contribution to knowledge of the balkan lepidoptera ii

Ecologica Montenegrina, 42, 2021, 1-44

Contribution to knowledge of the Balkan Lepidoptera II

(Lepidoptera: Macrolepidoptera)

STOYAN BESHKOV1 & ANA NAHIRNIĆ-BESHKOVA2

1, 2: National Museum of Natural History, Bulgarian Academy of Sciences, Tsar Osvoboditel Blvd.1, 1000 Sofia,

Bulgaria. E-mails: corresponding author: [email protected], [email protected]

Received 4 Aprl 2021 │ Accepted by V. Pešić: 26 April 2021 │ Published online 6 May 2021.

Abstract

Perizoma obsoletata (Herrich-Schäffer, 1838) is shown to be a part of the Balkan fauna. Pabulatrix pabulatricula

(Brahm, 1791) is new for the Western Balkans. One genus and 12 species are new and 20 species are reported for the

second time for Albania. For Montenegro, one genus and eight species are new and two species are recorded for the

first time. One genus and four species are new for the Republic of North Macedonia and another five species are

reported for the second time from that country. One genus and three species are new for Bosnia and Herzegovina. Four

species are reported for the second time and two for the third time for Serbia. Localities for Charissa pfeifferi (Wehrli,

1926) in Bulgaria are recorded for the first time. Data for some other interesting species from these countries and

Greece are also presented. Most specimens were collected at light in 2019. Illustrations of specimens and their genitalia

are given when necessary. Everted vesicas of some Geometridae species are illustrated for the first time.

Key words: butterflies, moths, Europe, Balkan countries, faunistic.

Introduction

The Balkan countries are explored unequally in terms of Macrolepidoptera, especially for nocturnal species.

It is difficult to find new taxa for countries such as Bulgaria and Greece, while Albania still remains perhaps

the most poorly investigated one. This is demonstrated here in that such a widespread and common species

as Orthosia cruda ([Denis & Schiffermüller], 1775) is reported as a new for Albania, and some other

common species recorded here have been reported only from a single locality in that country. In Montenegro

there is very incomplete information on occurrence of autumn and early spring species. We show this by

reporting here several common autumn species, which we were surprised to find, had not been previously

found in Montenegro. While Durmitor National park, Sea coast and other tourist areas are relatively well

explored, there is a lack of data for the other parts of that country. The situation in Bosnia and Herzegovina

is similar. The nocturnal Lepidoptera fauna of the Republic of North Macedonia is better explored, but

mainly by foreigners and the majority of the data are from only a few localities, while for large areas of the

country there is a lack of data. Finding a new species for that country is likely possible among difficult

groups and large genera, where examination of genitalia is necessary. Recording new localities of species

reported before from a single locality is, however, easy. Almost the same situation pertains in Serbia, where

Ecologica Montenegrina 42: 1-44 (2021)

This journal is available online at: www.biotaxa.org/em

http://dx.doi.org/10.37828/em.2021.42.1

http://www.biotaxa.org/em

BALKAN LEPIDOPTERA II

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in the last six years both of the present authors have published separately or together more than 60

Macrolepidoptera species new for this country.

Material and Methods

Collecting involved two portable light traps with an 8 watt actinic (368 nm) and 8 watt “Blacklight”, both

powered by 12 volt 9Ah batteries, as well as a Finnish “tent trap” with a 160 watt MV bulb at the top of the

pole and a 20 watt (368 nm) black light over the catching pot below. An additional 20 watt (368nm) lamp

was also positioned about 70 from the tent trap. The distance between the Finnish “tent trap” and the light

traps, as well as between the light traps themselves was sometime more than 1 km, as they were deployed in

different positions and habitats wherever possible. All traps ran throughout the night. All of the reported

material was collected by S. Beshkov & A. Nahirnić, so the names of the collectors are not mentioned for

each species.

Genitalia were fixed on glass slides in Euparal mountant after staining with a 2% Merbromin

solution. All genitalia slides were photographed by S. Beshkov with a trinocular Zeiss stereo microscope

Stemi 2000-C with axioCameRc 5s camera; solitary everted vesicas were photographed in alcohol before

mounting on glass. Genitalia slides prepared in 2021 were photographed by using self-modified transmitted

light. Insects and collecting localities were photographed with a Sony DSChX400v digital camera. Scale bar

is 1 cm. All trips were self-financed by the authors and undertaken in their own time.

Lepidoptera families are arranged according to Aarvik et al. (2017). Sequence and nomenclature of

families Erebidae and Noctuidae follow Fibiger et al. (2011) with subsequent changes incorporated from

recent taxonomic revisions. The Geometridae are arranged according to Hausmann & Sihvonen (2019).

When the specimen’s number is indicated, this means that they are in the collection of Stoyan Beshkov in the

National Museum of Natural History, Sofia (NMNHS). For the majority of the reported species the list of the

known localities is given, although some of the old reports for some problematic genera are not reliable

enough for inclusion. Faunistic records for some representatives of the genera Chasissa Curtis, 1825 and

Perizoma Hübner, 1825 before the last treatment of Müller et al. (2019) are not taken into account here.

Systematics

Limacodidae

Hoyosia cretica (Rebel, 1906) Greece, Amphissa district, NW of Eleonas Village, 691m, N38.559384°,

E022.387685°, 15.V.2016, Quercus coccifera, Cupressus, Juniperus on limestone area, 2 males at lamps and

light traps (Fig. 1). In this area H. cretica is known from the surroundings of Arachova and Delfi (Dufay,

1982). The nearest localities are in Thesalia, Peloponnesus, Thrace and the European part of Turkey

(Dardanelles) (de Freina & Witt, 1990).

Nymphalidae

Erebia gorge albanica Rebel, 1917 Albania, Mt. Tomor, Tomor peak (Abaz Aliu peak), 2379m, N40.6361°,

E20.1615°, eroded spots in high mountain dry grassy slopes, 15.VII.2018 (Fig. 2: a-f). The collecting locality

is illustrated in Beshkov & Nahirnić (2019b). Erebia gorge albanica Rebel is very remarkable and confusion

with other taxa is hardly believable. Specimens from Tomor have very small, in some specimens even

indistinct, ocelli. Wings in the males are very dark with almost indistinct pattern on both underside and

upperside. For comparison we illustrate two of the syntypes (Fig. 2: g-l) from North Albanian Alps (T.L:

Gropa e Shtrelit [Maja Stogut, Qafa e Strogut, north of Theth]) (Fig. 2: g-i) (Rebel, 1917a) and AL, Kakinja

[Maja e Kakisё], 2310m, 14.VII.1916 (Fig. 2: j-l) (source © Lepidoptera Image Collection, Natural History

Museum Vienna, H. Bruckner). We have not seen females from Tomor, therefore will not comment on

differences between the populations. According to Warren (1936): “There can never be any difficulty in

distinguishing albanica from any other gorge race; indeed, the difficulty is to believe that it belongs to the

same species. The specimens illustrated were lent to me by the captor, who stated that Rebel himself had

identified them as albanica, but that he found it hard to believe they were a race of gorge. The genitalia leave

BESHKOV & NAHIRNIĆ-BESHKOVA

Ecologica Montenegrina, 42, 2021, 1-44 3

no room for doubt, however…”. We also examined the genitalia (Gen. prep. 1./26.IV.2019, S. Beshkov) of a

single male specimen (Fig. 3). The specimens discussed and illustrated by Warren (1936) were from an

isolated mountain, Mali i Tomorrit in S. Albania and might show different external characters to E. gorge

albanica (Curvier et al., 2018). From Tomori E. gorge albanica is reported also by Rebel & Zerny (1931),

Micevski et al. (2015), Šašić et al. (2015) and Curvier at al. (2018), but it has been illustrated only in

monochrome in Warren (1936). Here Erebia gorge albanica is illustrated for the first time in colour.

According to Curvier et al. (2018) “collected males and females in five different localities and a direct

comparison of the collected specimens from these localities with specimens collected on the summit of Mali

i Tomorrit shows important variability within a single locality in the size of the ocelli and the width of the

orange band, but there is no obvious difference between populations from the N. Albanian Alps and Mali i

Tomorrit in S. Albania”. We would like to mentioned that not only the size of the ocelli but also the

completely rounded shape of the hind wings are differentiating characters against neighbouring populations

in the Dinaric mountains, e.g. Durmitor, Prenj, but also against E. gorge pirinica Buresch, 1918 from Pirin

and Rila (in latter populations with frequency of f. triopes.

Figure 1. Hoyosia cretica, male. GR, Eleonas 15.V.2016.

Erebia medusa ([Denis & Schiffermüller], 1775) Albania, Mt. Tomor, Tomor peak (Abaz Aliu peak),

2379m, N40.6361°, E20.1615°, 15.VII.2018. The illustrated male specimen (Fig. 4: a-b) has a reduced

number and small ocelli, and the orange band of the hindwings is restricted to very elongate spots in the

cells.

Erebia oeme (Hübner, [1804]) W Serbia, Javor Mts, below Vasilin Vrh near Lepojevići Village, 1489m,

N43.44140°0, E020.051601°, 02.VII.2019, meadows on limestone area with Juniperus communis near

Fagus forest. Genitalia checked.

Coenonympha orientalis Rebel, 1910 Albania, Korçё County, above Drenovё Village, 1050m, N40.588342°,

E020.806429°, 18.5.2017, river valley with Salix on serpentine slopes with Artemisia alba, Acantholimon,

etc., 1 male (Fig. 4: c-d).


4

Figure 2. Erebia gorge albanica from Albania. a. male, upperside. Mt. Tomor, 2379m, 15.VII.2018; b. male, underside.

Mt. Tomor, 2379m, 15.VII.2018; c. male, upperside. Mt. Tomor, 2379m, 15.VII.2018; d. male, underside. Mt. Tomor,

2379m, 15.VII.2018; e. male, upperside. Mt. Tomor, 2379m, 15.VII.2018; f. male, underside. Mt. Tomor, 2379m,

15.VII.2018; g. female syntype, upperside. Gropa e Shtrelit, 1800m, 3.VII.1916; h. female syntype, underside. Gropa e

Shtrelit, 1800m, 3.VII.1916; i. Label of the syntype of female, g-h; j. male syntype, upperside. Kakinja [Maja e Kakisё],

2310m, 14.VII.1916; k. male syntype, underside. Kakinja [Maja e Kakisё], 2310m, 14.VII.1916; l. Labels of Erebia

gorge albanica male syntype, j-k.

Drepanidae

Cilix asiatica O. Bang-Haas, 1907 Serbia, Stol Mt., northern foothill, above Luka Village, 807m,

N44.18334°, E022.13983°, 05.VI.2020, limestone slopes above Fagus forest, 1 female (Fig. 4: e). This is the

second report for Serbia after Beshkov & Nahirnić (2016) and the most northern one in the Balkans. North

Macedonia, Ohrid region, Petrina Planina - Galichica Mts, between Ohrid and Velestovo Village, 1005m,

N41.090632°, E020.827265°, 09.VI.2018, 1 male. In North Macedonia known from Strumica (Zolotuhin,

1999) and Demir Kapija surroundings (Beshkov et al., 2020b). Third report for North Macedonia. Albania,

Prespa Lake, near Pustec (=Liqenas) Village, 862m, N40.774238°, E020.903649°, 10.VII.2018, 1 male;

Korçë County, Mt. Ivan above Zvezdë Village on the road to Prespa Lake, 1088m, N40.733060°,



E020.880353°, 07.VI.2016, 1 male; Korçë County, Maliq municipality, above Klocë Village, near the radio

antenna, 1217m, N40.683921°, E020.694505°, 04.VII.2016, 2 females. In Albania known from Shkallё

(Beshkov & Nahirnić, 2018b), Vukopoles and Prespa Lake (Beshkov & Nahirnić, 2019b) and Boboshticë

(Beshkov et al., 2020b).

Figure 3. Erebia gorge albanica, male genitalia. Tomori, 2379, 15.vii.2018, Gen. prep. 1./26.IV.2019.

Geometridae

Geometrinae

Hemistola chrysoprasaria (Esper, 1795) Albania, Butrint Lake near the opening to the Sea, 18m,

N39.746394°, E019.996994°, 14.V.2017, and 02.VI.2019; Dajt Mt, Shkallë, 893m, N41.330302°,

E019.965265°, 29.VI.2017; Prespa Lake, near Pustec (=Liqenas) Village, 862m, N40.774238°,

E020.903649°, 10.VII.2018. Reported from Bicaj, Tirana, Shkodra and Miloti as Hemistola vernaria Hb.

(Rebel & Zerny, 1931), Uji Ftohte near Tepelene (Urbahn, 1966) as Hemistola immaculata Thunbg., from

Tirana, Marikaj, Burrel and Mbas Dejë (Misja, 1977) and from Mal Çajup (Beshkov et al., 2020b).


6

Eucrostes indigenata (Villers, 1789) Montenegro, Morača River Gorge, Milunovići Village N of Bioče,

160m, N42.5544°, E019.3299°, limestone slopes with Paliuris spina-christi, Quercus, Pistacia, etc., some

parts after fire, 04.VII.2019, 1 male (Fig. 4: f). New genus and new species for Montenegro, not represented

in Fauna Europaea website https://fauna-eu.org/cdm_dataportal/taxon/5fec94a5-7a91-4f64-bc16-

99e65cf06e49

Figure 4. a. Erebia medusa, male, upperside. AL, Mt. Tomor, 2379m, 15.VII.2018; b. Erebia medusa, male, underside.

AL, Mt. Tomor, 2379m, 15.VII.2018; c. Coenonympha orientalis, male, upperside. AL, Drenovё, 18.V.2017; d.

Coenonympha orientalis, male, underside. AL, Drenovё, 18.V.2017; e. Cilix asiatica, female. SRB, Stol, 5.VI.2020; f.

Euchrostes indigenata, male. MNE, Milunovići, 04.VII.2019; g. Narraga tessularia, male. SRB, Moklište, 29.VI.2020;

h. Pungeleria capreolaria, female. AL, Drenovё, 24.IX.2017; i. Charissa pullata, female. AL, Lurё, 09.VII.2019; j.

Charissa glaucinaria, female. AL, Lurё, 09.VII.2019; k. Charissa glaucinaria, female genitalia of 4: j. AL, Lurё,

09.VII.2019, Gen. prep. 1./24.III.2020; l. Charissa glaucinaria, male. AL, Munella, 1400m, 07.VII.2019; m. Charissa

glaucinaria, male. AL, Munella, 1400m, 07.VII.2019; n. Charissa glaucinaria, male genitalia of 4: l. AL, Munella,

1400m, 07.VII.2019, Gen. prep. 2./19.III.2020.

https://fauna-eu.org/cdm_dataportal/taxon/5fec94a5-7a91-4f64-bc16-99e65cf06e49

https://fauna-eu.org/cdm_dataportal/taxon/5fec94a5-7a91-4f64-bc16-99e65cf06e49



Ennominae

Dicrognophos sartata (Treitschke, 1827) Albania, Berat County, Vukopoles between Ibrolara and Vale,

Poliçan surroundings, 217m, N40.560005°, E020.093927°, 16.X.2016. Known from Vlora (Rebel, 1913),

Shkodra, Rogozhina, Elbasan, Durres, (Rebel & Zerny, 1931), Elbasan (Moucha, 1963), Butrint (Misja,

1990), Kakarriqi near Lezha, Limaçi, Dhermi (Beshkov & Misja, 1995), Zogaj (Beshkov, 1995).

Narraga tessularia (Metzner, 1845) Serbia, Svrljiške planine Mts, Bela Palanka District, above Moklište

Village, 616m, N43.27566°, E022.29339°, 29.VI.2020 (Fig. 4: g), limestone slopes with scattered pine trees

and Artemisia alba, 4 males at light; Serbia, Suva Planina Mt., Preslap, Divna Gorica above Bela Palanka

Town, 1186m, N43.19473°, E022.24400°, 30.VI.2020, dry limestone rocky slopes with Artemisia alba

above Fagus forest, 1 male at light. In Serbia N. tessularia is known from Crni Vrh (Vidlič Mt.) (Beshkov,

2017; Jakšić, 2017) and Šljivovički Vis (Jakšić, 2018). All localities are in eastern Serbia. Third locality in

Serbia. Narraga tessularia occurs in the more northern areas (Hungary, Austria) in alkaline habitats (ssp.

kasyi), not in calcareous ones.

Gerinia honoraria ([Denis & Schiffermüller], 1775) Albania, Berat County, between Ibrolara and Vale,

Poliçan district, 217m, N40.560005°, E020.093927°, 16.X.2016; Ionian Sea Coast, below Ilias, Dhermi

district, St. Theodor monastery, 140m, N40.131134°, E019.655874°, 15.V.2017, 2 males and 1 female; ibid,

03.VI.2019, 3 males; ibid, 153m, N40.131131°, E019.655340°, 17.X.2016, 2 males and 1 female; Delvina

Region, between Bistricё Village and Syri i Kaltёr, 127m, N39.931506°, E020.153775°, 13.V.2017, 1 male.

Reported from Vlora (=Valona) (Rebel, 1913; Rebel & Zerny, 1931) and Butrint (Misja, 1977; Beshkov et

al., 1996).

Pungeleria capreolaria ([Denis & Schiffermüller], 1775) Albania, Korçё County, above Drenovё Village,

1050m, N40.588342°, E020.806429°, 24.IX.2017, 1 female (Fig. 4: h). In Albania reported from Bështriq

(Rebel & Zerny, 1931) and Llogara (Misja, 1990). Third report for Albania.

Charissa obscurata ([Denis & Schiffermüller], 1775) Albania, Mali i Thatё Mt., above Korita Village,

1485m, N40.809441°, E020.834774°, 20.VIII.2017; Prespa Lake, near Pustec (=Liqenas) Village, 862m,

N40.774238°, E020.903649°, 19.VIII.2017; Tirana County, Malësia e Tiranës, (Highlands of Tirana), below

Maja e Fekenit (=Feken peak), west from Qafa e Selitës pass, 1133m, N41.3729°, E019.9992°, 15.VII.2018;

Korçё County, Mt. Kuq, Qarrit pass, west from Pepellash Village, 1318m, N40.481986°, E020.690849°,

22.VIII.2017. Known from Ploshtan and Qafa e Droshrës (Rebel & Zerny, 1931). Second report for Albania.

Charissa pullata ([Denis & Schiffermüller], 1775) Albania, Korçё County, Krushovё, above Voskopojё,

1296m, N40.6706°, E020.5674°, 11.VII.2018, serpentinite slopes with Buxus sempervirens, Pinus nigra,

Juniperus, etc.; above Kruja, 1134m, N41.5153°, E019.8039°, 18.VII.2018, limestone rocks and Pinus nigra

trees; Tirana County, Dajt Mt, near Shkallё Village, 893m, N41.330302°, E019.965265°, 29.VI.2017; Tirana

County, Malësia e Tiranës, (Highlands of Tirana), below Maja e Fekenit (=Feken peak), west from Qafa e

Selitës pass, 1133m, N41.3729°, E019.9992°, 15.VII.2018; Lurё Mt., above Fushё-Lurё Village, 1687m,

N41.7767°, E020.1989°, 09.VII.2019 (Fig. 4: i); Munella Mt., above Kimez Village, 1014m, N41.9710°,

E020.0686°, 06.VII.2019. Reported from Oroshi (Rebel, 1913), Bështriq and Trektani (Rebel & Zerny,

1931), Bizë (Urbahn, 1966), Qarrishtë, Berat, Lure, Balgjaj (Misja, 1977). Serbia, Kraljevo Region, Stolovi

Mt., Veliki Čukar, 688m, N43.601922°, E020.685631°, 01.VII.2019, S. Beshkov, A. Nahirnić, P. Jaksić, C.

Plant, A. King leg.

Charissa variegata (Duponchel, 1830) Montenegro, between Danilovgrad and Nikšić, Bogmilovići Village,

468m, N42.613625°, E019.013915°, 30.VI.2017, limestone slopes with Juniperus oxycedrus, Punica

granatum, Fraxinus ornus, Acer monspesullanum, Pistacia terebinthus, Thalictrum, Paliurus spina-christi;

Albania, Prespa Lake, near Pustec (=Liqenas) Village, 878m, N40.774171°, E020.902553°, 18.X.2017.

Known in Albania from Oroshi, Pasha Liman (Rebel, 1913), Kula e Lumës, Bicaj and Shkodra (Rebel &

Zerny, 1931), Zvezdë, Mal Çajup and Shkallë (Beshkov et al., 2020b). The report by Drenowski (1930) is

for the Republic of North Macedonia, not for Albania.


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Charissa ambiguata (Duponchel, 1830) Albania, Korçё County, Moravё Mt., above Dishnicё village,

1564m, N40.647811°, E020.857765°, 25.VI.2017; ibid, Dardhё, 1276m, N40.526116°, E020.825850°,

26.VI.2017. New species for Albania.

Charissa onustaria (Herrich-Schäffer, 1852) Albania, Mali i Thatё Mt., above Korita Village, 1485m,

N40.809441°, E020.834774°, 20.VIII.2017. Known from Kula e Lumës (Rebel & Zerny, 1931) and from

Pepellash (Beshkov et al., 2020b). The report by Drenowski (1930) is for the Republic of North Macedonia,

not for Albania. Third report for Albania.

Charissa glaucinaria (Hübner, 1799) Albania, Lurё Mt., above Fushё-Lurё Village, 1687m, N41.7767°,

E020.1989°, 09.VII.2019, 1 female (Fig. 4: g), Gen. prep. 1./24.III.2020, S. Beshkov on glass in euparal

(Fig. 4: k); Munella Mt., above Kimez Village, 1400m, N41.9491°, E020.0926°, 07.VII.2019, 2 males (Fig.

4: l-m), Gen. prep. 2./19.III.2020, S. Beshkov, male genitalia on glass in euparal (Fig. 4: n); Bjeshkët e

Nemuna Mts (=Prokletije Mts), Vermosh surroundings, Gropat e Selcёs, 1231m, N42.5405°, E019.6960°,

16.VIII.2018, genitalia checked, in microvial in glycerol. The reports by Rebel (1913) for Oroshi and Fandi

and by Rebel & Zerny (1931) from Kruma and Kula e Lumës perhaps concern another species, more likely

Ch. supinaria (Urbahn, 1966). Reported also by Misja (1977) from Qarrishtë and by Misja (1990) from Qafa

e Bordolecit but correct identification is questionable. In Albania Ch. glaucinaria was previously surely

proved only from Tomori, 2379m (Beshkov et al., 2020a). Here it is reported for a second time for Albania.

According to Müller et al. (2019) in the Balkan Peninsula Ch. glaucinaria occurs only in southern Bulgaria

and Greece. That is why in Müller et al. (2019) all previous reports for Montenegro (e.g. of Tomić et al.

(1990) and for North Macedonia are omitted as doubtful. Montenegro, Durmitor Mt., Nadgorje, 1735m,

N43.192505°, E019.044193°, 06.VIII.2015 (Fig 5: a-b), Gen. prep. 1./11.I.2021, S. Beshkov, male genitalia

on glass in euparal (Fig. 5: c). New species for Montenegro. North Macedonia, Pelister Mt., mountain house

"Shiroka", 1955m, N41.004735°, E021.168664°, 06.VIII.2016, 1 female (Fig. 5: d-e), Gen. prep.

1./11.V.2020, S. Beshkov (Fig. 5: f), on glass in euparal. New species for Northern Macedonia. Bulgaria, S

Pirin Mts, below Orelek Top, 1900m, N41.560062°, E023.616683°, 06.VIII.2010, 1 female (Fig. 5: g-h),

xeromontane area, S. Beshkov leg., genitalia checked, in microvial in glycerol.

Charissa pfeifferi (Wehrli, 1926) Greece, Peloponessos, Chelmos Mts, Kalavrita region, above Peristera,

1294m, N38.027607°, E022.240351°, 14.V.2016 (Fig. 6: a-b); ibid, between Kalavrita and Peristera, below

Gaidhourorachi summit, 1776m, N38.029995°, E022.218370°, 11.V.2017, 5 males (Fig. 6: c) and 7 females,

Gen. prep. 2./23.III.2020 S. Beshkov, male genitalia on glass in euparal (Fig. 6: d). Bulgaria, Eastern

Rhodopi Mts, Arda river valley, between Rabovo and Studen Kladenets villages, 173m, N41.609181°,

E025.650038°, 21.IV.2018 (Fig. 6: e-f), rocky river valley with Paliurus spina-christi, Gen. prep.

1./09.XII.2020 S. Beshkov, female genitalia on glass in euparal (Fig. 7: a); Eastern Rhodopi Mts, Arda river

valley, the bridge on Arda river near Madzharovo Town, 150m, N 41.643625°, E 025.872209°, 30.IV.2005,

S. Beshkov & D. Vassilev leg., 1 female (Fig. 7: b-c); Eastern Rhodopi Mts, below Komuniga Village,

494m, N41.796116°, E025.201705°, 14.V.2018, S. Beshkov & B. Zlatkov leg., 1 male. In Europe Ch.

pfeifferi is known only from Greece and Bulgaria (Müller et al., 2019) and recently it was reported also from

North Macedonia: Prilep (Tóth, 2019). In Bulgaria at present it is recorded only from Western and Eastern

Rhodopi Mts. Here Ch. pfeifferi is reported for the first time from Bulgaria with localities.The report of

Charissa intermedia (Wehrli, 1917) in Beshkov & Langourov (2011: 335, figs 8, 9) represents Ch. pfeifferi.

Furthermore, the reports of Charissa intermedia in Beshkov & Langourov (2004) and Ch. glaucinaria in

Beshkov & Langourov (2011) perhaps concern Ch. pfeifferi.

Charissa supinaria (Mann, 1854) Albania, Shkodra County, Vau i Dejës, 333m, N42.00283°, E019.68944°,

05.X.2019 (Fig. 7: d), Gen. prep. 2./23.4.2020, S. Beshkov, female genitalia on glass in euparal (Fig. 7: e);

Tirana County, Dajt Mt., near Shkallë village, 893m, N41.330302°, E019.965265°, 08.VI.2018, 1 female,

genitalia checked; Tirana County, Mali me Gropa Mts, above Shen Meri Village, 1405m, N41.3524°,

E020.0465°, 13.VIII.2018, one male, genitalia checked; ibid, 05.VI.2019 (Fig. 7: f), Gen. prep.

1./23.IV.2020, S. Beshkov, male genitalia on glass in euparal (Fig. 7: g); Mt. Thanës, near Bulqizë town,

above Plani i Bardhë village, N41°28'35", E020°09'18", 788m, 17.VIII.2018 (Fig. 8: a-d), Gen. prep.

3./23.IV.2020, S. Beshkov, male genitalia on glass in euparal (Fig. 8: e); ibid, 6.IV.2019 (Fig. 8: f, Fig. 9: a);

Korçё County, below Dardhё, above Boboshticë Village, 1225m, N40.540589°, E020.791964°, 19.V.2017,



10 males and two females, male genitalia checked; ibid, 04.VI.2018, 2 males and 1 female, male genitalia

checked; Tirana County, Dajti Mt, Qafa e Mollёs Pass, 665m, N41.3643°, E019.9657°, 04.IV.2019, 1

female, genitalia checked, in microvial in glycerol; Korçё County, Dardhё, 1276m, N40.526116°,

E020.825850°, 26.VI.2017, Korçё County, above Drenovё Village, 1050m, N40.588342°, E020.806429°,

18.V.2017; 1 male, genitalia checked; Bjeshkët e Nemuna Mts (=Prokletije Mts), Vermosh surroundings,

Gropat e Selcёs, 1231m, N42.5405°, E019.6960°, 16.VIII.2018, 1 female; Korçё County, Mt. Kuq, Qarrit

pass, west from Pepellash village, 1187m, N40.481719°, E020.6755930°, 05.VI.2018, 1 female. Reported

for Albania from Han i Grabom (Rebel, 1914), Kruma, Kula e Lumës, Zepë Mt., Oroshi, Fani (Rebel &

Zerny, 1931), Dajti: Shkall (Urbahn, 1966), Starë (Beshkov & Nahirnić, 2019a). Perhaps some of the old

reports for Ch. glaucinaria concern this species (see also under Ch. glaucinaria here). Montenegro, Gulf of

Kotor, Ledenice Village above Risan Town, 548m, N42.526652°, E018.705581°, 10.VIII.2015, abundant

(Fig. 9: b-h), Gen. prep. 1./18.III.2020, S. Beshkov, male genitalia on glass in euparal (Fig. 9: i). New

species for Montenegro.

Figure 5. a. Charissa glaucinaria, male, upperside. MNE, Durmitor, Nadgorje 6.VIII.2015; b. Charissa glaucinaria,

male, underside. MNE, Durmitor, Nadgorje 6.VIII.2015; c. Charissa glaucinaria, male genitalia of 5: a, b MNE,

Durmitor, Nadgorje 6.VIII.2015, Gen. prep. 1./11.I.2021; d. Charissa glaucinaria, female, upperside. RNM, Baba Mt,

1955m, 06.VIII.2016; e. Charissa glaucinaria, female, underside. RNM, Baba Mt, 1955m, 06.VIII.2016; f. Charissa

glaucinaria, female genitalia of 5: d, e RNM, Baba Mt, 1955m, 06.VIII.2016, Gen. prep. 1./11.V.2020; g. Charissa

glaucinaria, female, upperside. BG, Pirin Mts, Orelek, 06.VIII.2010; h. Charissa glaucinaria, female, underside. BG,

Pirin Mts, Orelek, 06.VIII.2010.

Charissa graecaria (Staudinger, 1871) (=certhiatus Rebel & Zerny, 1931) Albania, Tirana County, Mali me

Gropa, Central part S of plateau, 1501m, N41.3617, E020.0568, 16.VII.2018; Tomorri Mts, above Ujanik, on

the road to Abaz Aliu Top, 1739m, N40.6111, E020.1885, 13.VII.2018 (Fig. 10: a); ibid, Abaz Aliu Top,


10

2379m, N40.6361°, E020.1615°, 14.VII.2018, abundant; Albania, Mali i Thatё Mt., above Korita Village,

1485m, N40.809441°, E020.834774°, 20.VIII.2017. In Albania known from Bështriq and Korab (Rebel &

Zerny, 1931) and from two localities in the mountains Tomori and Mali me Gropë (Beshkov et al., 2020a).

Third report from Albania. North Macedonia, Galichica Mts., between Dvata Javora and Bulgarska Chuka

Top, 1587m, N40.990906°, E020.857791°, 03.VII.2016; Pelister Mt, mountain house "Shiroka", 1955m,

N41.004735°, E021.168664°, 06.VIII.2016, 2 females, genitalia of one checked, in microvial in glycerol. In

North Macedonia known from Korab, Šar Planina and Petrina Planina mountains (Pinker, 1968). Second

report for North Macedonia.

Figure 6. a. Charissa pfeifferi, male, upperside. GR, Peloponessos, Chelmos Mts, 1294m, 14.V.2016; b. Charissa

pfeifferi, male, underside. GR, Peloponessos, Chelmos Mts, 1294m, 14.V.2016; c. Charissa pfeifferi, male, upperside.

GR, Peloponessos, Chelmos Mts, 1776m, 11.V.2016; d. Charissa pfeifferi, male genitalia of 6: c GR, Peloponessos,

Chelmos Mts, 1776m, 11.V.2016, Gen. prep. 2./23.III.2020; e. Charissa pfeifferi, female, upperside. BG, E Rhodopi,

Rabovo, 21.IV.2018; f. Charissa pfeifferi, female, underside. BG, E Rhodopi, Rabovo, 21.IV.2018.

Dyscia raunaria (Freyer, 1851) Albania, Lurё Mt., above Fushё-Lurё Village, 1687m, N41.7767°,

E020.1989°, 09.VII.2019, 1 male; Tirana County, Mali me Gropa Mts, above Shen Meri Village, 1405m,

N41.3524°, E020.0465°, 17.VII.2018, 3 males; Tomorri Mts, above Ujanik, on the road to Abaz Aliu Top,

1739m, N40.6111°, E020.1885°, 13.VII.2018 (Fig. 10: b). Known in Albania from Mali me Gropë (Urbahn,

1966), Tomori, 2026m (Beshkov & Nahirnić, 2019), Boboshticë and Shkallë (Beshkov et al., 2020a). Third

report for Albania. The reports by Rebel & Zerny (1931) and by Misja (1990) for Dyscia conspersaria

([Denis & Schiffermüller], 1775) perhaps concern Dyscia raunaria.

Dyscia innocentaria sicanaria (Oberthür, 1923) Albania, Mali i Thatё Mt., above Korita Village, 1485m,

N40.809441°, E020.834774°, 20.VIII.2017. In Albania known from Poliçan (Urbahn, 1966), Starë, Plani i

Bardhë (Beshkov & Nahirnić, 2019a), Zvezdë and Mal Çajup (Beshkov et al., 2020b). Serbia, Svrljiške



planine Mts, Bela Palanka District, above Moklište Village, 616m, N43.27566°, E022.29339°, 29.VI.2020.

In Serbia known from Jelašnička klisura, Niš Region (Beshkov, 2015), two localities in Pčinja River Valley

(Beshkov & Nahirnić, 2016), Trnava Village above Preševo and Šljivovički Vis above Bela Palanka

(Beshkov & Nahirnić, 2017).

Figure 7. a. Charissa pfeifferi, female genitalia of Fig. 6: e, f BG, E Rhodopi, Rabovo, 21.IV.2018, Gen. prep.

1/09.XII.2020; b. Charissa pfeifferi, female, upperside. BG, E Rhodopi, Madzharovo, 30.IV.2005; c. Charissa pfeifferi,

female, underside. BG, E Rhodopi, Madzharovo, 30.IV.2005; d. Charissa supinaria, female. AL, Shkodra, Vau i Dejës,

05.X.2019; e. Charissa supinaria, female genitalia of 7: d AL, Shkodra, Vau i Dejës, 05.X.2019, Gen. prep.

2./23.IV.2020; f. Charissa supinaria, male. AL, Mali me Gropa, 1405m, 05.VI.2020; g. Charissa supinaria, male

genitalia of 7: f AL, Mali me Gropa, 1405m, 05.VI.2020, Gen. prep, 1./23.IV.2020.

Menophra abruptaria (Thunberg, 1792) Albania, Delvina Region, between Bistricё Village and Syri i Kaltёr,

127m, N39.931506°, E020.153775°, 13.V.2017, 1 male, Gen. prep. 1./08.V.2020, S. Beshkov, male genitalia

on glass in euparal (Fig. 10: c); Butrint Lake, near the opening to the Sea, 18m, N39.746394°,

E019.996994°, 22.X.2017 (Fig. 10: d), Gen. prep. 2./08.V.2020, S. Beshkov, male genitalia on glass in

euparal (Fig. 10: e). Reported for Albania from Tirana, Lezha, Mamurras, Gomen near Kavaja, and

Mbasdeje (Misja, 1977), Llogara, Butrint (Beshkov et al., 1996). The report by Beshkov (1995) for Dhermi

concerns another species - Menophra japygiaria (O. Costa) (Beshkov & Nahirnić, 2020a).


12

Menophra japygiaria (O. Costa, 1849) Albania, Butrint Lake, near the opening to the Sea, 18m,

N39.746394°, E019.996994°, 22.X.2017 (Fig. 11: a), Gen. prep. 3./08.V.2020, S. Beshkov, male genitalia on

glass in euparal (Fig. 11: b); Ionian Sea Coast, near Dhёrmi Village, Vlora Region, 24.XI.1993 (Fig. 11: c),

S. Beshkov leg., Gen. prep. 1./19.III.2020, S. Beshkov, male genitalia on glass in euparal (Fig. 11: d), ID

number BC ZSM Lep 105218 in BOLD database. This specimen is reported erroneously by Beshkov (1995)

as M. abruptaria (Beshkov & Nahirnić, 2020a). In Albania and on the Balkan Peninsula M. japygiaria is

known only from above Stan-Karbunara Village (Lushnjё County), Ionian Sea Coast near Dhërmi Village

(Beshkov & Nahirnić, 2020a) and Ardenica (Beshkov & Nahirnić, 2019a). Third report for Albania.

According to Müller et al. (2019) M. japygiaria is a West Mediterranean species with distribution restricted

to Portugal, Spain, southernmost France to Italy including Sardinia, Sicily and Malta. Outside Europe it is

known from North Africa: Morocco to Tunisia. The Balkan Peninsula is not included in the supposed range

of the species. With these new findings in Albania M. japygiaria is proved as a part of the Balkan’s fauna.

Figure 8. a. Charissa supinaria, male, upperside. AL, Bulqizë, Plani i Bardhë, 17.VIII.2018; b. Charissa supinaria,

male, underside. AL, Bulqizë, Plani i Bardhë, 17.VIII.2018; c. Charissa supinaria, male, upperside. AL, Bulqizë, Plani

i Bardhë, 17.VIII.2018; d. Charissa supinaria, male, upperside. AL, Bulqizë, Plani i Bardhë, 17.VIII.2018; e. Charissa

supinaria, male genitalia of 8: d AL, Bulqizë, Plani i Bardhë, 17.VIII.2018, Gen. prep. 3./23.IV.2020; f. Charissa

supinaria, male, upperside. AL, Bulqizë, Plani i Bardhë, 06.VI.2018

Menophra berenicidaria (Turati, 1924) Greece, Peloponnessos, Kivari near Argos, 130m, N37.506116°,

E022.726087°, 22.IV.2009, S. Beshkov, J. Buszko & B. Zlatkov leg. (Fig. 11: e) Gen. prep. 1./29.XII.2020,

S. Beshkov, male genitalia on glass in euparal (Fig. 11: f).

Cleorodes lichenaria (Hufnagel, 1767) Albania, Korçё County, Mt. Ivan above Zvezdё Village on the road

to Prespa Lake, 1088m, N40.733060°, E020.880353°, 06.VII.2016, limestone slopes with Buxus

sempervirens and Carpinus orientalis (Fig. 11: g); Korçё County, Dardhё, 1276m, N40.526116°,

E020.825850°, 26.VI.2017; meadow near stream with Salix, Fagus forest and hill with Astracantha (Fig. 11:

h). In Albania reported from Bizё (Urbahn, 1966) and Vermosh (Misja, 1990). Third report for Albania.



Figure 9. a. Charissa supinaria, male, underside. AL, Bulqizë, Plani i Bardhë, 06.VI.2018; b. Charissa supinaria,

male, upperside. MNE, Risan, Ledenice, 10.VIII.2015; c. Charissa supinaria, male, underside. MNE, Risan, Ledenice,

10.VIII.2015; d. Charissa supinaria, female, upperside. MNE, Risan, Ledenice, 10.VIII.2015; e. Charissa supinaria,

female, upperside. MNE, Risan, Ledenice, 10.VIII.2015; f. Charissa supinaria, female, underside. MNE, Risan,

Ledenice, 10.VIII.2015; g. Charissa supinaria, female, upperside. MNE, Risan, Ledenice, 10.VIII.2015; h. Charissa

supinaria, female, underside. MNE, Risan, Ledenice, 10.VIII.2015; i. Charissa supinaria, male genitalia. MNE, Risan,

Ledenice, 10.VIII.2015, Gen. prep. 1./18.III.2020.

Lycia graecarius (Staudinger, 1861) Albania, Mt. Thanës, near Bulqizë town, above Plani i Bardhë Village,

833m, N41.4798°, E020.1534°, 06.IV.2019 (Fig. 12: a-b), slopes with grass vegetation and scattered

Carpinus, Quercus, Pinus trees. In Albania known from Tomor (Rebel & Zerny, 1931). Second report for

Albania.

Agriopis marginaria (Fabricius, 1776) Albania, Mt. Thanës, near Bulqizë town, above Plani i Bardhë

Village, 833m, N41.4798°, E020.1534°, 06.IV.2019, slopes with grass vegetation and scattered Carpinus,

Quercus, Pinus trees (Fig. 12: c). Reported from Tirana (Misja, 1977) and Llogara (Beshkov et al., 1996).

Third report for Albania.


14

Figure 10. a. Charissa graecaria, male. AL, Mt. Tomor, 1739m, 13.VII.2018; b. Dyscia raunaria, male. AL, Mt.

Tomor, 1739m, 13.VII.2018; c. Menophra abruptaria, male genitalia. AL, Bistricё - Syri i Kaltёr, 127m, 13.V.2017,

Gen. prep. 1./08.V.2020; d. Menophra abruptaria, male. AL, Butrint, 22.X.2017; e. Menophra abruptaria, male

genitalia of 10: d AL, Butrint, 22.X.2017, Gen. prep. 2./02.VIII.2020.

Fagivorina arenaria (Hufnagel, 1767) Albania, Tirana County, Malësia e Tiranës, (Highlands of Tirana),

below Maja e Fekenit (=Feken peak), west from Qafa e Selitës pass, 1133m, N41.3729°, E019.9992°,

15.VII.2018, 2 males; Munella Mt, above Kimez Village, 1400m, N41.9491°, E020.0926°, 07.VII.2019 (Fig.

12: d). Reported from Plani [Plan] (Rebel, 1917b as angularia Thunberg), Bështriq (Rebel & Zerny, 1931),

Dajti, Mali me Gropa, Bizё (Urbahn, 1966) and Qarrishtë (Misja, 1977).

Nychiodes waltheri Wagner, 1919 Bulgaria, Central Stara Planina Mts, Chamdzha reserve near Hristo

Danovo Village, 570m, N42.733079°, E024.593715°, 31.VII.2014, rocky slopes in river valey with

Carpinus, Quercus, Pinus nigra leg S. beshkov; Eastern Stara Planina Mts, between Tushovitsa and Rish

villages, 430m, N42.966125°, E026.895938°, 17.VIII.2011, Quercus forest and clearing with Populus

tremula, etc.; Sliven Region, Sotirya Village, 327m, N42.684230°, E026.412620°, 06.VI.2017, S. Beshkov,

J. Toth & J. Bereczki leg., 1 male, genitalia checked; Eastern Rhodopi Mts, below Komuniga Village, 494m,

N41.796116°, E025.201705°, 14.V.2018, S. Beshkov & B. Zlatkov leg. According to Beshkov & Langourov

(2004) N. waltheri occurs from central Bulgaria to the Eastern Rhodopes and the Black Sea Coast close to

the border with Romania in the North and previous reports for N. dalmatina from Rhodopi Mts, central and

eastern Bulgaria must concern Nychiodes waltheri.

Nychiodes dalmatina (Wagner, 1909) Albania, Butrint Lake near the opening to Sea, 18m, N39.746394°,

E019.996994°, 14.V.2017. Known from Kula e Lumës, Shkodra (Rebel & Zerny, 1931), Mali me Gropa

(Urbahn, 1966), Zogaj (Beshkov & Misja, 1995). Reported as Nychiodes lividaria Hubner from Tirana,

Marikaj, Kune, Vlora, Bistricë (Misja, 1977).



Figure 11. a. Menophra japygiaria, male. AL, Butrint, 22.X.2017; b. Menophra japygiaria, male genitalia of 11: a AL,

Butrint, 22.X.2017, Gen. prep. 3./08.V.2020; c. Menophra japygiaria, male. AL, Dhёrmi, 24.IX.1993; d. Menophra

japygiaria, male genitalia of 11: c AL, Dhermi, 24.IX.1993, Gen. prep. 1./19.III.2020; e. Menophra berenicidaria,

male. GR, Peloponnessos, near Argos, 22.IV.2009 Gen. prep. 1./29.XII.2020; f. Menophra berenicidaria, male genitalia

of 11: e GR, Peloponnessos, near Argos, 22.IV.2009, Gen. prep. 1./29.XII.2020; g. Cleorodes lichenaria, male. AL,

Zvezdё, 06.VII.2016; h. Cleorodes lichenaria, male. AL, Dardhё, 26.VI.2017.

Sterrhinae

Idaea moniliata ([Denis & Schiffermüller], 1775) Albania, Korçё County, Mt. Kuq, Qarrit pass, below


16

Pepellash village, 999m, N40.4708°, E020.6722°, 12.VII.2019, stony slopes with Juniperus, Pinus nigra,

Quercus spp. In Albania known from Kula e Lumës and Bicaj (Rebel & Zerny, 1931), Burrel, Mamurras and

Ujanik (Misja, 1990), Pepellash, Benjë Novoselë and Mal Çajup (Beshkov et al., 2020b). The report by

Drenowski (1930) is for the Republic of North Macedonia, not for Albania. Montenegro, between

Danilovgrad and Nikšić, Bogmilovići Village, 468m, N42.613625°, E019.013915°, 30.VI.2017.

Scopula tessellaria (Boisduval, 1840) Albania, Korçё County, below Dardhё Village, 1276m, N40.526116°,

E020.825850°, 26.VI.2017 (Fig. 12: e). Known from Kishajt (Rebel & Zerny, 1931). Second report for

Albania.

Rhodostrophia calabra (Petagna, 1787) Bosnia & Herzegovina, Sarajevo Region, above Konjic, 652m,

N43.647227°, E017.987513°, 02.VII.2017, dolomites with Carpinus, Pinus, Acer, Quercus, 2 males (Fig. 12:

f).

Figure 12. a,-b. Lycia graecarius, male. AL, Bulqizë, Plani i Bardhë, 06.IV.2019; c. Agriopis marginaria, male. AL,

Bulqizë, Plani i Bardhë, 06.IV.2019; d. Fagivorina arenaria, male. Munella, 1400m, 07.VII.2019; e. Scopula

tessellaria, male. AL, Dardhё, 26.VI.2017; f. Rhodostrophia calabra, male. BiH, Bosnia, above Konjic, 02.VII.2017; g.

Chesias rufata, female. AL, Bulqizë, Plani i Bardhë, 06.IV.2019; h. Scotoptereyx ignorata, male genitalia with everted

veica. AL, Voskopojё, Krushovё 11.7.2018, Gen. prep. 2./21.IV.2020; i. Scotoptereyx ignorata, female genitalia. AL,

Voskopojё, Krushovё 11.7.2018, Gen. prep. 1./21.IV.2020.



Larentiinae

Chesias rufata (Fabricius, 1775) Albania, Mt. Thanës, near Bulqizë town, above Plani i Bardhë Village,

833m, N41.4798°, E020.1534°, 06.IV.2019 (Fig. 12: g), slopes with grass vegetation and single Carpinus,

Quercus, Pinus nigra trees. New genus and new species for Albania.

Scotopteryx ignorata (Huemer & Hausmann, 1998) Albania, Korçё County, Krushovё, above Voskopojё,

1296m, N40.6706°, E020.5674°, 11.VII.2018, serpentene slopes with Buxus sempervirens, Pinus nigra,

Juniperus, etc., gen. prep. 2./21.iv.2020, S. Beshkov, male genitalia with everted vesica (Fig. 12: h) and gen.

prep. 1./21.4.2020, S. Beshkov, female genitalia (Fig. 12: i). New species for Albania. Vesica has never been

everted and illustrated before.

Scotopteryx bipunctaria ([Denis & Schiffermüller], 1775) Albania, Tirana County, Mali me Gropa Mts,

1405m, N41.3524°, E020.0465°, 17.VII.2018 (Fig. 13: a), gen. prep. 1./23.III.2020, S. Beshkov, male

genitalia with everted vesica (Fig. 13: b). Vesica has never been everted and illustrated before. Known in

Albania from Bështriq, Ploshtan, Korab and Tomor (Rebel & Zerny, 1931), Mali me Gropa, Lure (Urbahn,

1966), Goricë, Qarrishtë, Vermosh and Butrint (Misja, 1977). The last locality for this species seems likely

to be incorrect because is on the sea coast. The report by Rebel & Zerny (1931) for Plav concerns

Montenegro.

Scotopteryx olympia Rezbanyai-Reser, 2003 Albania, Munella Mt, above Kimez Village, 1400m,

N41.9491°, E020.0926°, 07.VII.2019 (Fig. 13: c-e), gen. preps 3./19.III.2020 (Fig. 14: a) and 1./20.III.2020,

S. Beshkov, male genitalia with everted vesica (Fig14: b). New species for Albania. Vesica (Fig. 14: c-f) has

never been everted and illustrated before. It is very possible that Ortholithia octodurensis Favre, 1903 and

Ortholithia octodurensis f. obscura Urbahn, 1966 reported for Bizë by Friese (1967) and O. octodurensis f.

obscura from Urbahn (1966) concern Scotopteryx olympia, an endemic species for the Balkan Peninsula.

Catarhoe putridaria (Herrich-Schäffer, 1852) Montenegro, between Danilovgrad and Nikšić, Bogmilovići

Village, 468m, N42.613625°, E019.013915°, 30.VI.2017, limestone slopes with Juniperus oxycedrus,

Punica granatum, Fraxinus ornus, Acer monspesullanum, Pistacia terebinthus, Thalictrum, Paliurus spina-

christi, 1 female; Morača River Gorge, Milunovići Village, N of Bioče, 160m, N42.5544°, E019.3299°,

04.VII.2019, limestone slopes with Paliuris spina-christi, Quercus, Pistacia, etc. after fire, 1 female (Fig. 15:

a). In Montenegro known from Stari Bar (Pinker, 1968). Second report for Montenegro.

Catarhoe rubidata ([Denis & Schiffermüller], 1775) Albania, Tirana County, Dajti Mt, Qafa e Mollёs Pass,

665m, N41.3643°, E019.96547°, 04.VI.2019, 1 female (Fig. 15: b). Collected about 60 years ago from Mali

me Gropa (Urbahn, 1966). Second report for Albania.

Euphyia adumbraria (Herrich-Schäffer, 1852) Serbia, Suva Planina, Preslap, Divna Gorica above Bela

Palanka Town, 1186m, N43.19473°, E022.24400°, 30.VI.2020, dry limestone rocky slopes with Artemisia

alba above Fagus forest, 3 males (Fig.15 c-d), Gen. prep. 2./22.I.2021, S. Beshkov, male genitalia on glass

in euparal (Fig. 15: e). Second report for Serbia, previously known only from Savina Voda in Western Serbia

(Beshkov, 2020).

Euphyia frustata (Treitschke, 1828) Albania, Munella Mt, above Kimez Village, 1400m, N41.9491°,

E020.0926°, 07.VII.2019, 1 female (Fig. 15: f), genitalia checked because of the unusual coloration.

Anticlea derivata ([Denis & Schiffermüller], 1775) Albania, Korçё County, above Boboshticё Village,

1225m, N40.540589°, E020.791964°, 19.V.2017; Tirana County, Dajti Mt, Qafa e Mollёs Pass, 665m,

N41.3643°, E019.9657°, 04.IV.2019; Albania, Mt. Thanës, near Bulqizë town, above Plani i Bardhë village,

N41.4798°, E020.1534°, 833 m, 06.IV.2019. In Albania known only from Llogara (Beshkov et al., 1996).

Second report for Albania.

Eulithis prunata (Linnaeus, 1758) Albania, Prespa Lake, near Pustec (=Liqenas) Village, 849m,

N40.770632°, E020.908891°, 10.VII.2018, 1 male (Fig. 15: g). New species for Albania.


18

Figure 13. a. Scotopteryx bipunctaria, male. AL, Mali me Gropa Mts, 1405m, 17.VII.2018; b. Scotopteryx bipunctaria,

male genitalia with everted vesica. AL, Mali me Gropa Mts, 1405m, 17.VII.2018 Gen. prep. 1./23.III.2020; c-e.

Scotopteryx olympia, males. AL, Munella, 1400m, 07.VII.2019.

Ecliptopera silaceata ([Denis & Schiffermüller], 1775) North Macedonia, Pelister Mt., on the road to

Shiroka, 1493m, N41.031106°, E021.195290°, 02.VII.2016, clearings and small meadow in coniferous

forest, 1 male (Fig. 16: a). New genus and new species for North Macedonia.

Dysstroma truncata (Hufnagel, 1767) Albania, Tirana County, Mali me Gropa Mts, 1405m, N41.3524°,

E020.0465°, 05.VI.2019 (Fig. 16: b-c). Known from Qafa e Bordolecit (Misja, 1990). Second report for

Albania.

Horisme calligraphata (Herrich-Schäffer, 1852) Albania, Shkodra County, Bjeshkët e Nemuna Mts

(=Prokletije Mts), Radohima Mt., between Qafa e Thorës pass and Theth village, south-east of Shtegu peak,

1657m, N42.3854°, E019.7502°, 19.VII.2018. Known from Dajti (Urbahn, 1966) and Zvezdë (Beshkov et

al., 2020b). Third report for Albania.



Figure 14. a. Scotopteryx olympia, male genitalia. AL, Munella, 1400m, 07.VII.2019, Gen. prep. 3./19.III.2020; b.

Scotopteryx olympia, male genitalia with everted vesica. AL, Munella, 1400m, 07.VII.2019, Gen. prep. 1./20.III.2020;

c-f. Scotopteryx olympia, everted vesica. AL, Munella, 1400m, 07.VII.2019, Gen. prep. 1./20.III.2020.

Perizoma alchemillata (Linnaeus, 1758) Albania, Tirana County, Malësia e Tiranës, (Highlands of Tirana),


15.VII.2018. Reported from Bizё (Urbahn, 1966), Gasturan and Vermosh (Misja, 1990). Third report for

Albania.

Perizoma flavofasciata (Thumberg, 1792) Bulgaria, Ossogovo Mts, below Ruen (=Autotransport) chalet,

above Kyustendil town, 1512m, N42.1743°, E022.6342°, 19.VI.2020, near small stream in dark Fagus-Picea

forest; Eastern Rhodopi Mts, 1.5 km from the bridge on Arda River near Madzharovo to Borislavtsi Village,

170m, N41.651691°, E025.866440°, 19.V.2006, rocky slopes with Quercus and Acer monspessullanum; NW

Bulgaria, Vidin Region,Tsar Petrovo Village between Vidin and Kula towns, 223m, N43.968903°,

E022.650887°, 02.5.2010, meadows and light forest with Acer tataricum, Quercus, Ulmus, Acer campestre,

Crataegus. North Macedonia, Suva Planina Mt., Kozjak, Venec, 1070m, N41.890857°, E021.223850°,


20

01.VI.2018, limestone rocky area with Fraxinus ornus, Quercus, Pinus, Artemisia alba, etc. Greece, Eastern

Rhodopi Mts, Avdella Village between Kiprinos and Metaxades, 167m, N41.429756°, E026.200884°,

29.IV.2019, Quercus, Phylirea, Pistacia, Colutea; ibid., Erithropotamos (=Luda Reka) River near Alepohori

Village, 37m, N41.4392°, E026.2003°, 30.IV.2019, Populus alba, Fraxinus oxycarpa wet forest near river

and swamps.

Figure 15. a. Catarhoe putridaria, female. MNE, near Bioče, 04.VII.2019; b. Catarhoe rubidata, female. AL, Dajti Mt,

Qafa e Mollёs Pass, 04.VI.2019; c. Euphyia adumbraria, male. SRB, Suva Planina, Divna Gorica, 30.VI.2020; d.

Euphyia adumbraria, male. SRB, Suva Planina, Divna Gorica, 30.VI.2020; e. Euphyia adumbraria, male genitalia of

fig. 101. SRB, Suva Planina, Divna Gorica, 30.VI.2020, Gen. prep. 2./22.I.2021; f. Euphyia frustata, female. AL,

Munella, 1400m, 07.VII.2019; g. Eulithis prunata, male. AL, Pustec (=Liqenas), 849m, 10.VII.2018.

Perizoma obsoletata (Herrich-Schäffer, 1838) Montenegro, Bjelasica Mts, Biogradska Gora National Park,

above Biogradsko Jezero lake, above Katuni Dolovi Lalevića, on the road to Svatovsko Groblje, 1770m,

24.VII.2002, coniferous forest with small meadows and clearings, S. Beshkov & M. Langourov leg., 1 male

(Fig. 16: d), Gen. prep. 3./22.I.2021, S. Beshkov, male genitalia on glass in euparal (Fig. 16: e). Length of

the: uncus – 0.75mm; aedeagus 1.10mm; valva 1.35mm. According to Müller et al. (2019) all population

from the Balkan Peninsula belong to P. juracolaria. In Montenegro however both these species are surely



proved here (see also Perizoma juracolaria, below). New species for the Balkan Peninsula and Montenegro.

Differences between Perizoma obsoletata and Perizoma juracolaria are both external and internal. In the

male genitalia differences are in the shape of the valvae and the uncus as well as in the size of the uncus and

the aedeagus; male genitalia are illustrated and measurements are presented here for both species.

Figure 16. a. Ecliptopera silaceata, male. RNM, Baba Mt., 1493m, 02.VII.2016; b. Dysstroma truncata, female,

upperside. AL, Mali me Gropa Mts, 1405m, 05.VI.2019; c. Dysstroma truncata, female, underside. AL, Mali me Gropa

Mts, 1405m, 05.VI.2019; d. Perizoma obsoletata, male. MNE, Bjelasica, Biogradska Gora, 24.VII.2002; e. Perizoma

obsoletata, male genitalia of 16: d MNE, Bjelasica, Biogradska Gora, 24.VII.2002, Gen. prep. 3./22.I.2021; f. Perizoma

juracolaria, male. AL, Lurё, 9.VII.2019; g. Perizoma juracolaria, male genitalia of 16: f AL, Lurё, 9.VII.2019, Gen.

prep. 2./23.I.2021; h. Perizoma juracolaria, female. AL, Lurё, 9.VII.2019; i. Perizoma juracolaria, male. SRB, Stolovi,

5.VII.2017.


22

Figure 17. a. Perizoma juracolaria, male genitalia of 16: i SRB, Stolovi, 5.VII.2017, Gen. prep. 1./23.I.2021; b.

Perizoma juracolaria, female. SRB, Stolovi, 5.VII.2017; c. Perizoma juracolaria, male. MNE, Pivska Planina, Plužine,

18.VII.2019; d. Perizoma juracolaria, male genitalia. MNE, Pivska Planina, Plužine, 18.VII.2019, Gen. prep.

1./12.II.2021; e. Eupithecia venosata, female. AL, Dajti Mt, Qafa e Mollёs, 04.VI.2019; f. Eupithecia silenicolata,

female. SRB, Stolovi, 05.VII.2017.

Perizoma juracolaria (Wehrli, 1919) Albania, Lurё Mt, above Fushё-Lurё Village, 1687 m, N41.7767°,

E020.1989°, 09.VII.2019, 1 male (Fig. 16: f), Gen. prep. 2./23.I.2021, S. Beshkov, male genitalia on glass in

euparal (Fig. 16: g) and 4 females (Fig. 16: h). Length of the: uncus 0.06mm; aedeagus 1.45mm; valva



1.50mm. From Albania (Mali me Gropa) Perizoma obsoletata is reported by Urbahn (1966) as Cidaria

obsoletaria Herrich-Schäffer, 1838. Very possibly this report concerns P. juracolaria, although both these

species are sympatric on the Balkans, particularly in Montenegro (see also under Perizoma obsoletata,

above). This is the first documented record of P. juracolaria for Albania. Serbia, Kraljevo Region, Stolovi

Mt., Veliki Čukar, 688m, N43.601922°, E020.685631°, 05.VII.2017, 2 males (Fig. 16: i) Gen. prep.

1./23.I.2021, S. Beshkov, male genitalia on glass in euparal (Fig. 17: a) and 5 females (Fig. 17: b). Length of

the: uncus 0.06mm; aedeagus 1.45mm; valva 1.35mm.; ibid., 01.VII.2019, S. Beshkov, A. Nahirnić, P.

Jakšić, C. Plant, A. King leg., 1 female in coll. S. Beshkov. In Serbia P. juracolaria is surely confirmed by

Stojanović et al. (2018) from Mt. Čemerno, from where it is reported as a new for Serbia and illustrated with

male genitalia. Second report for Serbia. Montenegro, Pivska Planina, Plužine, 1474m, 18.VII.2019, P.

Jakšić leg., 1 male (Fig. 17: c), Gen. prep. 1./12.II.2021 S. Beshkov, male genitalia on glass in euparal (Fig.

17: d). Length of the: uncus 0.06mm; aedeagus 1.35mm; valva 1.35mm. In Müller et al. (2019) it is marked

on the map of Montenegro but without additional data. First faunistic report for Montenegro.

Eupithecia venosata (Fabricius, 1787) Albania, Tirana County, Dajti Mt, Qafa e Mollёs Pass, 665m,

N41.3643°, E019.96547°, 04.VI.2019, 1 female (Fig. 17: e). Reported from Mali me Gropa (Urbahn, 1966),

Mbasdejë, Qarrishtë and Vermosh (Misja, 1977). Third report for Albania.

Eupithecia silenicolata Mabille, 1867 Albania, Prespa Lake, near Pustec (=Liqenas) Village, 862m,

N40.774238°, E020.903649°, 10.VII.2018. The report by Rebel & Zerny (1931) is for Novo Selo, which is

in Serbia. New species for Albania. Serbia, Kraljevo Region, Stolovi Mt., Veliki Čukar, 688m, N43.601922°,

E020.685631°, 05.VII.2017, 1 male (Fig. 17: f). In Serbia known from three localities: Fruška Gora (Tomić

et al., 2002, Stojanović et al., 2010), Ovčar–Kablar Gorge (Đurić & Hric, 2015) and Kopaonik (Stojanović &

Šumarac, 2020).

Eupithecia extremata (Fabricius, 1787) Serbia, Stol Mt., northern foothill, above Luka Village, 807m,

N44.18334°, E022.13983°, 05.VI.2020, limestone slopes above Fagus forest, 1 female (Fig. 18: a). Known

from Južna Morava river in the outskirs of Niš (Pettersson, 1990) and Zaječar (Zečević & Radovanović,

1974; Zečević, 2002; Tomić et al., 2002).

Eupithecia breviculata ((Donzel, 1837) E. Serbia, Stol Mt., northern foothill, above Luka Village, 807m,

N44.18334°, E022.13983°, 05.VI.2020, limestone slopes above Fagus forest, 1 female (Fig. 18: b). In Serbia

known from Crni Vrh (Vidlič Mt.) (Beshkov, 2017; Jakšić, 2018) and Starac Mt. (Beshkov et al., 2020b).

Third locality for Serbia.

Lasiocampidae

Malacosoma franconica ([Denis & Schiffermüller], 1775) Albania, Korçё County, Krushovё, above

Voskopojё, 1296m, N40.6706°, E020.5674°, 11.VII.2018, serpentinite slopes with Buxus sempervirens,

Pinus nigra, Juniperus, etc., 1 female found resting during the day. Previously published for Albania only

from Tirana (Rebel & Zerny, 1931). Second report for Albania. Greece, Peloponessos, Taygetos Mt., Mt.

Profitis Ilias eastern slope, 21.VI.2016, leg. A. Nahirnić, one female found resting during the day. Bulgaria,

Burgas region, between Bryastovets and Rudnik villages, 225 m, N42.654175°, E27.482532°, 11.VI.2019, 1

female found resting during the day. Eastern Rhodopi Mts, Sheynovets Ridge, above Mezek Village, 535m,

N41.7178°, E026.0628°, 08.VI.2019, Fraxinux ornus, Phyllirea latifolia, Syringa vulgaris, Genista, Peonia,

Jasminum, Quercus, Asparagus acutifolius, 1 female found resting during the day. It is known from five

localities in Bulgaria (Ganev, 1984; Hristova & Beshkov, 2016). According to Hristova & Beshkov (2016)

M. franconica is Endangared (IUCN cathegory EN B2abiii) in Bulgaria.

Brahmaeidae

Lemonia strigata Rougeot et Viette, 1978 Albania, Gjirokaster County, Mt. Lunxhërisë, between Mal Çajup

and Erind village, 1015m, N40.182533°, E020.166066°, 02.X.2019, 2 males and 2 females (Fig.18: c-d);

Gjirokastër County, Këlcyrë (=Klisura) Gorge on Vjosa (=Aoos) River near Këlcyrë (=Klisura) Village,

176m, N40.29646°, E020.16260°, 03.X.2019, river valley with Platanus orientalis, Alnus glutinosa and


24

Paliurus spina-christi in maquis; Shkodra County, Vau i Dejës, 333m, N42.00283°, E019.68944°,

05.X.2019, open area with Quercus trojana, Quercus pubescens, Juniperus oxycedrus, Fraxinus ornus.

Perhaps all reports in the literature for Albania for Lemonia taraxaci ([Denis & Schiffermüller], 1775) refer

to Lemonia strigata, a species that is not rare in Albania from September to November from the sea coast to

about 1100m. a.s.l. Both species are split chronologically and by altitude and can be easily split by habitus as

well. Lemonia strigata usually has a dark median line which sometimes can be reduced (Beshkov &

Nahirnić, 2019a). North Macedonia, Gostivar, Suva Gora Mts, below Korito Village, 1091m, N41.823606°,

E020996654°, 26.IX.2016; Vardar-Pčinja, above Katlanovska Banja, 304m, N41.901710°, E021.690316°,

14.X.2016; Ohrid, Petrina Planina, between Ohrid and Velestovo, 1005m, N41.090632°, E020.827265°,

15.X.2016; Vardar River Valley, above Demir Kapija Town, on the road to Besvica Village, 244m,

N41.382783°, E022.195847°, 17.X.2017; Prilep, Babuna Planina, Pletvar Pass, 960m, N41.37008°,

E021.66988°, 27.IX.2019. It seems that Lemonia strigata is well presented in North Macedonia while

Lemonia taraxaci ([Denis & Schiffermüller], 1775) is not confirmed for that country.

Figure 18. a. Eupithecia extremata, female. SRB, Bor, Stol Mt., 05.VI.2020; b. Eupithecia breviculata, male. SRB,

Bor, Stol Mt., 05.VI.2020; c-d. Lemonia strigata, females. AL, Mt. Lunxhërisë, Mal Çajup, 02.X.2019; e. Acherontia

atropos, male. AL, Shkodra, Vau i Dejës, 05.X.2019; f. Daphnis nerii, male. AL, Korçё, Pepellash, 28.IX.2019; g.

Ocneria ledereri, male. BG, E Rhodopi Mts, Sheynovets, 08.VI.2019; h. Ocneria ledereri, female. BG, E Rhodopi Mts,

Sheynovets, 08.VI.2019; i. Diaphora luctuosa, male. AL, Dajti Mt, Qafa e Mollёs, 04.IV.2019; j. Zanclognatha

zelleralis, male. AL, Malësia e Tiranës, Qafa e Selitës, 15.VII.2018; k. Lygephila pastinum, female. AL, Qafa e Thorës

- Theth, 1657m, 19.VII.2018; l. Autophila ligaminosa, male. AL, Mali me Gropa, 1501m, 16.VII.2018.



Sphingidae

Acherontia atropos (Linnaeus, 1758) Albania, Shkodra County, Vau i Dejës, 333m, N42.00283°,

E019.68944°, 05.X.2019, 1 male, wingspan 100mm (Fig. 18: e). Reported from Oroshi (Rebel, 1913),

Durres (Eichler & Friese, 1965), Vukopoles (Nahirnić & Beshkov, 2017), Çorovodë and Tomori, 2379m

(Beshkov et al., 2020a).

Daphnis nerii (Linnaeus, 1758) Albania, Korçё County, Mt. Kuq, Qarrit pass, below Pepellash village,

999m, N40.4708°, E020.6722°, 28.IX.2019, 1 male, wingspan 103mm (Fig. 18: f); Tirana Region, Dajti Mt,

Qafa e Mollёs Pass, 665m, N41.3643°, E019.96547°, 24.XI.2019. Known from Shkodra (Eichler & Friese,

1965), Tirana and Golem near Kavajë (Misja, 1978), Tirana, The Botanical Garden (Beshkov, 1994).

Rethera komarovi drilon Rebel & Zerny, 1931 Albania, Munella Mt., above Kimez Village, 1014m,

N41.9710°, E020.0686°, 06.VII.2019, coniferous forest with Bruckenthalia spiculifolia. This species has not

been found in Albania for more than 100 years and is known from only a single locality from where was

described ssp. drilon (7-8.VI.1918, Brutti at the confluence of White Drin and Black Drin Rivers [near

Kukës]) (Rebel & Zerny, 1931, Eichler & Friese, 1965). Second report for Albania. Bulgaria, Eastern

Rhodopi Mts, below Komuniga Village, 494m, N41.796116°, E025.201705°, 14.V.2018, S. Beshkov & B.

Zlatkov leg.; Eastern Rhodopi Mts, Byala Reka, Zhultichalskoto Dere river near Meden Buk Village, 111m,

N41.379995°, E026.027538°, 03.V.2013 and 30.IV.2014. Rethera komarovi was known in Bulgaria from

Kresna Gorge, Alibotoush Mts and some more localities in Eastern Rhodopi only (Beshkov & Langourov,

2004). According to Hristova & Beshkov (2016) R. komarovi is Vulnerable (IUCN cathegory VU B2abiii) in

Bulgaria.

Notodontidae

Drymonia velitaris (Hufnagel, 1766) Serbia, Bela Palanka District, Šljivovički Vis Mt., above Šljivovik

Village, 925m, N43.141432°, E022.386693°, 21.VI.2017, 1 male; Kraljevo Region, Stolovi Mt., Veliki

Čukar, 688m, N43.601922°, E020.685631°, 01.VII.2019, S. Beshkov, A. Nahirnić, P. Jakšić, C. Plant, A.

King leg. In Serbia known only from Stara Planina Mt., above Gornji Krivodol Village (Beshkov, 2017).

Second report for Serbia.

Erebidae

Lymantriinae

Ocneria ledereri (Millière, 1869) Bulgaria, Eastern Rhodopi Mts, Sheynovets Ridge, above Mezek Village,

535m, N41.7178°, E026.0628°, 08.VI.2019, Fraxinux ornus, Phyllirea, Syringa, Genista, Peonia, Jasminum,

Quercus, Asparagus acutifolius, 1 male (Fig. 18: g) and 1 female (Fig. 18: h). In Bulgaria Ocneria ledereri is

known from some more localities in the south in Eastern Rhodopes and from Kresna Gorge (Beshkov &

Langourov, 2004). Findings of males are infrequent.

Arctiinae

Diaphora luctuosa (Geyer, [1831]) Albania, Tirana County, Dajti Mt, Qafa e Mollёs Pass, 665m, N41.3643°,

E019.9657°, 04.IV.2019, 1 male (Fig. 18: i). In Albania D. luctuosa is known from Kula e Lumёs (Rebel &

Zerny, 1931), Boga Village (Beshkov, 1995), Vermosh (Beshkov et al., 1996), Zvezdё and Boboshticё

(Beshkov et al., 2020b).

Herminiinae

Zanclognatha zelleralis (Wocke, 1850) Albania, Tirana County, Malësia e Tiranës, (Highlands of Tirana),


15.VII.2018, 1 male (Fig. 18: j). In Albania Z. zelleralis is known from Kula e Lumёs (Rebel & Zerny, 1931;

Heinicke, 1965), Krraba and Uji Ftohte southern of Tepelene (Heinicke, 1965). Third report for Albania.


26

Toxocampinae

Lygephila pastinum (Treitschke, 1826) Albania, Shkodra County, above Boga Village, Bjeshkët e Nemuna

Mts (=Prokletije Mts), Radohima Mt. between Qafa e Thorës pass and Theth village, south-east of Shtegu

peak, 1657m, N42.3854°, E019.7502°, 19.VII.2018, 1 female (Fig. 18: k). In Albania this species was known

from only a single locality: Dishnicё (Beshkov & Nahirnić, 2018b). Second report for Albania.

Autophila ligaminosa (Eversmann, 1851) Albania, Tirana County, Mali me Gropa, Central part S of plateau,

1501m, N41.3617°, E020.0568°, 16.VII.2018 (Figs 18: l, 19: a-b). In Albania known from Strelcë (Beshkov

& Nahirnić, 2018a). Second report for Albania. Perhaps the reports for Autophila anaphanes Boursin, 1940

and Autophila cataphanes (Hübner, [1813]) for Oroshi (Rebel, 1913; Rebel & Zerny, 1931; Heinicke, 1965),

Llogara and Vlora (Misja, 1980) concern A. ligaminosa, the species surely proved for Albania.

Boletobiinae

Odice arcuina (Hübner, 1790) North Macedonia, near Vardar River Valley above Demir Kapija Town on the

road to Besvica Village, 244m, N41.382783°, E022.195847°, 18.VIII.2017 (Fig. 19: c-d). In North

Macedonia known from Valandovo, Stari Dojran and Drenovo (Thurner, 1964).

Eublemma ostrina (Hübner, [1808]) North Macedonia, near Vardar River Valley above Demir Kapija Town

on the road to Besvica Village, 244m, N41.382783°, E022.195847°, 05.V.2017 (Fig. 19: e); Prilep Region,

Babuna Planina, Pletvar Pass, 960 m, N41.3701°, E021.6699°, 27.IX.2019 (Fig. 19: f). Polymorphic and

polyvoltine species with great differences in colour and size between generations.

Metachrostis dardouini (Boisduval, 1840) North Macedonia, near Vardar River Valley above Demir Kapija

Town on the road to Besvica Village, 244m, N41.382783°, E022.195847°, 05.V.2017 (Fig. 19: g). Second

report for North Macedonia, previously known only from Suva Planina: Kozjak, Venec (Beshkov &

Micevski, 2019).

Erebinae

Catocala elocata (Esper, 1787) Bulgaria, Sakar Mts, near Matochina Village, 167m, N41.8533,°

E026.5505°, 05.X.2020, Gen. prep. 2./21.XI.2020, S. Beshkov, male genitalia with everted vesica on glass in

euparal (Fig. 19: h-k). Catocala elocata is a supposedly common species in Bulgaria and in the Balkan

Peninsula, but there is a question whether the closely similar species Catocala deducta Eversmann, 1843

may be present as well. Examination of the genitalia is necessary.

Noctuidae

Cuculliinae

Cucullia lactucae ([Denis & Schiffermüller], 1775) Albania, Munella Mt, above Kimez Village, 1400m,

N41.9491°, E020.0926°, 07.VII.2019, 1 male (Fig. 20: a), Gen. prep. 1./27.III.2020, S. Beshkov, male

genitalia with everted vesica on glass in euparal (Fig. 20: b-d). New species for Albania.

Cucullia dracunculi (Hübner, [1813]) Albania, Munella Mt, above Kimez Village, 1400m, N41.9491°,

E020.0926°, 07.VII.2019, 1 female (Fig. 20: e), Gen. prep. 1./29.III.2020, S. Beshkov, on glass in euparal

(Fig. 20: f). From Albania Cucullia dracunculi was reported as new for the Balkan Peninsula from Shkallë

(Beshkov & Nahirnić, 2019a). In Albania known also from Plani i Bardhë (Beshkov & Nahirnić, 2019b).

Third report for Albania and for the Balkan Peninsula.

Shargacucullia verbasci (Linnaeus, 1758) Albania, Tomorri Mts, above Ujanik, Abaz Aliu Top, 2379m,

N40.6361°, E020.1615°, 14.VII.2018, 2 females (Fig. 21: a-b), genitalia of one checked. Both specimens

have unusual appearance with reduced to missing rusty coloration. In Albania Sh. verbasci is known from

Tirana (Rebel & Zerny, 1931; Heinicke, 1965) and Dhermi (Beshkov et al., 1996). Third report for Albania.



Figure 19. a. Autophila ligaminosa, male of 18: l, underside. AL, Mali me Gropa, 1501m, 16.VII.2018; b. Autophila

ligaminosa, male. AL, Mali me Gropa, 1501m, 16.VII.2018; c. Odice arcuina, male, upperside. RNM, Demir Kapija,

Besvica, 18.VIII.2017; d. Odice arcuina, male of 19: c underside. RNM, Demir Kapija, Besvica, 18.VIII.2017; e.

Eublemma ostrina, male. RNM, Demir Kapija, Besvica, 05.V.2017; f. Eublemma ostrina, male. RNM, Prilep, Pletvar,

960m, 27.IX.2019; g. Metachrostis dardouini, male. RNM, Demir Kapija, Besvica, 05.V.2017; h. Catocala elocata,

male genitalia. BG, Sakar Mt., Matochina, 05.X.2020, Gen. prep. 2./21.XI.2020; i-k. Catocala elocata, everted vesica.

BG, Sakar Mt., Matochina, 05.X.2020, Gen. prep. 2./21.XI.2020.

Eustrotiinae


28

Figure 20. a. Cucullia lactucae, male. AL, Munella Mt, 1400m, 07.VII.2019; b. Cucullia lactucae, male genitalia of 20:

a AL, Munella Mt, 1400m, 07.VII.2019, Gen. prep. 1./27.III.2020; c-d. Cucullia lactucae, everted vesica of 20: a AL,

Munella Mt, 1400m, 07.VII.2019, Gen. prep. 1./27.III.2020; e. Cucullia dracunculi, female. AL, Munella Mt, 1400m,

07.VII.2019; f. Cucullia dracunculi, female genitalia of 20: e AL, Munella Mt, 1400m, 07.VII.2019.

Phyllophila obliterata (Rambur, 1833) Serbia, Kraljevo Region, Mt. Stolovi, Veliki Čukar, 688m,

N43.601922°, E020.685631°, 05.VII.2017, 1 male. In Serbia it is known from the northern part of the

country (Stojanović et al., 2011) and from Nacionalni park “Kopaonik” (Stojanović & Šumarac, 2020).

Pseudozarba bipartita (Herrich-Schäffer, 1850) Albania, Mt. Thanës, near Bulqizë town, above Plani i

Bardhë village, N41.4762°, E020.1554°, 767 m, 20.VII.2018, 1 male (Fig. 21: c) and one female (Fig. 21: d);



Delvina district, above Muzinё Village, on the road to Dhrovjan, 389m, N39.914449°, E020.201717°,

12.VII.2018, 1 male (Fig. 21: e). In Albania known from a single locality: Stan-Karbunarë (Beshkov, 2018).

Second report for Albania.

Acontiola moldavicola (Herrich-Schäffer, [1851]) Montenegro, between Danilovgrad and Nikšić,

Bogmilovići Village, 468m, N42.613625°, E019.013915°, 30.VI.2017, limestone slopes with Juniperus

oxycedrus, Punica, Fraxinus ornus, Acer monspessulanum, Pistacia terebinthus, Thalictrum, Paliurus spina-

christi, one female (Fig. 21: f) and 2 males (Fig. 21: g) in light trap. At light this species was collected

recently also in Greece (De Vrieze, 2001) and in Albania (Beshkov & Nahirnić, 2018b). From Montenegro

reported from Kotor more than 60 years ago (Carnelutti & Michieli, 1958). Second report for Montenegro.

Figure 21. a-b. Shargacucullia verbasci, females. AL, Mt. Tomor, 2379m, 14.VII.2018; c. Pseudozarba bipartita,

male. AL, Bulqizë, Plani i Bardhë, 20.VII.2018; d. Pseudozarba bipartita, female. AL, Bulqizë, Plani i Bardhë,

20.VII.2018; e. Pseudozarba bipartita, male. AL, Delvina, Muzinё, 12.VII.2018; f. Acontiola moldavicola, female.

MNE, Bogmilovići, 30.VI.2017; g. Acontiola moldavicola, male. MNE, Bogmilovići, 30.VI.2017; h. Acronicta

euphorbiae, female. AL, Dajti Mt, Qafa e Mollёs, 30.IX.2019; i. Calliergis ramosa, male. AL, Mali me Gropa, 1405m,

05.VI.2019.

Acronictinae

Acronicta euphorbiae ([Denis & Schiffermüller], 1775) Albania, Tirana County, Dajti Mt, Qafa e Mollёs

Pass, 665m, N41.3643°, E019.9657°, 30.IX.2019, 1 female (Fig. 21: h).


30

Oncocnemidinae

Calliergis ramosa (Esper, 1786) Albania, Tirana County, Mali me Gropa, 1405m, N41.3524°, E020.0465°,

05.VI.2019, stony slopes with Artemisia alba, Saturea, Juniperus, 1 male (Fig. 21: i). In Albania known

from Llogara (Misja, 1980) and Boboshticё (Beshkov et al., 2020). Third locality for Albania.

Epimecia ustula (Freyer, [1835]) Albania, Tirana County, Mali me Gropa, 1405m, N41.3524°, E020.0465°,

17.VII.2018, 1 male. In Albania known from Dhermi (Beshkov, 1995) and Vlora (Beshkov et al., 1996).

Montenegro, between Danilovgrad and Nikšić, Bogmilovići Village, 468m, N42.613625°, E019.013915°,

30.VI.2017, limestone slopes with Juniperus oxycedrus, Punica granatum, Fraxinus ornus, Acer

monspessulanum, Pistacia terebinthus, Thalictrum, Paliurus spina-christi, 1 male (Fig. 22: a). In

Montenegro known from Bar (Carnelutti & Michieli, 1958) and the from the Gulf of Kotor, erroneously as a

new species for Montenegro (Beshkov, 2017). Third locality for Montenegro.

Figure 22. a. Epimecia ustula, male. MNE, Bogmilovići, 30.VI.2017; b. Nyctobrya amasina, male. AL, Çorovodë,

Çerenisht, 09.VIII.2018; c. Proxenus hospes, male. AL, Shkodra, Vau i Dejës, 05.X.2019; d. Nonagria typhae, female.

AL, Syri i Kaltër, 12.VII.2018; e. Pabulatrix pabulatricula, male. BiH, Bosnia, Pribošijevići near Rogatica,

04.VII.2017; f. Apamea lithoxylaea, male. AL, near Pustec (=Liqenas), 849m, 10.VII.2018; g. Tiliacea cypreago, male.

AL, Tepelene, Bëncës River Gorge, 04.X.2019; h. Episema tersa, male. MNE, Ibarska Klisura, near Rožaje, 07.X.2019;

i. Episema korsakovi, male. AL, Gjorokaster, Mt. Lunxhërisë, Mal Çajup, 02.X.2019.



Teinoptera lunaki (Boursin, 1940) Bulgaria, Osogovo Mts, Piyanets Ridge, Smolichano Village,

“Ravnishteto” place, 705m, N42.146681°, E022.806455°, 08.VI.2017, dry warm slopes near Quercus forest

S. Beshkov, J. Toth & J. Bereczki legs, as a day time copula on a flower of high sticky Silene sp. cf. Silene

italica (L.) Pers. In Bulgaria its northernmost locality was in Kresna Gorge (Beshkov, 2000). The new

locality seems to be the northernmost not only for Bulgaria, but also for the whole range of the species which

is endemic for the southern Balkans (T.L.: Ohrid).

Bryophilinae

Nyctobrya amasina (Draudt, 1931) Albania, Osum river canyon, Çorovodë town south, opposite of Çerenisht

village, N40.4846°, E020.2362°, 09.VIII.2018, 2 males (Fig. 22: b), genitalia checked. Reported as new for

Albania by Beshkov & Nahirnić (2018a), and subsequently found in some more localities (Beshkov &

Nahirnić, 2018b; Beshkov & Nahirnić, 2019a; Beshkov et al., 2020a).

Noctuinae

Proxenus hospes (Freyer, [1831]) Albania, Shkodra County, Vau i Dejës, 333m, N42.00283°, E019.68944°,

05.X.2019, 1 male (Fig. 22: c). Known from: 6 km E of Tirana to Elbasan (Beshkov & Misja, 1995),

Vukopoles (Nahirnić & Beshkov, 2017); Starë; Librazhd; Parpunje; Ardenica; Syri i Kaltër (Beshkov &

Nahirnić, 2019a).

Nonagria typhae (Thunberg, 1784) Albania, Gjirokastër County, Finiq municipality, Syri i Kaltër spring,

N39.923070°, E020.191716°, 155 m, 12.VII.2018, 1 female (Fig. 22: d). In Albania known from Stan-

Karbunarë (Beshkov & Nahirnić, 2018a), Butrint Lake and Prespa Lake (Beshkov & Nahirnić, 2018b). Third

report for Albania.

Pabulatrix pabulatricula (Brahm, 1791) Bosnia & Herzegovina, Rogatica, Pribošijevići village, 643m,

N43.783897°, E018.985954°, 04.VII.2017, meadow, gardens, Quercus forest, 1 male (Fig. 22: e), genitalia

checked. New genus and new species for Bosnia & Herzegovina and the Western Balkans. In the Balkan

Peninsula P. pabulatricula is known from two localities close to each other, near the southern Black Sea

Coast of Bulgaria (Hacker, 1996; Beshkov, 2000; Beshkov, 2017).

Apamea lithoxylaea ([Denis & Schiffermüller], 1775) Albania, Prespa Lake, near Pustec (=Liqenas) Village,

849m, N40.770632°, E020.908891°, 10.VII.2018, 1 male (Fig. 22: f). New species for Albania.

Oligia versicolor (Borkhausen, 1792) Bosnia & Herzegovina, Sarajevo County, Bjelašnica Mt, Gornja

Grkarica, 1175m, N43.730122°, E018.284521°, 01.VII.2017, meadow in coniferous forest, 1 male, genitalia

checked. For Bosnia & Herzegovina O. versicolor is mentioned as present there (e.g. Vasić, 2002) but

without locality and we hardly believe the reliability of such reports if the genitalia were not checked. This

seems to be the first reliable report for Bosnia & Herzegovina.

Tiliacea cypreago (Hampson, 1906) Albania, Gjirokastër County, Bëncës River Gorge, ca 3 km SW from

Bëncë Village, Tepelene District, 555m, N40.24188°, E019.97882°, 04.X.2019 (Fig. 22: g). In Albania

known only from the northern parts of the country: Plani i Bardhë, Qafa e Mollës, and Starë (Beshkov &

Nahirnić, 2019a). Second report for Albania.

Episema tersa ([Denis & Schiffermüller], 1775) Albania, Shkodra County, Vau i Dejës, 333m, N42.00283°,

E019.68944°, 05.X.2019. In Albania known from Llogara, Zvezdë, Butrint (Beshkov & Nahirnić, 2018a)

and Plani i Bardhë (Beshkov & Nahirnić, 2019a). Third report for Albania. Montenegro, Ibarska Klisura

Gorge, near Rožaje, 1027m, N42.85862°, E020.20956°, 07.X.2019, 1 male (Fig. 22: h). According to Vasić

(2002) and Zečević (2002) Episema tersa is known in all neighboring countries of Serbia. At that time,

however, Montenegro and Serbia were one country. Here Episema tersa is reported as a new species for

Montenegro.


32

Episema korsakovi (Christoph, 1885) Albania, Gjirokaster County, Mt. Lunxhërisë, between Mal Çajup and

Erind village, 1015m, N40.182533°, E020.166066°, 02.X.2019 (Fig. 22: i). Reported as a new species for

Albania from Zvezdë and Drenovë (Beshkov & Nahirnić, 2018a). Second report for Albania. North

Macedonia, Prilep Region, Babuna Planina, Pletvar Pass, 960 m, N41.3701, E021.6699, 27.IX.2019, 1 male

(Fig. 23: a).

Cleoceris scoriacea (Esper, [1789]) Albania, Korçё County, Mt. Kuq, Qarrit pass, below Pepellash village,

999m, N40.4708°, E020.6722°, 28.IX.2019 (Fig. 23: b).

Cirrhia gilvago ([Denis & Schiffermüller], 1775) North Macedonia, near Vardar River Valley, above Demir

Kapija Town on the road to Besvica Village, 244m, N41.382783°, E022.195847°, 17.X.2017, maquis,

Juniperus exelsa, Pistacia, Phyllirea, Astragalus, etc. Known from Drenovo (Thurner, 1964). Second report

for North Macedonia.

Figure 23. a. Episema korsakovi, male. RNM, Prilep, Pletvar Pass, 27.IX.2019; b. Cleoceris scoriacea, male. AL,

Korçё, Pepellash, 28.IX.2019; c. Anchoscelis humilis, female. MNE, near Bioče Village, 195m, 06.X.2019; d.

Anchoscelis helvola, male. MNE, Ibarska Klisura, near Rožaje, 07.X.2019; e. Atethmia centrago, male. AL,

Gjorokaster, Mt. Lunxhërisë, Mal Çajup, 02.X.2019; f. Ammoconia senex, male. AL, Malësia e Tiranës, Qafa e Selitës,

1222m, 01.X.2019; g. Aporophyla canescens, male. AL, Shkodra, Vau i Dejës, 05.X.2019; h. Aporophyla canescens,

male. MNE, near Bioče Village, 195m, 06.X.2019; i. Dasypolia templi vecchimontium, male. AL, Malësia e Tiranës,

Qafa e Selitës, 1222m, 01.X.2019.



Anchoscelis litura (Linnaeus, 1758) Albania, Krraba Mt, near Gracen Village, 783m, N41.16092°,

E019.96299°, 29.IX.2019, Quercus-Carpinus forest with Juniperus oxycedrus, Cystus, Saturea, 1 male,

genitalia checked. In Albania known only from Prespa Lake (Beshkov & Nahirnić, 2018c). Second report for

Albania. Montenegro, Ibarska Klisura Gorge, near Rožaje, 1027m, N42.85862, E020.20956, 07.X.2019, 1

male, genitalia checked. In Montenegro known from Durmitor and Tara canyon (Carnelutti et al., 1991).

Anchoscelis humilis ([Denis & Schiffermüller], 1775) Montenegro, Morača River Valley, near Bioče

Village, 195m, N42.5273°, E019.3549°, 06.X.2019, 1 female (Fig. 23: c). In Montenegro known from

Durmitor and Tara canyon (Carnelutti et al., 1991).

Anchoscelis helvola (Linnaeus, 1758) Montenegro, Ibarska Klisura Gorge, near Rožaje, 1027m, N42.85862°,

E020.20956°, 07.X.2019, 1 male (Fig. 23: d). In Montenegro known from Durmitor and Tara canyon

(Carnelutti et al., 1991).

Cosmia pyralina ([Denis & Schiffermüller], 1775) Bosnia & Herzegovina, Rogatica, Pribošijevići village,

643m, N43.783897°, E018.985954°, 04.VII.2017, meadow, gardens, Quercus forest, 1 male. New species

for Bosnia & Herzegovina.

Atethmia centrago (Haworth, 1809) Albania, Krraba Mt, near Gracen Village, 783m, N41.16092°,

E019.96299°, 29.IX.2019, Quercus-Carpinus forest with Juniperus oxycedrus, Cystus, Saturea; Albania,

Gjirokaster County, Mt. Lunxhërisë, between Mal Çajup and Erind village, 1015m, N40.182533°,

E020.166066°, 02.X.2019 (Fig. 3: e). In Albania reported from Gorica near Korçё, Butrint and Borsh (Misja,

1976). Second report for Albania.

Dryobotodes servadeii Parenzan, 1982 Albania, Shkodra County, Vau i Dejës, 333m, N42.00283°,

E019.68944°, 05.X.2019, open area with Quercus trojana, Quercus pubescens, Juniperus oxycedrus,

Fraxinus ornus, 1 male. In Albania known from Ilias (Beshkov & Nahirnić, 2018a), Muzinё – Dhrovjan

(Beshkov & Nahirnić, 2018b) and Starë (Beshkov & Nahirnić, 2019a).

Antitype jonis (Lederer, 1865) Albania, Tirana County, Dajti Mt, Qafa e Mollёs Pass, 665m, N41.3643°,

E019.9657°, 30.IX.2019, 1 male; Tirana County, Mali me Gropa Mts, NW of Qafa e Selitёs Pass, 1222m,

N41.37706°, E020.01645°, 01.X.2019, 3 males and 2 females; Gjirokaster County, Mt. Lunxhërisë, between

Mal Çajup and Erind village, 1015m, N40.182533°, E020.166066°, 02.X.2019, 1 male. In Albania known

only from Korab (Rebel & Zerny, 1931; Heinicke, 1965) and from Plani i Bardhë (Beshkov & Nahirnić,

2019a). North Macedonia, Prilep Region, Babuna Planina, Pletvar Pass, 960 m, N41.3701°, E021.6699°,

27.IX.2019, 1 male; Galichica Mts, above Ohrid Town on the road to Oteshevo, above Peshtani Village,

1480m, 10.IX.1997, rocky area near Quercus forest S. Beshkov, V. Gashtarov, M. & K. Beshkovi leg;

Galitchica Mts, above Oteshevo, near the Livada Pass, 1530m, N40.955365°, E020.839439°, 19.X.2007,

mountain steppe above Fagus forest S. Beshkov, J. Buszko, B. Zlatkov leg; Gostivar, Suva Gora Mts, below

Korito Village, 1091m, N41.823606°, E020996654°, 26.IX.2016, slopes with Artemisia and Eryngium spp.

Genitalia of all male specimens checked. In North Macedonia Antitype jonis is reported from Korab

(Thurner, 1964), but this may be the same male specimen reported from Korab by Rebel & Zerny (1931) for

Albania (also the opinion of Hacker (1989)). Species proved to occur in North Macedonia. Known also from

the Greek part of Macedonia and Epirus (Wegner, 2011).

Ammoconia senex (Geyer, [1828]) Albania, Tirana Region, Mali me Gropa Mts, NW of Qafa e Selitёs Pass,

1222m, N41.37706°, E020.01645°, 01.X.2019, 1 male (Fig. 23: f).

Aporophyla canescens (Duponchel, 1826) Albania, Shkodra County, Vau i Dejës, 333m, N42.00283°,

E019.68944°, 05.X.2019, open area with Quercus trojana, Quercus pubescens, Juniperus oxycedrus,

Fraxinus ornus, 1 male (Fig. 23: g). Montenegro, Morača River Valley, near Bioče Village, 195m,

N42.5273°, E019.3549°, 06.X.2019 (Fig. 23: h). New species for Montenegro.

Dasypolia templi vecchimontium Ronkay & Varga, 1985 Albania, Tirana County, Mali me Gropa Mt., NW

of Qafa e Selitёs Pass, 1222m, N41.37706°, E020.01645°, 01.X.2019, 1 male (Fig. 23: i). In Albania known


34

only from Zvezdë (Beshkov & Nahirnić, 2018a). Second locality for Albania.

Polymixis rufocincta (Geyer, 1828) Montenegro, Ibarska Klisura Gorge, near Rožaje, 1027m, N42.85862°,

E020.20956°, 07.X.2019, 1 male (Fig. 24: a). In Montenegro known from Ulcinj (Carnelutti & Michieli,

1958), Bistrica in Durmitor Mts and Tara canyon (Carnelutti et al., 1991).

Panolis flammea ([Denis & Schiffermüller], 1775) Albania, Mt. Thanës, near Bulqizë town, above Plani i

Bardhë village, N41.4798°, E020.1534°, 833m, 06.IV.2019 (Fig. 24: b). Known from Llogara (Beshkov et

al., 1996). Second report for Albania.

Orthosia miniosa ([Denis & Schiffermüller], 1775) Albania, Tirana County, Dajti Mt., Qafa e Mollёs Pass,

665m, N41.3643°, E019.9657°, 04.IV.2019, Quercus-Corylus-Acer forest, 1 female (Fig. 24: c). Known

from Butrint (Beshkov et al., 1996). Second report for Albania.

Orthosia cruda ([Denis & Schiffermüller], 1775) Albania, Tirana County, Dajti Mt., Qafmolla Pass, 665m,

N41.3643°, E019.9657°, 04.IV.2019, Quercus-Corylus-Acer forest, 2 females (Fig. 24: d-e). New species for

Albania. This generally common and widespread species is not reported before from Albania, perhaps

because of its early flight period when night collecting is not always comfortable.

Figure 24. a. Polymixis rufocincta, male. MNE, Ibarska Klisura, near Rožaje, 07.X.2019; b. Panolis flammea, male.

AL, Bulqizë, Plani i Bardhë, 06.IV.2019; c. Orthosia miniosa, female. AL, Dajti Mt, Qafa e Mollёs, 04.VI.2019; d-e.

Orthosia cruda, females. AL, Dajti Mt, Qafa e Mollёs, 04.VI.2019; f. Perigrapha rorida, male. AL, Bulqizë, Plani i

Bardhë, 06.IV.2019; g. Perigrapha i-cinctum slovenica, male. AL, Bulqizë, Plani i Bardhë, 06.IV.2019; h. Perigrapha

i-cinctum slovenica, female. AL, Bulqizë, Plani i Bardhë, 06.IV.2019; i. Conisania renati meszarosi, female. SRB,

Rtanj Mt., 885m, 04.VI.2020.



Perigrapha rorida (Frivaldsky, 1835) Albania, Mt. Thanës, near Bulqizë town, above Plani i Bardhë village,

N41.4798°, E020.1534°, 833 m, 06.IV.2019, 1 male (Fig. 24: f). Known from Rogozhina (Rebel & Zerny,

1931, Heinicke, 1965), Zogaj, Levan and Dhermi (Beshkov et al., 1996).

Perigrapha i-cinctum slovenica Michielie, 1966 Albania, Mt. Thanës, near Bulqizë town, above Plani i

Bardhë village, N41.4798°, E020.1534°, 833 m, 06.IV.2019, 20 males (Fig. 24: g) and 3 females (Fig. 24: h).

New species for Albania.

Conisania renati meszarosi Varga & Ronkay, 1991 Bulgaria, W. Stara Planina Mts, Chavdar Chalet above

Bunovo Village, 15.VIII.2010, V. Gashtarov leg., one dead and dry specimen on a window. Localities in

Bulgaria are summarized in Beshkov (2017). Serbia, Sokobanja district, Rtanj Mt., Vrtače, above Mužinac

village, 885m, N43.75875°, E021.88979°, 04.VI.2020, limestone slopes with dolines with Prunus spinosa,

Syringa vulgaris, Fraxinus ornus, Quercus, Artemisia alba, etc., 3 males and 3 females (Fig. 24: i). In Serbia

known from Crni Vrh summit above Pirot Town and Stara Planina above Gornji Krivodol Village (Beshkov,

2017) and Šljivovički Vis above Bela Palanka Town (Beshkov & Nahirnić, 2017). According to Beshkov

(2017) C. renati meszarosi is an endemic Balkan taxon, known only from western Bulgaria and Eastern

Serbia. Third report for Serbia.

Hadena gueneei (Staudinger, 1901) Morača River Gorge, Milunovići Village N of Bioče, 160m, N42.5544°,

E019.3299°, 04.VII.2019, limestone slopes with Paliuris spina-christi, Quercus, Pistacia, etc. after fire, 1

male (Fig. 25: a), Gen. prep. 1./26.III.2020, S. Beshkov, male genitalia with everted vesica (Fig. 25: b-c).

New species for Montenegro.

Hadena vulcanica urumovi (Drenowski, 1931) Albania, Tirana County, Mali me Gropa, Central part S of

plateau, 1501m, N41.3617, E020.0568, 16.VII.2018 (Fig. 25: d).

Hadena luteocincta (Rambur, 1834), Albania, Tirana County, Malësia e Tiranës (Highlands of Tirana),


15.VII.2018, 1 male (Fig. 25: e). A more certain identification would be possible from examination of the

female genitalia with measurement of the ovipositor.

Hadena caesia xanthophoba (Schawerda, 1922) Albania, Munella Mt, above Kimez Village, 1400m,

N41.9491°, E020.0926°, 07.VII.2019, 1 female (Fig. 26: a); Lurё Mt, above Fushё-Lurё Village, 1687 m,

N41.7767°, E020.1989°, 09.VII.2019, 1 male (Fig. 26: b).

Hadena clara macedonica Boursin, 1959 Albania, Tirana County, Mali me Gropa, Central part S of plateau,

1501m, N41.3617°, E020.0568°, 16.VII.2018, 1 female (Fig. 26: c) and 1 male (Fig. 26: d), Gen. prep.

1./25.III.2020 S. Beshkov, male genitalia with everted vesica on glass in euparal (Fig. 26: e). Everted vesica

before mounting, photographed in alcohol is illustrated on Fig. 26: f-g. New species for Albania. Otherwise

the distribution of Hadena clara macedonica Boursin is restricted to Greece and North Macedonia (T.L.:

Ohrid, Petrina Planina, 1600m). The reports for Bulgaria are wrong and the result of misidentification

(Beshkov, 2000).

Hadena drenowskii drenowskii (Rebel, 1930) North Macedonia, Suva Planina Mt., Kozjak, Venec, 1046m,

N41.8922°, E021.2304°, 07.VII.2018, limestone rocky area with Fraxinus ornus, Quercus, Pinus, Artemisia

alba, etc. 1 male (Fig. 27: a). In North Macedonia known from the SW part of the country: Petrina Planina

and Karaorman (Thurner, 1964) and Galichica Mt. (Hacker, 1996). Third report for North Macedonia.

Enterpia laudeti (Boisduval, 1840) Serbia, Svrljiške Planine Mts, Bela Palanka District, above Moklište

Village, 616m, N43.27566°, E022.29339°, 29.VI.2020, limestone slopes with scattered pine trees, 1 male. In

Serbia it is known from Gornji Krivodol Village (Beshkov, 2017) and from Starac (Beshkov et al., 2020b).

Third locality for Serbia.

Mythimna congrua (Hübner, [1817]) Albania, Tirana County, Dajti Mt, Qafa e Mollёs Pass, 665m,

N41.3643°, E019.9657°, 30.IX.2019, 1 female (Fig. 27: b). Known from Durres (Heinicke, 1965), Tirana,


36

Vlora, Butrint, Fusha-Kuqe (Misja, 1980), Curila near Durres, Dhermi (Beshkov, 1995).

Figure 25. a. Hadena gueneei, male. NME, Milunovići, 04.VII.2019; b. Hadena gueneei, male genitalia with everted

vesica of 25: a NME, Milunovići, 04.VII.2019, Gen. prep. 1./26.III.2020; c. Hadena gueneei, everted vesica of 25: a

NME, Milunovići, 04.VII.2019, Gen. prep. 1./26.III.2020; d. Hadena vulcanica urumovi, male. AL, Mali me Gropa,

1501m, 16.VII.2018; e. Hadena luteocincta, male. AL, Malësia e Tiranës, Qafa e Selitës, 1133m, 15.VII.2018.

Eriopygodes imbecilla (Fabricius, 1794) Bosnia & Herzegovina, Sarajevo County, Bjelašnica Mt, Gornja

Grkarica, 1175m, N43.730122°, E018.284521°, 01.VII.2017, meadow in coniferous forest, 1 male. Known

from Derventa and Vučija Bara near Gacko (Rebel, 1904).

Agrotis catalaunensis (Millière, 1873) (= syricola Corti & Draudt, 1933) Albania, Korçё County, Mt. Kuq,

Qarrit pass, below Pepellash village, 999m, N40.4708°, E020.6722°, 28.IX.2019, 1 male (Fig. 27: c). Known

from Velipoja (Beshkov, 1995), Divjaka, Fikasi, Levan, Dhermi, Butrint (Beshkov et al., 1996). Third report

for Albania.

Agrotis trux (Hübner, [1824]) Albania, Gjirokaster County, Mt. Lunxhërisë, between Mal Çajup and Erind

village, 1015m, N40.182533°, E020.166066°, 02.X.2019, 1 male (Fig. 27: d). Known from Tirana, Divjaka

(Misja, 1980), Zogaj (Beshkov, 1995) and Librazhd (Beshkov & Nahirnić, 2019a).



Figure 26. a. Hadena caesia xanthophoba, female. AL, Munella Mt, 1400m, 07.VII.2019; b. Hadena caesia

xanthophoba, male. AL, Lurё Mt, 09.VII.2019; c. Hadena clara macedonica, female. AL, Mali me Gropa, 1501m,

16.VII.2018; d. Hadena clara macedonica, male. AL, Mali me Gropa, 1501m, 16.VII.2018; e. Hadena clara

macedonica, male genitalia with everted vesica of 26: d AL, Mali me Gropa, 1501m, 16.VII.2018, Gen. prep.

1.25.III.2020; f-g. Hadena clara macedonica, everted vesica of 26: d before mounting. AL, Mali me Gropa, 1501m,

16.VII.2018, Gen. prep. 1.25.III.2020.


38

Figure 27. a. Hadena drenowskii drenowskii, male. RNM, Mt. Kozjak, Venec, 1046m, 07.VII.2018; b. Mythimna

congrua, female. AL, Dajti Mt, Qafa e Mollёs, 30.IX.2019; c. Agrotis catalaunensis, male. AL, Korçё, Pepellash,

28.IX.2019; d. Agrotis trux, male. AL, Mt. Lunxhërisë, Mal Çajup, 02.X.2019; e. Cerastis rubricosa, male. AL,

Bulqizë, Plani i Bardhë, 06.IV.2019; f. Noctua janthe, female. SRB, Stolovi, 05.VII.2017; h. Spaelotis senna contorta,

female. SRB, Suva Planina, 1153m, 30.VI.2020.

Cerastis rubricosa ([Denis & Schiffermüller], 1775) Albania, Mt. Thanës, near Bulqizë town, above Plani i

Bardhë village, N41.4798°, E020.1534°, 833m, 06.IV.2019, 1 male (Fig. 27: e). Known from Beshtriku

(Rebel & Zerny, 1931, Heinicke, 1965), Shkodra (Rebel & Zerny, 1931, Heinicke, 1965, Misja, 1980), Fusha

Lekli near Tepelene (Beshkov et al., 1996).

Lycophotia porphyrea ([Denis & Schiffermüller], 1775) Serbia, Stolovi Mt., Veliki Čukar, 688m,

N43.601922°, E020.685631°, 05.VII.2017 and 01.VII.2019. Albania, Munella Mt., above Kimez Village,

1014m, N41.9710°, E020.0686°, 06.VII.2019, coniferous forest and meadows with Bruckentalia spiculifolia;

ibid, 1400m, N41.9491°, E020.0926°, 07.VII.2019; Lurё Mt, above Fushё-Lurё Village, 1687m, N41.7767°,

E020.1989°, 09.VII.2019; Pinus, Fagus forest, Polygonium bistorta, wetlands, stonny slopes. In Albania



known only from Quarrishta [Qarrishtë] (Misja, 1980). Second report for Albania. Bosnia & Herzegovina,

above Konjic, 652m, N43.647227°, E017.987513°, 02.VII.2017, dolomites with Carpinus, Pinus nigra,

Acer, Quercus, 4 males. In Bosnia and Herzegovina known from Plaša as Agrotis strigula Thnbg. (Rebel,

1904). Findings of Lycophotia porphyria in Serbia and in Bosnia and Herzegovina in localities of low

altitude are surprising. In Bulgaria, North Macedonia and Albania it is a mountain species found mostly at

altitudes of 1400-2000m.a.s.l. with few exceptions. It inhabits the coniferous zone in the mountains where

the possible food plant – Bruckentalia spiculifolia – is present. In the localities of low altitude in Serbia and

Bosnia and Herzegovina mentioned here Bruckentalia spiculifolia is also present, which seems to be the

reason why Lycophotia porphyria is present there.

Chersotis anatolica (Draudt, 1936) North Macedonia, Pelister Mt, above mountain house "Shiroka" 2200m,

06.VIII.2016, B. Zlatkov leg.; Gostivar, Suva Gora Mts, below Korito Village, 1378m, N41.811671°,

E021.016744°, 13.VIII.2016, dry slopes with Artemisia and Eryngium spp. The first report for North

Macedonia is that of Beshkov (2000) and up to now Ch. anatolica was known from that country only from

Korab (Huemer et al., 2011) and from Galichica Mt. where it is sympatric with Chersotis elegans

(Eversmann, 1837) (Beshkov, 2014). Both species are also sympatric in Shar planina and in Timfy Mts. in

Northern Greece (Z. Varga, pers. comm. 30.04.2021).

Noctua janthe (Borkhausen, 1792) Serbia, Kraljevo Region, Stolovi Mt., Veliki Čukar, 688m, N43.601922°,

E020.685631°, 05.VII.2017, serpentinite with Silene, Artemisia, Quercus, Fraxinus ornus, 1 female (Fig. 27:

f). In Serbia N. janthe is known from Jelova Gora, Užice Region (Dodok, 2007), Vasilin Vrh of Javor Mt.

(Beshkov, 2017; Beshkov & Nahirnić, 2020b), Suva Planina above Mokra Village and Stara Planina Mts,

above Gornji Krivodol Village (Beshkov, 2017).

Divaena haywardi (Tams, 1926) Serbia, Svrljiške planine Mts, Bela Palanka District, above Moklište

Village, 616m, N43.27566°, E022.29339°, 29.VI.2020, limestone slopes with planted scattered Pinus nigra

trees, 1 male. In Serbia known from Vzganica in Stara Planina (Stojanović et al., 2014), Suva Planina Mts

above Bojanine vode and Vidlič Mt. above Pirot Town (Beshkov, 2015) and Trnava Village above Preševo

(Beshkov & Nahirnić, 2017).

Spaelotis senna contorta (Rebel & Zerny, 1931) Serbia, Suva Planina above Bela Palanka, 1153m,

N43.19413°, E022.25566°, 30.VI.2020, limestone slopes with Artemisia alba, etc., in Fagus forest (Fig. 27:

g).

Xestia castanea (Esper, 1798) Albania, Krraba Mt, near Gracen Village, 783m, N41.16092°, E019.96299°,

29.IX.2019, Quercus-Carpinus forest with Juniperus oxycedrus, Cystus, Satureja. Known from Vorra (Rebel

& Zerny, 1931; Heinicke, 1965), Voskopoja (Plociennik, Pawlikiewicz & Pabis, 2009), Starë; Parpunje;

Qafa e Mollës and Ardenica (Beshkov & Nahirnić, 2019a).

Xestia cohaesa (Herrich-Schäffer, [1849]) Albania, Krraba Mt, near Gracen Village, 783m, N41.16092°,

E019.96299°, 29.IX.2019, Quercus-Carpinus forest with Juniperus oxycedrus, Cystus, Satureja. In Albania

known from Goricë [near Korçё] (Beshkov et al., 1996), Vukopoles (Nahirnić & Beshkov, 2017), Starë,

Plani i Bardhë and Ardenica (Beshkov & Nahirnić, 2019a). North Macedonia, Prilep Region, Babuna

Planina, Pletvar Pass, 960 m, N41.3701°, E021.6699°, 27.IX.2019 (Fig. 28: a); Vardar-Pčinja, above

Katlanovska Banja, 304m, N41.901710°, E021.690316°, 14.X.2016, Pistacia, Paliurus spina-christi, dry

rocky grasslands. In North Macedonia X. cohaesa is known from Ohrid and Drenovo (Thurner, 1964). The

second report from North Macedonia.

Nolidae

Nola harouni Wiltshire, 1951 North Macedonia, Vardar River Valley, near Demir Kapija Town, on the road

to Klisura village, 155m, N41.389202°, E022.294770°, 20.X.2016, 1 male (Fig. 28: b), Gen. prep.

1./13.I.2021, S. Beshkov, male genitalia on glass in euparal (Fig. 28: c). New species for North Macedonia.

In Europe Nola harouni is known from several localities in Greece and from Bulgaria: Vrachanska Planina

and Kresna Gorge; localities are summarized in Beshkov (2017).


40

Figure 28. a. Xestia cohaesa, male. RNM, Prilep, Pletvar Pass, 27.IX.2019; b. Nola harouni, male. RNM, Demir

Kapija, Klisura, 155m, 20.X.2016; c. Nola harouni, male genitalia of 28: b RNM, Demir Kapija, Klisura, 155m,

20.X.2016, Gen. prep. 1./13.I.2021.

Acknowledgments

We are glad to thank Dr Predrag Jakšić (Belgrade, Serbia) for the Perizoma juracolaria he gave us for

dissection and to Mr Viktor Gashtarov (Sofia, Bulgaria) for the data for Conisania renati meszarosi

published here. We are thankful to Dr Sabine Gaal-Haszler (Natural History Museum Vienna, Austria) for

information and photos of Erebia gorge albanica syntypes illustrated here and to Dr Zoltán Varga

(Debrecen, Hungary) for some important comments incorporated in the text. Dr Mark Shaw (Edinburg,

Scotland, UK) is thanked for linguistic corrections.

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