landscape than

    previously, with the removal/ decline of alder carr leaving only single trees or small stands, and

    probably the decline of woodland in general within the region. The species present are mainly of

    open-country species probably associated with rough grassland, including creeping buttercup,

    docks and ragged-robin. Other species such as common nettle indicate some disturbance and

    nitrogen enrichment of the soil, perhaps associated with livestock. Seeds of yellow water-lily

    continue to suggest areas of still to standing water in the vicinity, while species of water-crowfoot

    may be found within both slow-flowing rivers and muddy wet areas at the rivers edge. Similar

    evidence can be seen in the upper deposits from Staines Moor following the decline in alder,

    suggesting an open landscape with meadows and some arable cultivation (Keith-Lucas 2000).

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    Waterlogged Wood

    Catherine Barnett

    Introduction

    Identification was undertaken on ten randomly selected waterlogged wood fragments from each of

    the same four 1 litre waterlogged samples analysed for plant macrofossil remains through the

    sedimentary sequence of Feature 4271. A fine slice was taken from each fragment along three

    planes (transverse section (TS), radial longitudinal section (RL) and tangential longitudinal section

    (TL) and identification undertaken at up to 400x magnification, according to the anatomical

    characteristics described by Schweingruber (1990) and Butterfield and Meylan (1980).

    Identification was to the highest taxonomic level possible, usually that of genus and nomenclature

    is according to Stace (1997). The results are shown in Table 3 below.

     Although the total number of fragments examined was relatively small, differences can be seen in

    the range of taxa represented that coincide with the three phases defined on the basis of

    waterlogged plant macrofossil content.

    Table 3: Waterlogged Wood Identif ications Feature 4271

    Context,depth

    Sampleno.

    Identification Common name No. offragments

    Macrophase

    4316, 0-0.25m

    1086 Salix/ Populus sp. willow/ aspen type 2 3

    Fraxinus excelsior ash 2 (1 twig, 1mature)

    Pomoideae Pomaceous fruits (e.g. hawthorn,apple, pear, whitebeam)

    1

    Carpinus betulus hornbeam 1

    Betula sp. birch 1

    Quercus sp. oak 1 Alnus glutinosa alder 1

    4322, 0.67-0.77m

    1092 Betula sp. Twigwood birch 2 2

     Alnus glutinosa alder 3 (2 twig, 1mature)

    Corylus avellana hazel 1

    cf. Corylus/ Alnus  cf. hazel/ alder 2

    Unidentifiable - 2

    4326, 0.77-0.87m

    1096  Alnus glutinosa alder 4 2

    Salix/ Populus sp.twigwood

    willow/ aspen type 4

    Unidentifiable - 2

    4328, 1.31-1.42m

    1098 Salix/ Populus sp. Twigand roundwood

    willow/ aspen type 10 1

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    Interpretation

    PHASE 1: LATEGLACIAL TO EARLY POST-GLACIAL AND EARLY MESOLITHIC.

    Only one wood type was identified in the oldest sample, willow and aspen cannot be differentiated

    on the basis of their wood anatomy and both could be present in this early Holocene context.

    However, given the lack of aspen macrofossils, usually produced in abundance if present, local

    importance of willow is suggested for the floodplain and wetland edges. All fragments were small

    and immature and it is possible dwarf forms of willow are represented.

    PHASE 2: LATER MESOLITHIC TO NEOLITHIC/ BRONZE AGE.

     A continuation in the importance of damp habitats is indicated by the continued presence of willow/

    aspen and the colonisation of alder. Although alder clearly came to be dominant in the wetland

    zone by the plant macrofossil and pollen data, the wood assemblage also supports an

    interpretation of the local establishment of open canopy mixed woodland on marginal drier areas,

    as represented by the open-loving taxa, hazel and birch.

    PHASE 3: BRONZE AGE/ IRON AGE TO ROMANO-BRITISH.

    Despite the reversion to a broadly open environment as suggested by the pollen data, it can be

    demonstrated that willow and alder continued to grow at the channel edge and what woodland or

    stands of trees that remained locally were rich and varied in taxa. The first appearance of ash

    (Fraxinus excelsior ), hornbeam (Carpinus betulus), oak (Quercus sp.) and Pomoideae occurred in

    this phase. The latter cannot be differentiated to species but may include taxa such as hawthorn,

    apple, and whitebeam. All types represented are native and could have established locally but

    management of this resource, in general decline in later prehistory due to clearance for agriculture,

    might also have taken place nearby. Indeed worked elder, ash and oak have been reported for the

    Middle Bronze Age and Late Iron Age at Stratford Box (Valler and Crockett 2011)

    DISCUSSION

     A detailed Lateglacial to mid-Holocene sedimentary sequence has been described. This is

    important in that continued waterlogging has allowed preservation of a range of

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    palaeoenvironmental remains. The information on past conditions provided from these remains,

    including wood and macrofossils of very local origin, has combined to provide a detailed picture of

    the changing landscape over this period, enhanced by the provision of absolute dates for the main

    peat body (Table 4).

    Table 4: Summary of Palaeoenvironmental Resul ts Feature 4271

    Context m ODDepth(m)

    Summary descrip tion PhaseMacrofossiland woodphase

    Localpollenzone

    Blytt andSernander period

    43160.953-0.703

    0-0.25

    Edge deposit with alluvialinput comprises humic finesand and silt with abundantshells and wood

    Romano-British

    Phase 3

    -

    Sub-Boreal-

    Sub-Atlantic4317 0.703-0.403 0.25-0.55 Humic silt alluviumLate Prehistoric(LBA/ IA)

    Zone 5

    43180.403-0.283

    0.55-0.67

    Calcareous shelly fluvialsands and tufa, truncatedtop of 4319Hiatus

    4319-4324

    0.283-0.153

    0.67-0.8

    Intercalating edge andalluvial/spring deposits ofbanded humic sands siltysands and organic siltysands with reworked tufa late Mesolithic/ Early

    Neolithic to ?EarlyBronze Age

    Phase 2 Zone 4 Atlantic4325

    0.153-0.123

    0.8-0.83

    70% twig and wood inorganic sand

    4326

    0.123-

    0.083

    0.83-

    0.87

    Dirty calcareous sand, may

    have truncated top of 4327Hiatus

    43270.083-0.053

    0.87-0.89

    Fibrous herbaceous fenpeat

    early Mesolithic upto 8450-8240 cal BC

    Phase 1

    Zone 3 Boreal4327

    0.053-0.033

    0.89-0.92

    Humified silty woody peatwith chalky inwash

    early Post-glacial(9469-9270 cal BCat base 1.31m,9140-8730 cal BC at0.98m

    43270.033--0.357

    0.92-1.31

    Humified Phragmites peatwith occasional silt inwash

    Zone 2 Pre-Boreal

    Zone 1 - (Lateglacial)

    4328-0.357--0.357

    1.31-1.42

    Silt and humic silt alluviumLateglacial-earlyPost-glacial - -

    4329

    -0.357-

    -0.717

    1.42-

    1.67

    Humic banded silt alluvium

    (over fluvial gravels)

     An open and initially relatively sparsely vegetated landscape has been demonstrated for the

    terminal Lateglacial to early Post-glacial period. The local cold stage environment (context 4329-

    4328, Phase 1, Zone 1) was of open damp sedge, reed and grass dominated floodplain with

    extensive open moving water. Low-growing shrubs, notably juniper, crowberry, dwarf birch and

    willow, grew on the edge and on drier raised areas of the Pleistocene gravel terrace. Pine

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    woodland established on drier areas early on (base of 4327, Phase 1, Zone 2). Colonisation of a

    wider range of plants then occurred in continued response to the climatic amelioration at the start

    of the Holocene, with open taxa such as hazel expanding, followed by mixed deciduous woodland

    shrubs and tree types such as oak and elm (context 4327, Phase 1, Zone 3), indicating the

    transition from the pre-Boreal to the Boreal period. Fen plant types increasingly dominated the

    floodplain and marginal wetland areas and a thick terrestrial fen peat deposit grew away from the

    channel. This continued into at least the early Mesolithic, the exact date is unknown due to erosion

    of the top of the peat by later floodwaters.

    The lower sequence adds to a small but growing body of environmental evidence for landscape

    evolution gained from wetland sedimentary sequences of Lateglacial to early Post-glacial date in

    the River Thames tributaries such as the Lea (e.g. Allison et al. 1952; Godwin 1984; Lewis 1991,

    2000; Lewis et al. 1992), Colne (e.g. Lacaille 1963; Gibbard and Hall 1982; Keith-Lucas 2000) and

    Kennet Valley (Holyoak 1980; Chisham 2004). Such sequences not only have potential to record

    climatic and vegetation changes but are of raised potential to inform on local early Mesolithic

    activity, notably where charred remains and microcharcoal occur. In addition, parts of these

    floodplains were accessible and where buried soils and peat horizons exist there is the potential for

    often deeply buried in situ early Mesolithic archaeological remains and should remain a focus.

    The sedimentary hiatus during the Mesolithic period is reflected in the vegetation record at

    Stratford Box. The pollen, plant macrofossil and wood assemblages all attest to the subsequent

    spread of dense alder carr on and along the floodplain edge (context 4326-4319, Phase 2, Zone 1)

    during the Atlantic/ late Mesolithic period and the occurrence of small layers. Reworked pieces of

    tufa indicate highly calcareous springs were active under warm wooded conditions in the local

    area. Hazel, oak and elm continued to grow on the terrace with hawthorn, dogwood and elder.

     A second, longer hiatus has been recorded, following repeated fluctuation between stable

    terrestrial, alluvial and spring sedimentation in context 4319, the sequence was truncated once

    more by fluvial inundation, also reflected in the abrupt change to Phase 3 in the macrofossil and

    wood assemblage and Zone 5 in the pollen. Although potentially caused by a local channel shift,

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    this inundation may in part relate to the wider phenomenon of the Thames III transgression

    observed elsewhere in London (Devoy 1979, 1980; Gibbard 1994; Haggart 1995). The upper

    humic alluvial and wetland edge deposits record the late prehistoric environment at Stratford Box,

    most useful in providing a landscape context for the late prehistoric activity demonstrated at the

    site (see Valler and Crockett 2011).

    By the Middle Bronze Age/ Iron Age the floodplain was rich in herb taxa, with areas of still water

    and boggy patches persisting while the immediate area had been opened up by human activity.

    The establishment of rough pasture, with corresponding disturbed and nitrogen enriched soils and

    possible arable cultivation is suggested by the pollen spectrum and macrofossils. An increase in

    the variety of woody taxa at this time however indicates that although trees were fewer in number,

    they had increased in diversity, with the possibility of local management raised. The local late

    prehistoric community clearly exploited the woodland resource, as demonstrated by the presence

    of Middle Bronze Age and Late Iron Age stake alignments and wood working chips. That these

    follow the edge of the hollow/channel, and were pushed into the top of the earlier peat at a time

    when an increase in alluviation has been demonstrated, is interesting. It appears the stakes were a

    direct response to increasing water levels and minor flood events, either an attempt to stabilise the

    wetland edge to protect agricultural land or as part of a structure to enable continued access to the

    area, perhaps for water or grazing.

    The plant macrofossil assemblage from the top of the sequence contained a fig seed and charred

    glumes of spelt wheat in the uppermost deposit (context 4316), which, even if locally reworked,

    suggests a Romano-British date for at least the top sample and also indicates nearby settlement of

    this date despite a lack of use of the water’s edge at this location at the time (see Valler and

    Crockett 2011).

     ACKNOWLEDGEMENTS

    The assistance of Mark Turner and Helen Glass, who monitored the project on behalf of RLE is

    gratefully acknowledged, as is the on-site assistance of Kevin MacFarlane (RLE). Wessex

     Archaeology would also like to thank Union Railways (North) Limited (URN) for commissioning the

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    fieldwork project. Acknowledgement is also due to the on-site staff of contractors Skanska, who

    gave their kind assistance and helped us achieve our aims. The project was monitored by Dr Jane

    Sidell and Nick Truckle (then of English Heritage Greater London Archaeological Advisory

    Service), whose comments and advice throughout the project are acknowledged.

    The fieldwork was managed by Mick Rawlings and Andy Crockett for Wessex Archaeology and

    directed in the field by Hilary Valler, ably assisted by Vic Tomalin as Project Supervisor, and a field

    team too numerous to mention here – this report would not have been possible without their long

    hours dedicated to the project.

    The post-excavation programme was managed by Andy Crockett, on behalf of the Oxford Wessex

     Archaeology Joint Venture, in close consultation with Stephen Haynes of RLE. In addition to the

    many colleagues who provided invaluable peer review for this work, the principal authors would

    particularly like to thank Sarah Wyles and Laura Catlin for their assistance during post-excavation

    analysis. Pollen preparation was undertaken at the Palaeoecology Laboratory of the School of

    Geography, University of Southampton, and radiocarbon determinations provided by the Rafter

    Radiocarbon Laboratory, New Zealand.

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    London: Hodder and Stoughton

    MOORE, WEBB & COLLINSON (1991), P D Moore, J A Webb and M E Collinson Pollen analysis.

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    PELLING (2003), R Pelling ‘Charred plant remains’, 73-76 in P. Hutchings, Ritual and riverside

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    G Blackwell, C E Buck, G S.Burr, K B Cutler, P E Damon, R L Edwards, R G Fairbanks, M

    Friedrich, T P Guilderson, A G Hogg, K A Hughen, B Kromer, G McCormac, S Manning, C Bronk

    Ramsey, R W Reimer, S Remmele, J R Southon, M Stuiver, S Talamo, F W Taylor, J van der

    Plicht and C E Weyhenmeyer ‘IntCal04 terrestrial radiocarbon age calibration, 0-26 cal kyr BP’,

    Radiocarbon, 46 (3), 1029-1058

    RITCHIE et al. (2008), K Ritchie with M J Allen, C Chisham, N Cooke, J Crowther, R Gale,R., G P

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    26

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    B

    3845BH

    3846BH

    3847BH   3848BH

    3849BH

    3850BH 3952BH

    Greater London

    London Borough of Newham

    The Site

    Stratford Box

    Site outline

    Evaluation trench

    Excavation Areas

    Key:

            5        3        8        5        0        0

    185500

    R                      i                       v                 e                r                  L                      

    e                a                 

     ARC3839TT

    Balancing pond

     ARC3841TT ARC3842TT

     ARC3843TT

     ARC3844TT

            5        3        8        0        0        0

    185000

    184500

    Line of High Speed 1

    Borehole

    Line of High Speed 1

    Contains Ordnance Survey data © Crown Copyright and database right 2012.

    500 m0

    Rivers

    Railtrack

    2 1

    Figure 1. Site and trench location plan

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       A  r  e  a  o   f  e  x  c  a  v  a   t   i  o  n

       T   i  m   b  e  r

       H  o   l   l  o  w

       4   2   7   1

       K  e  y  :

       R   i  v  e  r  s

       S  e  c   t   i  o  n

       4   1   4   9

       4   1   5   3

       4   1   3   5

       4   1   3   4

       4   1   3   2

       4   1   4   1

       4   1   4   4

       4   0   7   9

       4   0   8   1

       4   0   8   3

       4   0   1   5

       4   2

       7   1

       4   2   6   4

       4   2   7   1

       4   1   4   8

       4

       1   1   4

       4   2   7   4

       4   1   1   0

       4   2   3   4

       4   1   6   9

       4   2   0   6

       4   2   0   3

       4   2   1   0

       4   1   2   5

       4   1   5   5

       4   2   2   5

       4   2   4   3

       (   F   i  g .   3   )

       1   0  m

       0

       A  r  e  a   2

      C  h a  n  n e

      l s e a   R

      i  v e  r

       A  r  e  a   1

    Waterworks River 

       M  o  d  e  r  n  s  e  r  v i  c  e

       F   i  g  u  r  e   2 .   S   t  r  a   t   f  o  r   d   B  o  x  e  x  c  a  v  a   t   i  o  n   A  r  e  a  s   1  a  n   d   2 ,  s   h  o  w   i  n  g   l  o  c  a   t   i  o  n  o   f   H  o   l   l  o  w

       4   2   7   1

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    0.5 m0

    0.953 maOD

    4316

    4317

    4318

    4319

    4321

    4323

    43204322

    43244325

    4326

    4327

    4327

    4328

    4329

    1085

    0.072 maOD

    1084

    Key:

    Bark / timber fragments

    Chalk

    Gravel

    N S

    Silt / alluvium

    Peat

    Sand

    Sand and silt with alluvium

    Chamfered step

    Figure 3. Section through the sedimentary sequence filling hollow 4271

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         T     R     E     E     S     &     S     H     R     U     B     S

         S     t    r    a     t .    B     E

         T     U   L    A   B

         E     T     U   L

        A    c    f .     N

        A     N    A

         P    I     N     U

         S

         U   L     M

         U     S     Q

         U     E     R     C

         U     S

         P     O     P     U

       L     U     S

         T    I   L    I

        A   C    A     R

         P    I     N     U

         S    B     E

         T     U   L     U

         S

        A   L     N     U

         S     G   L     U

         T    I     N     O

         S    A

         S     C     R     O

         P     H     U   L

        A     R    I    A

         C     E    A     E

         U     N     D    I

         F     F .

        J     U     N    I

         P     E     R     U

         S     C     O

         M     M

         U     N    I     S

         R     H    A     M

         N     U     S

         C    A     T     H

        A     R     T    I

         C     U     S

         S     O     R    B

         U     S     T     Y

         P     E

         P     R     U     N

         U     S    /     M

        A   L     U     S

         T     Y

         P     E

         C     O     R     Y

       L     U     S

        A     V     E   L   L

        A     N    A

         S    A   L    I

         X   c    f .     V

        I    B     U     R

         N     U     M

         E     R    I     C

        A   R    A     N     U

         N     C     U   L

        A     C     E    A

         E     U     N

         D    I     F     F .

         R    A     N     U

         N     C     U   L

         U     S     T     Y

         P     E

         T     H    A   L

        I     C     T     R

         U     M

         S    I     N    A

         P    I     S

         T     Y     P     E

         H     Y     P     E

         R    I     C     U

         M

         H     E   L    I

        A     N     T     H

         E     M     U     M

         C    A     R     Y

         O     P     H

         Y   L   L    A

         C     E    A     E

         U     N     D    I

         F     F .

         C     E     R    A

         S     T    I     U

         M     T     Y

         P     E

         C     H     E     N

         O     P     O

         D    I    A     C

         E    A     E

         F    A    B    A

         C     E    A     E

         U     N     D    I

         F     F .

         M     E     D

        I     C    A     G

         O     T     Y

         P     E

         R     O     S    A

         C     E    A     E

         U     N     D    I

         F     F .

         F    I   L    I

         P     E     N     D     U

       L    A

         P     O     T

         E     N     T    I

       L   L    A

         T     Y     P     E

         D     R     Y    A

         S     O

         C     T     O

         P     E     T     U

       L    A

         S    A     N     G

         U    I     S     O

         R    B    A

         O     F     F

        I     C    I     N

        A   L    I     S

        A     P    I    A

         C     E    A     E

         S    A     N     G

         U    I     S     O

         R    B    A

         M    I     N

         O     R

         E     U     P     H

         O     R    B

        I    A

         R     U     M

         E     X      P     O   L     E

         M     O

         N    I     U     M

         C    A     E     R

         U   L     E     U

         M

         R     H    I     N

        A     N     T     H

         U     S     T     Y

         P     E

         O     D     O

         N     T    I     T

         E     S     T     Y

         P     E

         M     E     N

         T     H    A

         T     Y     P     E

         H     E     R     B     S

         Z    o    n    e

         Z    o    n    e     5

         Z    o    n    e     4

         Z    o    n    e     3

         Z    o    n    e     2

         Z    o    n    e     1

         2     0

         %

         P    o     l     l    e    n     S    u    m

         0 .     4

         8

         0 .     5

         6

         0 .     6

         4

         0 .     7

         2

         0 .     8

         0

         0 .     8

         8

         0 .     9

         6

         1 .     0

         4

         1 .     1

         2

         1 .     2

         0

         1 .     2

         8

         1 .     3

         6

        D   e   p  t   h     m

         6

         0

         4     0

         2     0

         6     0

         4     0

         2     0

         8     0

         6     0

         4     0

         2     0

         8     0

         2     0

         2     0

         I    n     t    e    r    c    a     l    a     t     i    n    g     h    u    m     i    c

        s    a    n     d    s ,    s     i     l     t    y    s    a    n     d    s    a    n     d     t    u     f    a

         P    e    a     t    y    s    a    n     d  -     h    u    m     i    c

        e     d    g    e     d    e    p    o    s     i     t

         C    a     l    c    a    r    e    a    s    s    a    n     d    y    a     l     l    u    v     i    u    m

         P    e    a     t

         S     i     l     t    y    p    e    a     t    w     i     t     h    c     h    a     l     k

         i    n    w    a    s     h

         S    a    n     d    y

        s     i     l     t    a     l     l    u    v     i    u    m

         H     i    a     t    u    s

         S     i     l     t    a     l     l    u

        v     i    u    m

         E    r    o    s     i    o    n    a     l     b    o    u    n     d    a    r    y

         F     i    g    u    r    e     4 .

         P    o     l     l    e    n     d     i    a    g    r    a    m

         f    o    r     H    o

         l     l    o    w     4     2     7     1

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         P   L    A     N

         T    A     G

         O     M

        A    J     O

         R     T     Y

         P     E

         P   L    A     N

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         N     C     E     O

       L    A     T    A

         P   L    A     N

         T    A     G

         O     C     O

         R     O     N     O

         P     U     S

         T     Y     P     E

         R     U    B    I

        A     C     E    A

         E

        B    I     D     E

         N     S     T     Y

         P     E

        A     N     T     H

         E     M    I     S

         T     Y     P     E

        A     R     T     E

         M    I     S

        I    A

         C    I     R     S

        I     U     M

         T     Y     P     E

         C     E     N     T

        A     U     R     E

        A     N    I     G

         R    A     T     Y

         P     E

         S     E     R     R

        A     T     U   L    A

         T     Y     P     E

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         E    A     E

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        A     C     E    A

         E

       L    A     R     G

         E     P     O

        A     C     E    A

         E

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        I     T     R    I

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         T     Y     P     E

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         G     E     T

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         T     Y     P     H

        A   L    A

         T    I     F     O

       L    I    A

         T     Y     P     E

         T     Y     P     H

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         G     U     S

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        A     T     Y

         P     E

         C     Y     P     E

         R    A     C     E

        A     E     E     Q

         U    I     S     E     T

         U     M

         1     0     0

         R    o     b     S    c    a     i     f    e     2     0     0     6

         6     0

         4     0

         2     0

         2     0

         6     0

         4     0

         2     0

         %

         S    u    m

         2     0

         4     0

         2     0

         8     0

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         2     0     4     0

         2     0     0     4     0     0     6     0     0     8     0     0     1     0     0     2     0     0     3     0     0     4     0     0

         %

         S    u    m

        +     M    a    r    s     h

         %

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         F    I     E     D

        /     D     E     G

         R    A     D     E

         D

         P     T     E

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         R      W     O

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        A     S     T     R

         U     M     P     R

         E   -     Q

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         R     N    A     R

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         T     R     E     E

         S    /     S     H

         R     U    B     S

        /     H     E     R

        B     S

         M    A     R

         S     H

         S     P     O

         R     E     S   M

        I     S     C .

         T     O     T    A

       L     P     O

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