chapter 4 ecology of weeds in rice...

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61 Chapter 4 Ecology of Weeds in Rice Fields Rice is the most important cereal crop in the developing world and is the staple food of over half of the world’s population (Juliano, 1993). Rice cultivation is thought to be the oldest form of intensive agriculture by man (Fernando, 1977). The geographical site of rice domestication is not yet definitely known and hence remains a matter of conjecture. It is general belief that rice cultivation in Sri Lanka was started by Indo-Aryan immigrants before about 540 B.C (more than 2500 years ago), where it was probably grown as a dry land crop (Grist, 1965; Perera, 1980). Rice-growing environment is varying drastically within countries. Therefore, scientists and specialists from different disciplines and parts of the rice-growing world formed an international committee to determine terminology for rice growing ecosystems. Khush (1984) described the outcome, where the world rice land environments are classified into five major categories based on water regime, drainage, temperature, soil type and topography. India is the largest rice growing country, over 43 percent population has been dependent on rice as food grain. Rice is grown under four different ecological zones, with the irrigated ecology accounting for the largest area and highest production and productivity closely followed by rainfed shallow lowlands. Rainfed upland is just one half of the rainfed lowland area that produces less than one fifth of it. Region-wise, the predominantly rainfed eastern zone accounts for the largest area and production but with the lowest productivity, while the largely irrigated north and south zones together accounting for slightly less area produce one and a half times more than that of eastern India with a distinct yield edge. Western Ghat region of Maharashtra is under rain fed conditions and its ecological parameters were not studied so far. Present ecological study of weeds from rice field has been carried out in ten sites. Bhor and Velhe talukas of Pune district are rich in rice cultivation. Five localities were selected for experiment namely Karanje, Karnawad, Kiwat, Nigudghar and Salekarvasti from Bhor. Similarly in Velhe taluka Ambawane, Gujawani, Pabe, Vinzar and Wanjale were selected as representative villages.

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Page 1: Chapter 4 Ecology of Weeds in Rice Fieldsshodhganga.inflibnet.ac.in/bitstream/10603/77990/11/11... · 2018-07-08 · 61 Chapter 4 Ecology of Weeds in Rice Fields Rice is the most

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Chapter 4

Ecology of Weeds in Rice Fields Rice is the most important cereal crop in the developing world and is the staple

food of over half of the world’s population (Juliano, 1993). Rice cultivation is thought to

be the oldest form of intensive agriculture by man (Fernando, 1977). The geographical site

of rice domestication is not yet definitely known and hence remains a matter of conjecture.

It is general belief that rice cultivation in Sri Lanka was started by Indo-Aryan immigrants

before about 540 B.C (more than 2500 years ago), where it was probably grown as a dry

land crop (Grist, 1965; Perera, 1980).

Rice-growing environment is varying drastically within countries. Therefore,

scientists and specialists from different disciplines and parts of the rice-growing world

formed an international committee to determine terminology for rice growing ecosystems.

Khush (1984) described the outcome, where the world rice land environments are

classified into five major categories based on water regime, drainage, temperature, soil

type and topography.

India is the largest rice growing country, over 43 percent population has been

dependent on rice as food grain. Rice is grown under four different ecological zones, with

the irrigated ecology accounting for the largest area and highest production and

productivity closely followed by rainfed shallow lowlands. Rainfed upland is just one half

of the rainfed lowland area that produces less than one fifth of it. Region-wise, the

predominantly rainfed eastern zone accounts for the largest area and production but with

the lowest productivity, while the largely irrigated north and south zones together

accounting for slightly less area produce one and a half times more than that of eastern

India with a distinct yield edge. Western Ghat region of Maharashtra is under rain fed

conditions and its ecological parameters were not studied so far.

Present ecological study of weeds from rice field has been carried out in ten sites.

Bhor and Velhe talukas of Pune district are rich in rice cultivation. Five localities were

selected for experiment namely Karanje, Karnawad, Kiwat, Nigudghar and Salekarvasti

from Bhor. Similarly in Velhe taluka Ambawane, Gujawani, Pabe, Vinzar and Wanjale

were selected as representative villages.

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• Soil Sample Collection and Analysis:

Soil is the outer loose material of earth’s surface which is distinctly different from

the underlying bedrock and the region which support plant life. Agriculturally, soil is the

region which supports the plant life by providing mechanical support and nutrients

required for growth. It is the region where most of the physical, biological and biochemical

reactions related to decomposition of organic weathering of parent rock takes place. Soil is

an admixture of five major components viz. organic matter, mineral matter, soil-air, soil

water and soil microorganisms/living organisms. The amount of these components varies

with locality and climate. Different types of inorganic compounds containing various

minerals are present in soil. Amongst them the dominant minerals are Silicon, Aluminum,

iron and others like Carbon, Calcium, Potassium, Manganese, Sodium, Sulphur,

Phosphorus etc. are in trace amount. Rice grown areas are essential to know the

physiochemical properties present in the soil. In this connection, Soil samples from 10 sites

of both the talukas were collected and analysed for physiochemical parameters such as pH,

EC, organic carbon, N, P and K. It is also essential to know the Zn, Cu, Iron and

Manganese present in the soil.

Table 4.1. Soil analysis

Taluka  Village  pH  EC  N %  P %  K % Zn Cu  Fe  Mn

(ppm) (ppm)  (ppm) (ppm)Bhor  Karanje  7.18  0.12 0.43 0.001 0.016 0.81 7.00  4.00 11.20

  Karnawad  6.20  0.08 0.36 0.014 0.032 1.31 7.00  17.00 31.40  Kiwat  7.17  0.18 0.25 0.007 0.011 0.64 6.00  5.00 3.20  Nigudghar  5.19  0.10 0.54  0.013  0.021 2.45  10.60  54.00  14.30   Salekarvasti  6.80  0.21 0.32 0.011 0.084 1.98 6.50  1.00 32.40         

Velhe  Ambavane  7.20  0.22 0.22 0.013 0.10 2.28 7.30  15.00 29.90  Gunjavane  6.80  0.08 0.13 0.012 0.13 0.70 4.90  17.00 28.80  Pabe  5.56  0.11 0.56 0.008 0.09 1.17 6.70  55.00 28.70  Vinzar  7.34  0.20 0.30 0.006 0.11 4.20 9.40  17.00 31.80  Wanjale  6.60  0.11 1.34 0.009 0.11 1.70 11.60  41.00 34.60

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Micro- elements in soils:

Soils have been defined differently by geologists, agriculturists, chemists,

engineers, environmentalists and soil scientists but the commonly accepted definition of

soil is that, “Soil is natural body formed out of weathered material and it is defined as a

three dimensional body at the earth surface which supports plants and distinct layers of

mineral and organic components “(Kale and Gupta, 2001). Parent materials of soil play a

key role in concentrating various micro-elements (including heavy metals) in the soils by

releasing them into the soils during soil formation processes. Metals like Zn, Cu, Mn and

Fe naturally enter into the soils due to weathering of parent material. Present study is an

attempt made to know micro-elements in different soil types from 10 rice growing sites of

Bhor and Velhe talukas of Pune District.

I.2 Soil micro-flora:

Soil is an excellent culture media for the growth and development of various

microorganisms. Soil is not an inert static material but a medium pulsating with life. Soil is

now believed to be dynamic or living system. Soil contains several distinct groups of

microorganisms and amongst them bacteria, fungi, actinomycetes, algae, protozoa and

viruses are the most important. But bacteria are more numerous than any other kinds of

microorganisms. Microorganisms form a very small fraction of the soil mass and occupy a

volume of less than one percent. In the upper layer of soil (top soil up to 10-30 cm depth

i.e. Horizon A), the microbial population is very high which decreases with depth of soil.

Each organism or a group of organisms are responsible for a specific change /

transformation in the soil. The final effect of various activities of microorganisms in the

soil is to make the soil fit for the growth and development of higher plants. Soil flora

(micro flora) e.g. Bacteria, fungi, Actinomycetes, Algae etc. Relative proportion /

percentage of various soil microorganisms are: Bacteria-aerobic (70%), Anaerobic (13%),

Actinomycetes (13%), Fungi/moulds (3%) and others (Algae Protozoa viruses) 0.2-0.8 %.

Soil organisms play key role in the nutrient transformations. In present study soil samples

from four locations from Bhor and Velhe were studied for their micro-flora in relation to

bio-chemical results (Plate 4.1).

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Table 4.2 Colony characters:

Sample

description 

Salekarvasti  Kiwat   Wanjale  Pabe 

Media  Luria Agar.  Luria Agar   Luria Agar  Luria Agar 

Size  0.5 cm  o.2 cm  0.1 cm  0.5 cn 

Shape  Circular  Circular  Circular  Irregular 

Colour  Off white  Off white  Chalky white  Off white 

Elevation  Flat  Flat  Flat  Flat 

Consistency  Butyrous  Butyrous  Butyrous  Butyrous 

Opacity  Opaque  Opaque  Opaque  Translucent 

Margin  Entire  Entire  Entire  Wavy 

Incubation time

and temp. 

24 hrs at 37 o c  24 hrs at 37 o c  24 hrs at 37 o c  24 hrs at 37 o c 

Gram staining  Gram postive

cocci 

Gram negative

short rods 

Gram negative

Short rods 

Gram negative

Short thin rods 

Motility  Non-motile  Motile  Non-motile  Motile  

Table 4.3Biochemical results:

Test  Kiwat  Salekarvasti   Pabe  Wangale 

Mannitol  No Acid gas   Acid  Acid  No Acid & gas 

Glucose  Acid  Acid & gas  Acid  No Acid & gas 

Maltose  No Acid & gas  Acid  No Acid & gas  No Acid & gas 

Xylose  Acid  No Acid & gas  Acid  No Acid & gas 

Cellobiose  No Acid & gas  No Acid & gas  Acid and gas  No Acid & gas 

Arabinose  No Acid & gas  No Acid & gas  No Acid & gas  No Acid & gas 

H2S production  Negative  Negative  Negative  Negative 

Starch

hydrolysis 

No hydrolysis  Positive  No hydrolysis  Positive 

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The results obtained from colony characters and biochemical findings and referring

standard Bergey’s Manual of determinative bacteriology. Bacteria like Alcaligenes,

Halomonas, Rhizobacterium Meriisculus may be present in the soil samples.

II. Cultivation practices of rice:

Rice crop has wide diversity in their characters, growth pattern and maturity from

60 to over 200 days at varied elevations. Bhor and Velhe regions lie in Western ghat region

of Maharashtra having elevation from sea level 1500 m. This area falls under heavy

rainfall to medium rainfall.

The major crop in both areas is Paddy/Rice production during the monsoon season

(Khariff). Local people prepare land for rice cultivation with local agricultural implements.

Land preparation for seedlings of rice:

Selection of land for seedling preparation is defined by each farmer depending

upon land holding. Raab is traditional slash and burn agricultural practice and is the core

economic activity of the local communities comprising Kunabis, Konkanas, Varlies and

Mahadev kolis.

Seedbed preparation is performed by farm yard manure or dry cow dung of first

layer, second layer of plant twigs, Butea monosperma (Lamk.) Taub., Calycopteris

floribunda (Roxb.) Poir., Holarrhena antidysenterica A.DC., Tectona grandis L., Bridelia

squamosa Gehrm. and Terminalia crenulata Roth. and third layer of different types of

grasses leaf litter. It is not uniform in all the regions of Bhor and Velhe. It changes

according to the availability of the plant material. The process of raab cultivation were

analysed by taking interviews of 20 farmers in both the regions.

After clearing of ash from the seedbed, seeds are sown 2-3 days after Rohini

Nakshatra is initiated in a bed of 40-50 sq. m. depending upon the land holder’s seedling

transplanting area.

Fields are ploughed a number of times in standing water (puddling). Puddling of

soil facilitates easy transplanting of seedlings. Generally, seedlings are one to one and 15

cm in length and spacing between two spots is 22.5 cm, and at each spot about four

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seedlings are transplanted. Under the wet system of cultivation ,continuous standing of

water into fields is observed (Plate 4.2).

III. Land races of rice grown in the region

India’s rice possesses wide diversity in their characters. They vary in maturation

pattern from 60 to over 200 days and can grow in varied elevations. At one extreme is the

deep water rice growing between 6 and 15 cm of water and at the other rice is grown with

an annual rainfall of barely 500 mm. This traditional practice was propagated based on the

intimate knowledge of rice varieties then prevalent in ancient India followed by the

varietals choice (Sarawgi and Rastogi, 2000). Maharashtra state has rich diversity of rice

due to variation in soil, climate and choice of local people. Kulkarni et al. (1998) reported

25 land races of rice like Kala rice (aromatic rice), Varangal, Rajguda, Kolambi, Tam,

Raibhog, Halva, Garva, and White rice collected from Western ghat of Maharashtra.

In many parts of the Bhor and Velhe regions, different varieties are still chosen for

needs, definitely not for the yields alone. They were chosen for their ability to withstand

droughts or floods, resistance to pests, susceptibility to disease, salinity tolerance, time of

maturity, size of the grains, colour, aroma, taste, keeping qualities, nutritional values. Early

maturity rice landraces are preferred by farmers. Patil et al. (2009) recorded different types

of landraces used by local people for mitigating climatic conditions in Bhor region.

Special land races were selected like Kolamba with early maturity and less water

requirement, Taka, Varangal, Jeera and Ekakadi which are drought resistant and suitable

in coarse sand. Kalbhat and Basamati are scented type of rice and generally grown in the

middle of the field due to destruction of crop before maturity by wild animal like Bison.

The yield of rice is very low but has economic value. Some improved rice varieties are

Indryani, Komal, Poonum, Rashipunam, Ratnagiri-24, Shan, Anupum, Indum,etc. Local

landraces and improved rice varieties are playing major role in agriculture of Bhor and

Velhe (Plate 4.3, 4.4).

IV. Weeds along the bunds of rice fields:

In many parts of India especially in Chhattisgarh ,rice bunds cover about 10% area

of rice fields. The bunds are a potential source of useful weeds. More than 60 weeds in

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bunds are identified as useful weeds (Oudhia, 1999a). In ancient Indian literature, it is

mentioned that every plant on the earth is useful for human being, crops and animals.

Bunds in rice fields are rich in biological diversity. These weeds can act as allelophathic

effects. The term Allelopathy means the injurious effects of one upon another. It represents

that the plant against plant aspect of the broader field of chemical ecology. Allelochemicals

which inhibited the growth of some species at certain concentrations may stimulate the

growth of same or different species at lower concentrations. Positive (inhibitory)

allelopathic effects of any weed on other weeds can be exploited to develop eco-friendly,

cheap and effective ‘green herbicides’. Green herbicides containing green allelochemicals

which are an integral part of eco or organic farming (Oudhia, 1999b). They represent the

floristic composition of the area and may act as green herbicide. In present ecological

study of rice, weeds along the bunds are documented.

Total 208 plant species were recorded on bund of rice field, which includes Herbs-

118, Shurbs-31, under Shrub-2, Trees-39, Twiner-5, Climber-13. In this 208 plants, 173

genera, 208 species belong to 63 families. Dominant family are Fabaceae 20 genera and 27

species, Asteraceae 19 genera and 20 species, Poaceae 13 genera and 14 species,

Euphorbiaceae 7 genera and 7 species, Convolvulaceae 6 genera and 8 species,

Lamianeace 5 genera and 8 species, Malvaceae 5 genera and 7 species, Verbanaceae 5

genera and 5 species, Caeaslpinaceae 4 genera and 6 species (Plate 4.5).

Bund weeds are classified according to family, genera and species to know the

richness of plant diversity along the bunds in table : Some Pteridophytes are also recorded

on bunds like Adintum philippense L., Actinopteris dichotoma Kuhn.

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Table 4.4 Family, Genera and Species wise weed distribution along Bunds: 

Sr. No.  Family  Genus Total 

Species1  Acanthaceae  3  5 

2  Agavaceae  1  1 

3  Amaranthaceae  5  6 

4  Anacardiaceae  2  2 

6  Ancistrocladaceae  1  1 

7  Annonaceae  1  2 

8  Apiaceae  1  1 

9  Apocynaceae  1  1 

10  Arecaseae  1  1 

11  Asclepidaceae  3  4 

12  Asteraceae  19  20 

13  Balsaminaceae  1  1 

14  Bombaceae  1  1 

15  Brassicaceae  1  1 

16  Caesalpinaceae  4  6 

17  Caricaceae  1  1 

18  Celastraceae  1  1 

19  Cleomaceae  1  2 

20  Combretaceae  2  4 

21  Commelinaceae  2  2 

22  Convolulaceae  6  8 

23  Cucurbitaceae  5  5 

24  Cyperaceae  1  2 

25  Elatinaceae  1  1 

26  Eriocaulaceae  1  1 

27  Euphorbiaceae  7  7 

28  Fabaceae  20  27 

29  Gentianaceae  2  2 

30  Lamiaceae  5  8 

31  Lecythidaceae  1  1 

32  Liliaceae  1  1 

33  Lytheraceae  3  3 

34  Meliaceae  1  1 

35  Malvaceae  5  7 

36  Martyniaceae  1  1 

37  Meliaceae  2  2 

38  Menispermaceae  1  1 

39  Mimosaceae  3  5 

40  Moraceae  2  2 

41  Moringaceae  1  1 

42  Musaceae  1  1 

43  Myrisinaceae  1  1 

44  Myrtaceae  3  3 

45  Nyctaginaceae  1  1 

46  Orchidaceae  1  1 

47  Oxalidaceae  1  1 

48  Papaveraceae  1  1 

49  Periplocaceae  2  2 

50  Plumbaginaceae  1  1 

51  Poaceae  13  14 

52  Polygonaceae  1  1 

53  Rhamnaceae  1  2 

54  Rubiaceae  1  1 

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Sr. No.  Family  Genus Total 

Species55  Rutaceae  1  1 

56  Santalaceae  1  1 

57  Sapindaceae  3  3 

58  Scrophulariaceae  4  5 

59  Solanaceae  3  5 

60  Teliaceae  3  5 

61  Verbenaceae  5  5 

62  Vitaceae  3  3 

63  Zygophyllaceae  1  1 

V. Weeds in rice field Weeds are group of plants of which are neither desirable nor economical values to

surroundings. They have special characteristic such as tolerance to environmental changes,

production of large numbers of seeds and high competitive ability, rapid growth through

vegetative phase to flowering, discontinuous germination and great longevity of seeds

(Monaco et al., 2002). Therefore understanding biology of weeds is very useful to improve

weed management strategies. In rice cropping system, knowledge of the biological

identification

distribution and life cycle of pre-harvest and post harvest as well as ecological aspects is

taken into consideration during the present study. Collection of pre and post harvest weeds

was made by using five quadrate of 1x1 m for each site. The weed diversity were recorded

and richness were calculated by quantitative estimation (Goel and Mitra, 2000) (Plate 4.6,

4.7) (Table 4.5 – 4.14).

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Table 4.5 Weed diversity of rice fields in Village - Karanje

  VILLAGE - KARANJE F RF D  RD A RA IVIPRE  Brachiaria reptans (L.) Gard 60 9.68 1.40  5.60 2.33 4.12 19.40HARVEST  Chloris baraata Sw.  40 6.45 1.80  7.20 4.50 7.95 21.60  Cyperus difformis L.  60 9.68 2.20  8.80 3.67 6.48 24.96  Dactyloctenium.aegyptium (L) Willd 40 6.45 0.80  3.20 2.00 3.53 13.19  Dichanthium annulatum (Forssk) Staf. 20 3.23 1.00  4.00 5.00 8.83 16.06  Digitaria stricta Roth ex R. & S. 40 6.45 2.80  11.20 7.00 12.37 30.02  Dinebra retroflexa (Vahl.) Panz. 40 6.45 1.40  5.60 3.50 6.18 18.24  Eragrostis uniloides (Retz.)Nees ex Steud 60 9.68 2.40  9.60 4.00 7.07 26.34  Impatiens balsamina L  100 16.13 5.60  22.40 5.60 9.89 48.42  Mnesithea granularis (L.) Koning & Soszf. 20 3.23 1.80  7.20 9.00 15.90 26.33  Mollugo oppositifolia L  20 3.23 0.80  3.20 4.00 7.07 13.49  Physalis longifolia Nutts 20 3.23 0.20  0.80 1.00 1.77 5.79  Smithia bigemina Dalz  60 9.68 2.40  9.60 4.00 7.07 26.34  Tridax procumbens L.  40 6.45 0.40  1.60 1.00 1.77 9.82     POST  Achyranthus aspera L  60 6.38 2.20  3.77 3.67 5.03 15.18HARVEST  Alysicarpus tetragonolobous Edg.J. 100 10.64 12.40  21.23 12.40 17.03 48.90  Ammania baccifera L.  100 10.64 15.60  26.71 15.60 21.42 58.77  Boerhavia erecta L.  40 4.26 0.40  0.68 1.00 1.37 6.31  Cardiospermum helicacabumL. 80 8.51 0.40  0.68 0.50 0.69 9.88  Celosia argentea L.  60 6.38 4.20  7.19 7.00 9.61 23.19  Crotolaria filipes Benth  100 10.64 3.80  6.51 3.80 5.22 22.36  Desmodium triflorum (L) D C 100 10.64 4.20  7.19 4.20 5.77 23.60  Digera muricata (L.) Mart. 20 2.13 0.20  0.34 1.00 1.37 3.84  Leucus celiata Benth  60 6.38 6.40  10.96 10.67 14.65 31.99  Parthenium hysterophorus L 80 8.51 3.60  6.16 4.50 6.18 20.85  Salvia aegyptiaca L  80 8.51 4.00  6.85 5.00 6.86 22.22  Sida acuta Burm. F.  40 4.26 0.60  1.03 1.50 2.06 7.34  Zornia gibbosa Span  20 2.13 0.40  0.68 2.00 2.75 5.56

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Table 4.6 Weed diversity of rice fields in Village - Karnawad

  VILLAGE - KARNAWAD F RF D  RD A RA IVIPRE  Biophytum sensitivium DC 60 20 2.60  20 4.33 13.20 53.20HARVEST  Celosia argentea L  40 13.33 1.60  12.31 4.00 12.18 37.82   Commelina subulata Roth 20 6.67 1.40  10.77 7.00 21.32 38.76   Cymbopogon citratus(DC.) Stapf. 40 13.33 1.20  9.23 3.00 9.14 31.70   Cynodon dactylon pers 40 13.33 3.00  23.08 7.50 22.84 59.25   Digitaria stricta Roth ex R. & S. 80 26.67 2.40  18.46 3.00 9.14 54.27   Dinebra retrotlexa (Vahl.) Panz. 20 6.67 0.80  6.15 4.00 12.18 25.00       POST  Ammannia baccifera L. 60 5.66 4.20  7.09 7.00 9.28 22.03HARVEST  Blumea solidaginoides (Poir.) Dc. 40 3.77 0.60  1.01 1.50 1.99 6.78   Boerahaia erecta L.  80 7.55 3.40  5.74 4.25 5.63 18.92   Celosia argentia L.  100 9.43 2.80  4.73 2.80 3.71 17.87   Chrozophora prostrata Dalz 80 7.55 3.60  6.08 4.50 5.96 19.59   Chrozphora rottleri (geis) Juss 40 3.77 1.00  1.69 2.50 3.31 8.78   Cleome chelidonii L. F. 80 7.55 3.40  5.74 4.25 5.63 18.92   Cleome felina L.F.  20 1.89 0.20  0.34 1.00 1.33 3.55   Eclipta prostrata (L.) L. 60 5.66 2.40  4.05 4.00 5.30 15.02   Euphorbia hirta L  80 7.55 7.40  12.5 9.25 12.26 32.31   Indigotera cordifolia Heyne ex Roth 20 1.89 0.20  0.34 1.00 1.33 3.55   Salvia aegyptiaca L  80 7.55 4.80  8.11 6.00 7.95 23.61   Spharanthus indicus L 100 9.43 18.40 31.08 18.40 24.39 64.90   Tridax procumbens L 80 7.55 3.60  6.08 4.50 5.96 19.59   Triomfetta rhombidea Jacq. 80 7.55 2.00  3.38 2.50 3.31 14.24   Xanthium indicum Kooen 60 5.66 1.20  2.03 2.00 2.65 10.34

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Table 4.7 Weed diversity of rice fields in Village - Kiwat

  VILLAGE - KIWAT F RF D  RD A RA IVIPRE  Ageratum conyzoides L.  40 9.52 1.00 6.58 2.50 7.37 2.35HARVEST  Boerahavia erecta L.  20 4.76 0.20 1.32 1.00 2.95 9.03   Cajanus scarabaeoides (L) Du petit Thou. 20 4.76 0.20 1.32 1.00 2.95 9.03   Cassia obtusifolia L.  40 9.52 0.60 3.95 1.50 4.42 17.89   Celosia argentea L.  40 9.52 0.60 3.95 1.50 4.42 17.89   Croton bonplandianus Baill. Adansonia 20 4.76 0.40 2.63 2.00 5.90 13.29   Cyperus rotundus L.  80 19.05 5.80 38.16 7.25 21.38 78.58   Dinebra retroflexa(Vahl)Panz 60 14.29 2.80 18.42 4.67 12.76 46.47   Lagascea.mollis cav.  40 9.52 0.60 3.95 1.50 4.42 17.89   Persicaria glabra (Willd) Gomez de la 20 4.76 1.40 9.21 7.00 20.64 34.61   Setaria pumila (Poir.) R. & S. 40 9.52 1.60 10.53 4.00 11.79 31.84      

POST  Alternanthera sessilis R.Br.Dc. 100 11.11 10.80 35.76 10.80 20.52 67.39HARVEST  Alysicarpus tetragonolobus Edg. 60 6.67 0.80 2.65 1.33 2.53 11.85   Amaranthus spinosus L.  20 2.22 0.60 1.99 3.00 5.70 9.91   Ammania baccifera L.  40 4.44 0.60 1.99 1.50 2.85 9.28   Caesulla axillris Roxb  80 8.89 2.80 9.27 3.50 6.65 24.81   Canscora diffusa(Vahl) R.Br.ex.R.&S. 40 4.44 1.00 3.31 2.50 4.75 12.51   Cassia absus L.  40 4.44 0.60 1.99 1.50 2.85 9.28   Cassia mimosoides L.  20 2.22 0.20 0.66 1.00 1.90 4.78   Crotolaria filipes Benth  40 4.44 1.60 5.3 4.00 7.60 17.34   Datura quercifoliaH.B.&K. 20 2.22 0.20 0.66 1.00 1.90 4.78   Flaveria trineriva (Sprenq) C.Mohr. 40 4.44 0.60 1.99 1.50 2.85 9.28   Impatiens balsamina L.  20 2.22 0.80 2.65 4.00 7.60 12.47   Ipomoea aquatica Forssk. 20 2.22 0.20 0.66 1.00 1.90 4.78   Launaea procumbens (Roxb.) Ramayya & Rajgopal 40 4.44 0.80 2.65 2.00 3.80 10.89   Solanum anquivi Lam.   20 2.22 0.20 0.66 1.00 1.90 4.78   Spharanthus indicus L  80 8.89 3.20 10.6 4.00 7.60 27.08   Trichodesma indicum (L.)Lenn 60 6.67 1.20 3.97 2.00 3.80 14.44   Trimfetta rhomboidea Jacq. 40 4.44 0.60 1.99 1.50 2.85 9.28   Verbascum chinese (L) Sant 80 8.89 2.40 7.95 3.00 5.70 22.54   Xanthium indicum Koen  40 4.44 1.00 3.31 2.50 4.75 12.51

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Table 4.8 Weed diversity of rice fields in Village - Nigudghar

  VILLAGE - NIGUDGHAR F RF D  RD A RA IVIPRE  Acalypa ciliata Forrsk 20 5.56 1.00  4.67 5.00 9.15 19.37HARVEST  Commelina paleata Hassk 20 5.56 0.80  3.74 4.00 7.32 16.61   Cynodon dactylon Pers 40 11.11 3.60  16.82 9.00 16.46 44.40   Eriocaulon cinereum R.Br. 40 11.11 4.60  21.5 11.50 21.04 53.64   Impatiens balsamia L. 60 16.67 3.00  14.02 5.00 9.15 39.83   Leucas indica R.Br.  40 11.11 1.40  6.54 3.50 6.40 24.06   Oxalis corniculata L  40 11.11 4.20  19.63 10.50 19.21 49.94   Parthenium hysterophorus L. 60 16.67 1.00  4.67 1.67 3.05 24.39   Smithia bigemina Dalz. 40 11.11 1.80  8.41 4.50 8.23 27.75       POST  Achyranthus aspera L 60 9.09 1.60  2.92 2.67 4.29 16.30HARVEST  Ageratum conyzoides L. 100 15.15 4.80  8.76 4.80 7.72 31.63   Alternanthera pungens HB&K 20 3.03 0.20  0.36 1.00 1.61 5.00   Alternanthera tenella colla Va tenella Veldk 100  15.15  11.00  20.07  11.00  17.68  52.91    Argemone mexicana L 20 3.03 0.40  0.73 2.00 3.22 6.98   Croton bonplandianus Baill.Adansonia 60 9.09 2.00  3.65 3.33 5.36 18.10   Cyperus rotundus L.  100 15.15 20.00 36.5 20.00 32.15 83.80   Euphorbia geniculata ortea 40 6.06 0.80  1.46 2.00 3.22 10.74   Malvastrum coromandelianum (L.) Garcke 20 3.03 0.20  0.36 1.00 1.61 5.00   Portulaca oleracea L 100 15.15 13.40 24.45 13.40 21.54 61.15   Solanum nigrum L.  40 6.06 0.40  0.73 1.00 1.61 8.40

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Table 4.9 Weed diversity of rice fields in Village - Salekarvasti

  VILLAGE - SALEKARVASTI F RF D  RD A RA IVIPRE  Acalypha indica Linn. 60 10.71 1.80  6.52 3.00 4.76 21.99HARVEST  Andropogon pumilus Roxv. 20 3.57 1.20  4.35 6.00 9.51 17.43   Arthraxon lanceolatus (Roxb) Hochst 40 7.14 1.00  3.62 2.50 3.96 14.73   Celosia argentea L  60 10.71 4.20  15.22 7.00 11.10 37.03   Commelina subulata Roth 20 3.57 1.20  4.35 6.00 9.51 17.43   Cynodon dactylon Pers 40 7.14 5.20  18.84 13.00 20.61 46.59   Cyperus rotundus L.  40 7.14 1.60  5.80 4.00 6.34 12.28   Eragrostis bifaria Wt. 20 3.57 0.80  2.90 4.00 6.34 12.81   Impatiens balsamina L. 80 14.29 2.60  9.42 3.25 5.15 28.86   Oxalis corhiculata L  40 7.14 2.60  9.42 6.50 10.30 26.87   Parthenium hysterophorus L. 60 10.71 2.60  9.42 4.33 6.87 27.00   Setaria pumila (Poir.) R. & S. 80 14.29 2.80  10.14 3.50 5.55 29.98       POST  Amaranthus spinosus L. 60 7.32 1.60  3.01 2.67 3.65 13.97HARVEST  Alternanthera sessilis (L) R.Br. 100 12.2 4.00  7.52 4.00 5.47 25.19   Amaranthus viridis L. 60 7.32 1.40  2.63 2.33 3.19 13.14   Caesulla axillris Roxb 100 12.2 9.00  16.92 9.00 12.31 41.43   Centaurium meyeri (Bunge) druce 80 9.76 8.40  15.79 10.50 14.37 39.91   Cyanodon dactylon Benth ex Dc 40 4.88 2.20  4.14 5.50 7.53 16.54   Exacum pedunculatum L. 20 2.44 2.00  3.76 10.00 13.68 19.88   Gomphrena serrata L. 40 4.88 0.60  1.13 1.50 2.05 8.06   Leucus longifolia Benth. 60 7.32 2.00  3.76 3.33 4.56 15.64   Ocimum aratissium L 80 9.76 2.00  3.76 2.50 3.42 16.94   Parthenium hysterophorus L 100 12.2 13.00 24.44 13.00 17.79 54.42   Verbascum chinese (L) Sant 80 9.76 7.00  13.16 8.75 11.97 34.89

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Table 4.10 Weed diversity of rice fields in Village - Ambavane

  VILLAGE - AMBAVANE F RF D  RD A RA IVIPRE  Alternanthera sessilis L. 20 5.56 0.60  3.7 3.00 7.74 17.00HARVEST  Chenopodium album L. 20 5.56 0.80  4.94 4.00 10.32 20.82   Cyperus difformis L.   80 22.22 4.20  25.93 5.25 13.55 61.70   Digera muricata (L.)Mart. 40 11.11 0.60  3.7 1.50 3.87 18.69   Echinochloa colona(L)Link 40 11.11 1.40  8.64 3.50 9.03 28.79   Eragrostis uniolodes(Retz)Neesex steud 40 11.11 2.00  12.35 5.00 12.90 36.36   Eriocaulon cinereum R.Br. 40 11.11 3.40  20.99 8.50 21.94 54.03   Fimbristylis dichotoma(L) Vahl 40 11.11 1.60  9.88 4.00 10.32 31.31   Panicum psilopodium Trin 40 11.11 1.60  9.88 4.00 10.32 31.31       POST  Cajanus scarabaeoides (L) Du. Petit. Thou 60 13.64 1.80  10.34 3.00 11.44 35.42HARVEST  Chorozophora prostrata Dalz 60 13.64 1.60  9.2 2.67 10.17 33.00   Crotolaria filipes Benth 20 4.55 0.20  1.15 1.00 3.81 9.51   Lagasca mollis 0Cav  20 4.55 0.20  1.15 1.00 3.81 9.51   Laucas celiata Benth  40 9.09 1.00  5.75 2.50 9.53 24.37   Ocimum gratissimum L 60 13.64 2.20  12.64 3.67 13.98 40.26   Parthenium hysterophorus L. 100 22.73 8.40  48.28 8.40 32.02 103.20   Xanthium indicum Koen 20 4.55 0.20  1.15 1.00 3.81 9.51   Zornia gibbosa span.  60 13.64 1.80  10.34 3.00 11.44 35.42

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Table 4.11 Weed diversity of rice fields in Village - Gunjavane

  VILLAGE - GUNJAVANE F RF D  RD A RA IVIPRE  Bacopa moneri (L.) Wettst. 20 5.56 1.80 12.16 9.00 18.85 36.57HARVEST  Brachiaria eruciformis (J.E.Sm.) Griseb 40 11.11 3.00 20.27 7.50 15.71 47.09   Brachiaria reptans(L)1Gard. 20 5.56 1.00 6.76 5.00 10.47 22.78   Caesulia axillaris Roxb. 40 11.11 0.60 4.05 1.50 3.14 18.31   Cassia absus L.  20 5.56 0.20 1.35 1.00 2.09 9.00   Celosia argentea L.  20 5.56 2.00 13.51 10.00 20.94 40.01   Eleusine coracana (L)Gaerth 20 5.56 0.80 5.41 4.00 8.38 19.34   Eragrostis cillaris (L)R.Br. 40 11.11 1.00 6.76 2.50 5.24 23.10   Lagasca mollis Cav  20 5.56 0.20 1.35 1.00 2.09 9.00   Smitha bigemina Ait Dalz 80 22.22 3.40 22.97 4.25 8.90 54.10   Sonchus oleraceus L. 40 11.11 0.80 5.41 2.00 4.19 20.70       POST  Alysicarpus tetragonolobus Edg.J. 40 10 1.00 6.25 2.50 6.94 23.19HARVEST  Centaurium meyeri (Bunge) Druce 60 15 1.40 8.75 2.33 6.48 30.23   Chenopodium murale L. 40 10 3.00 18.75 7.50 20.83 49.58   Cleome chelidonii L  60 15 1.40 8.75 2.33 6.48 30.23   Commelina subalata Roth 60 15 2.60 16.25 4.33 12.04 43.29   Cynodon dactylon Pers. 40 10 2.60 16.25 6.50 18.06 44.31   Ecilipta prostrata (L) L. 40 10 3.20 20 8.00 22.22 52.22   Euphorbia geniculata L 40 10 0.60 3.75 1.50 4.17 19.92   Ludwigia octovalvis (Jacq).Ravan 20 5 0.20 1.25 1.00 2.78 9.03

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Table 4.12 Weed diversity of rice fields in Village – Pabe

  VILLAGE - PABE F RF D  RD A RA IVIPRE  Acalypa cilliata Forssk 40 10.53 0.80 5.41 2.00 4.84 20.77HARVEST  Amaranthus spinosus L. 20 5.26 1.00 6.76 5.00 12.10 24.12   Argemone mexicana L. 20 5.26 0.20 1.35 1.00 2.42 9.03   Brachlaria reptans (L.) Gard. 60 15.79 2.60 17.57 4.33 10.48 43.84   Commelina diffusa Burm.f. 40 10.53 1.40 9.46 3.50 8.47 28.45   Cyanotis cristata(L.) D.Don. 20 5.26 1.40 9.46 7.00 16.94 31.66   Digitaria stricta Roth ex R. & S. 40 10.53 1.40 9.46 3.50 8.47 28.45   Dinebra retroflexa (Vahl.) Bor. 20 5.26 0.60 4.05 3.00 7.26 16.58   Eleusine coracana (L.) Gaerth. 20 5.26 0.80 5.41 4.00 9.68 20.35   Impatiens balsamia L. 80 21.05 4.00 27.03 5.00 12.10 60.18   Sprobolus spicatus(Vahl.)Kunth. 20 5.26 0.60 4.05 3.00 7.26 16.58       POST  Achyranthus aspera L 20 2.13 0.20 0.48 1.00 1.62 4.23HARVEST  Alternanthera sessilis (L) R.Br.ex.DC 60 6.38 3.00 7.18 5.00 8.12 21.68   Amaranthus roxburghianus Nevski 40 4.26 1.20 2.87 3.00 4.87 12.00   Cassia uniflora Mill  40 4.26 1.00 2.39 2.50 4.06 10.71   Corchorus olitorius L.ex.steud. 80 8.51 4.60 11 5.75 9.34 28.86   Crotalaria filipes Benth. 80 8.51 3.80 9.09 4.75 7.72 25.32   Cyperus deformis L.  100 10.64 11.80 28.23 11.80 19.17 58.04   Ecilipta prostrata (L) L. 40 4.26 1.00 2.39 2.50 4.06 10.71   Ergrostis unioloides (Retz.)Nees 80 8.51 2.80 6.7 3.50 5.69 20.90   Evolvulus alsinoides (L.) L 40 4.26 1.20 2.87 3.00 4.87 12.00   Exacum pedunculatum L. 60 6.38 2.40 5.74 4.00 6.50 18.62   Nicandra physalodes (L.) Gaerth 60 6.38 2.80 6.7 4.67 7.58 20.66   Plectranthus mollis (Ait) Spr. 80 8.51 1.80 4.31 2.25 3.66 16.47   Portulace oleracea L  60 6.38 1.60 3.83 2.67 4.33 14.54   Tridax procumbens L 40 4.26 1.00 2.39 2.50 4.06 10.71   Verbascum chinese (L) Sant. 60 6.38 1.60 3.83 2.67 4.33 14.54

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Table 4.13 Weed diversity of rice fields in Village - Vinzar

  VILLAGE - VINZAR F RF D  RD A RA IVIPRE  Cassia uniflora Mill.  20 5.26 0.40  3.7 2.00 8.96 17.92HARVEST  Echinochloa colona(L)Link 60 15.79 1.80  16.67 3.00 13.43 45.89   Eclipta prostrata (L.)L. 60 15.79 1.80  16.67 3.00 13.43 45.89   Eragrostis uniolodes(Retz)Neesex steud 60 15.79 2.20  20.37 3.67 16.42 52.58   Iscache globosa (Thunb) O.Ktze.Rev. 40 10.53 1.40  12.96 3.50 15.67 39.16   Portulaca oleracea L. 60 15.79 1.00  9.26 1.67 7.46 32.51   Prisicaria glabra (Willd) Gomez dla Maza. 40 10.53 0.80  7.41 2.00 8.96 26.90   Spheranthus indicus L. 40 10.53 1.40  12.96 3.50 15.67 39.16       POST  Alternanthera tenella .colla Veldk 80 14.81 2.60  14.77 3.25 6.62 36.21HARVEST  Amaranthus roxburghianus Nevski 40 7.41 0.40  2.27 1.00 2.04 11.72   Amaranthus spinosus L. 20 3.7 0.60  3.41 3.00 6.11 13.22   Argemone mexicana L. 20 3.7 0.20  1.14 1.00 2.04 6.88   Blainvillea acmella (L.) Philip 20 3.7 0.40  2.27 2.00 4.07 10.05   Blumea obliqua (L.) Druce 20 3.7 0.20  1.14 1.00 2.04 6.88   Boerahavia erecta L.  20 3.7 0.40  2.27 2.00 4.07 10.05   Canscora diffusa (Valh) R.Br. 20 3.7 1.00  5.68 5.00 10.19 19.57   Pogostemon deccanensis (Panigr.)Pers 20 3.7 0.20  1.14 1.00 2.04 6.88   Chloris barbata Sw.  40 7.41 3.20  18.18 8.00 16.30 41.89   Cleome chelidonii L.  20 3.7 0.40  2.27 2.00 4.07 10.05   Cynodon dactylon Pers. 40 7.41 3.80  21.59 9.50 19.35 48.35   Echinochloa colona (L) Link 60 11.11 1.20  6.82 2.00 4.07 22.00   Euphorbia hirta L.  40 7.41 0.80  4.55 2.00 4.07 16.03   Trimfetta rhomboidea Jacq. 60 11.11 1.40  7.95 2.33 4.75 23.82   Xanthium indicum Koen. 20 3.7 0.80  4.55 4.00 8.15 16.40

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Table 4.14 Weed diversity of rice fields in Village - Wanjale

  VILLAGE - WANJALE F RF D  RD A RA IVIPRE  Brachiaria eruciformis (J.E.Sm.) Griseb 80 26.67 3.40 32.69 4.25 20.00 76.36HARVEST  Caesulina axillris Roxb. 60 20 2.40 23.08 4.00 18.82 61.90   Cajanus scarbaeoides(L) Da Petit Thou 20 6.67 0.40 3.85 2.00 9.41 19.92   Digitaria stricta Roth ex R. & S. 40 13.33 2.20 21.15 5.50 25.88 60.37   Eriocaulon cinereum R.Br. 40 13.33 0.80 7.69 2.00 9.41 30.44   Nicandra physalodes (L.) Gaerten 20 6.67 0.20 1.92 1.00 4.71 13.30   Polygala erioptera DC 40 13.33 1.00 9.62 2.50 11.76 34.71       POST  Achyranthus aspera L. 20 4.55 0.60 4.62 3.00 12.71 21.87HARVEST  Ammania baccifera L. 100 22.73 3.80 29.23 3.80 16.10 68.06   Corchorus olitorius L. 20 4.55 0.20 1.54 1.00 4.24 10.32   Cullen corylifolia (L) Medik Vorles 20 4.55 0.60 4.62 3.00 12.71 21.87   Ecilipta prostrata (L) L. 40 9.09 0.40 3.08 1.00 4.24 16.41   Euphorbia hirta L.  40 9.09 1.00 7.69 2.50 10.59 27.38   Parthenium hysterophorus L. 100 22.73 3.80 29.23 3.80 16.10 68.06   Salvia aegyptiaca L.  40 9.09 1.40 10.77 3.50 14.83 34.69   Trichodesma indicum (L) Lenn 60 13.64 1.20 9.23 2.00 8.47 31.34

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Effect of weed control and weeds on rice and straw yield

In this connection, 1 x 1 plot were selected from 5 villages from Bhor and 5

villages from Velhe taluka. Weed index is low as 22.86 in village Karnawad, rice yield 270

gram with weeds and 350 gram without weeds. Higher weed index is 31.82 in village

Karanje, rice yield 300 gram with weeds and 440 gram without weeds.

In Velhe taluka, 5 villages were selected for rice yield with and without weeds.

Wanjale has 530 and 890 gram rice yield with weeds and without weeds respectively,

having weed index 40.45. On other hand maximum weed index is 51.11 in village

Ambavane having rice yield 220 gram with weeds and 450 gram without weeds.

Straw yield in without weed and with weed plots were recorded from Bhor taluka.

Village Kiwat has maxiumum 680 gram straw with weeds and 820 grams without weeds.

Similarly, Salekarvasti has minimum straw yield with 420 in with weeds plot and 700 in

without weeds plot

Velhe taluka has 5 village, Pabe village has minimum yield 90 gram straw in with

weeds and 260 gram without weeds. Similarly, 890 gram and 980 maximum yield of straw

in with and without weeds plots in village Wanjale.

Table 4.15 Yield of Rice

TALUKA  VILLAGE RICE YIELD

WITH WEEDS (GM)

RICE YIELD WITHOUT WEEDS

(GM)

WEED INDEX 

  Karnawad  270 350 22.86   Karanje  300 440 31.82   Kiwat  310 410 24.39   Nigudghar  220 310 29.03   Salekarvasti  270 360 25.00        VELHE  Ambavane  220  450  51.11    Gujawani  260  490  46.94    Pabe  50  90  44.44    Vinzar  240  440  45.45    Wanjale  530  890  40.45 

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Graph 4.1 Effect of weeds on rice yield in Bhor Taluka

Graph 4.2 Effect of weeds on rice yield in Velhe Taluka

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Table 4.16 Yield of Straw

TALUKA  VILLAGE  RICE STRAW YIELD WITH WEEDS (GM) 

RICE STRAW YIELD WITHOUT WEEDS (GM) 

  Karnawad  560 680    Karanje  610 950    Kiwat  680 820    Nigudghar  540 780    Salekarvasti  420 700        VELHE  Ambavane  450 480    Gujawani  490 670    Pabe  90 260    Vinzar  440 650    Wanjale  890 980 

Graph 4.3 Effect of weeds on straw yield in Bhor Taluka

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Graph 4.4 Effect of weeds on straw yield in Velhe Taluka

Experimental studies on viability of weed seeds was carried out in laboratory:

Most of the agricultural soils contain a large reservoir of weed seeds which

germinate over time. The number and type of seeds in the reservoir are determined by a

field’s cropping history, edaphic characters (moisture-holding capacity and pH), past weed

control practices, tillage and land preparation practices, rate of germination and seedling

growth of each species (Zamdahl et al., 1988). The soil seed bank is a product of the past

and represents the potential future of the above ground plant community. As a result, seed

banks are very important to the ecology of all types of communities (macro-phytes and

micro-phytes) and for recruitment of species, particularly those that rely mostly or totally

on non-vegetative means of reproduction.

Zamora and Thill (1989) stated that the soil seed banks are important components of

vegetation dynamics affecting both ecosystem resistance and resilience caused the viable

seeds on the soil surface and in the soil for many years comprised an enormous reservoir of

dormant and non-dormant seeds. There are important elements of seed bank dynamics like

seed bank depletion, unsuccessful germination, seed movement and seed mortality as a

result of ageing and attack by pathogens. The soil seed bank divided into surface seed bank

(active seed bank) and buried seed bank (dormant seed bank). Buhier et al. (1997),

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EFFECT OF WEEDS ON STRAW YIELD

Rice yield with weeds Rice yield without weeds

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mentioned that weed seed bank characteristic influence both the weed populations that

occur in a field and the success of weed management. Weeds produce a lot of weed seeds.

Most of the seeds entering the seed bank come from annual weeds (Rao, 2000). New seeds

may enter the seed bank through many sources, but the largest source is plants producing

seed within the field.

During the field survey, 10 sites were selected for collection of soil samples and

viability of weed seeds in laboratory was recorded. Some common monocot and dicot

weeds were growing in pot culture with given addition water for their survival. Common

weeds collected are: Smithia bigemina Dalz., Biophytum sensitivum DC., Polygala

erioptera DC., Alysicarpus teteragonolous Edgew., Alternanthera sessilis Br., Crotalaria

filipes Benth., Cassia absus L., Trichodesma indicum Br. Ageratum conyzoides L., Cassia

mimosoides L., Evolvulus alsinoides L., Corchorus olitorius L., Nicandra physeloides

Benth., Euphorbia hirta L., Celosia argentea L., Achyranthus aspera L., Oldenlandia sp.,

Eragrostis bifaria Wt., Commelina sp., Impatiens sp. , Arthraxon sp., Digitaria sp.,

Euphorbia sp., etc.

Table 4.17 Viability studies of Weed seeds

S. No. 

Villages  Total No. of Plants

Dicotyledons  Monocotyledons 

1  Salekarvasti  31 29 2 2  Karanje  60 - 60 3  Karnawad  36 2 34 4  Kiwat  16 11 5 5  Nigudghar  27 3 24 6  Pabe  22 - 22 7  Gunjavani  19 1 18 8  Wanjale  109 2 107 9  Vinzar  16 6 10 10  Ambavane  79 14 65 

Number of dicotyledons and monocotyledons germination observed after one year. In

village Wanjale soil maximum growth of weeds is seen i.e. 109 out of which 107 are

monocots. Soil sample of village Kiwat shows minimum weeds 16, out of which 11 are

dicot and 5 monocot and village Vinzar out of 16, 6 dicot and 10 monocot weeds. Pabe and

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Karanje soils have no dicot weed flora. This indicates that soil seed bank in above villages

from Bhor and Velhe taluka has no uniform seed viability.

Ash Value of rice husk and grains:

Rice (Oryza sativa L.) production is an important part of the national economy.

India is one of the world's largest producers of rice, accounting for 20% of all world rice

production. Moreover, the country has biggest area under rice cultivation, as it is one of the

principal food crops. Maharashtra is one of the major states in rice production; totally it

represented 1.48 M ha. land under rice cultivation which is 3.4% of total area of India and

gives 2.47 M.t of productivity. (Tiwari, 2002)

Demand for rice is growing every year and it is estimated that in 2010 and 2025

AD the requirement would be 100 and 140 million tones respectively. To sustain present

food self-sufficiency and to meet future food requirements, India has to increase its rice

productivity by 3 per cent per annum. (Thiyagarajan and Selvaraju, 2001) By considering

increase demand the productivity and maintained quality of economically impotent crop of

rice in India led the researchers to develop different cultivars and cultivation practices. The

performance of many quality characteristics depends greatly on environmental conditions,

sowing practices and weeds associated with it which alters the physicochemical properties

of rice (Araus et al, 1998 and Bryant et al, 2012)

Ash value/content is directly associated with environmental conditions and is of

great interest in defining optimal values of grain quality for a particular region. (Rharrabtia

et al, 2003 and Mansor et al, 2012) The perusal of literature shows very few studies have

taken into consideration the effects of cultural practices on weed distribution and rice

yields. Therefore, weed management strategies were chosen taking into account all the

factors affecting weedy rice populations in order to reduce the impacts of the weed to an

economically acceptable level.

In present study, Different five localities of Bhor and Velhe taluka from Pune,

Maharashtra were selected. The main objective of study was calculated effect of different

environmental conditions and different weed management practices on quality and

productivity of rice.

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Rice is one of the major economical crops in India. The grain quality is most

important parameter and having direct impact on acceptance of it in domestic and global

markets. The performance of many quality characteristics depends greatly on

environmental conditions and cultivation practices. In fact, grain ash value with respect to

total ash percentage, acid insoluble ash percentage and water soluble ash percentage are

important quality features for rice which influences under high-transpiration environments.

In cultivation practices i.e. application of fertilizer, weeding practices and water

availability shows correlation with quality and productivity of rice. The total ash value,

acid-insoluble ash value and water soluble ash value for each control and uncontrolled

grain samples were determined by standard method. The mean of six readings and standard

deviation was calculated.

Table 4.18 Ash Analysis of Rice Samples

TALUKA  VILLAGE  WATER SOLUBLE ASH ACID INSOLUBLE ASH

CONTROL SAMPLE CONTROL  SAMPLEBHOR  Karanje  5.34  5.29  4.57  4.59 

  Karnawad  5.48  5.26  4.63  4.63   Kiwat  7.19  5.86  5.53  5.82   Nigudghar  5.68  5.18  5.40  4.61   Salekarvasti  5.52  5.50  5.26  5.34            

VELHE  Ambawane  5.63  5.64  5.08  5.05   Gunjavani  5.24  5.27  4.77  4.78   Pabe  6.57  6.38  6.17  5.38   Vinzar  5.20  5.58  5.10  5.48   Wanjale  4.06  4.06  5.07  5.07 

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Graph 4.5 Water soluble ash

Graph 4.6 Acid Insoluble ash

0.00

1.00

2.00

3.00

4.00

5.00

6.00

7.00

8.00

KA

RA

NJE

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WATER SOLUBLE ASH

CONTROL SAMPLE

0.00

1.00

2.00

3.00

4.00

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6.00

7.00

KA

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ACID INSOLUBLE ASH

CONTROL SAMPLE

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Ash value

The comparative ash analysis shows that as the variation in environment and

weeding practices show positive correlation with ash value. In Bhor Kiwat show highest

ash value as compared to different locality listed in Table 1. and fig 1. Velhe, Pabe show

highest ash values compared to different locality listed in Table 2. and fig 2.

Other associates in rice field:

Algae The occurrence of blue green algae in cultivated paddy fields in India was first

reported in India in 1939. Studies in last 3-4 decades on this group of organisms have been

not only to understand their role in the conservation of nitrogen but also in the

understanding of the problem of nitrogen metabolism. The process of nitrogen fixation by

blue green algae is a highly oxygen sensitive reaction, can take place in the same cells that

release oxygen during photosynthesis. Ecologically these organisms are the most adaptive

in nature. They occur in all extremes of physically and nutritionally diverse habitats, both

aquatic and terrestrial ecosystems besides being present as air spores. In the aquatic

environ hey can be found as thermophilic and cryophilic forms in fresh and saline water.

Among the terrestrial forms these can be found in all types of soils and even in sub aerial

situations (Anand, 1993). Algal flora is an idea of utilizing it as biofertilizer as an

alternative or supplementary source of nitrogen for rice is a potential biological system

under low cost rice production technology. The resulting nitrogen fertility by algal

application has permitted moderate but constant productivity in fields where no application

of Nitrogen fertilizer (Kannaiyan, 1993). In this respect, algal flora from rice fields from

Bhor and Velhe has been documented with proper identification (Plate 4.7).

Chroococcus minutus (Kuetz) Naeg.

Cells spherical, hemispherical or oblong, single or in groups of 2-4; sheath

common as well as individual, distinct and colourless. Cells light blue in colour.

Occurrence: Paddy field of Hatnoshi, Taluka: Bhor.

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Gloeocapsa rupestris Kuetz

Cells in colonies, colonies many assigned irregularly in expanded mass. Individual

colony 2-4 celled, cells spherical with sheath, sheath yellow brown in colour with

concentric layers, cells blue green in colour.

Occurrence: Paddy field of Bare, Taluka: Bhor.

Aphanocapsa roseana de Bary.

Thallus blue green in colour, colonial, amorphous, mucilaginous, sheath

homogenous and colourless, cells rounded to oval loosely arranged.

Occurrence: Paddy field of Kari, Taluka: Bhor.

Oscillatoria princeps Vaaucher ex Gomont

Trichome straight, blue green or violet not constricted at cross walls, slightly

attenuated at the tip and bend. End cell slightly capitates, rounded with thick

membrane.

Occurrence: Paddy field of Vinzar, Taluka: Velhe

Nostoc punctiforme (Kuetz) Hariot.

Thallus light blue green, globose, filament densely entangled, sheath delicate,

hyaline, mucous trichomes broad, cells short, barel shaped. Cells are blue green

heterocysts subpsherical.

Occurrence: Paddy field of Margasani, Taluka: Velhe .

Anabaena variabilis Kuetzing ex Born et Flas.

Thallus light blue green, gelatinous, trichomes broad, constricted at cross walls.

Heterocyst nearly rounded akinetes away from heterocyst, smooth barrel shaped

long colourless.

Occurrence: Paddy field of Dhanep, Taluka: Velhe.

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Aulosira firtschii Bharadwaja

Thallus dark blue green, wooly, attached to the paddy leaves. Filaments

unbranched, broad straight, sheath colourless, trichome slightly constricted, light

blue green in colour. Heterocyst intercalary and single.

Occurrence: Paddy field of Nigudghar, Taluka: Bhor.

Scytonema chiastum Geitler.

Thallus blue green, filaments slightly curved, broad, single as well as multibranced.

Pseudobranches long narrower than main filament, constricted at cross walls,

heterocyst single.

Occurrence: Paddy field of Ambawane , Taluka: Velhe.

Gloeotrichia intermedia (Lemm.) Geitler

Thallus smooth, rounded, light brown in colour. Filaments very thin, sheath close

to trichome, brownish, trichome broad, construction at the joints, cells flattened at

the base and barraled shaped. Heterocyst spherical, single and broad.

Occurrence: Paddy field of Bhutonde, Taluka: Bhor.

Fungi in rice field:

Plant pathogens are major cause of diseases in crops and plants. These plant

pathogens devastated our crops and led us to devise methods and to search cultivars for

resistant. In case of rice crop several diseases are reported such as Bacterial Blight, rice

blast, Aggregate sheath spot, Black kernel Brown spot, Downy mildew, False smut, etc.

During the field survey in rice fields from Bhor and Velhe, the infected paddy samples

were collected during dry season shortly before harvest period. About half kilogram of the

samples were collected from different villages in both talukas. Collected samples were

labelled, packed in polythene and taken to the laboratory for further identification. The

samples were identified at Agharkar Research Institute in the department of mycology and

plant pathology group.

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Table 4.19 List of Fungi from Bhor and Velhe Taluka

Sr. No.  Name of Village Identified Remark 

1  Nazare   Nigrospora sphaerica (Sacc.) Mason 

2  Bare  Cladosporium oxysporum Berk. & Curt. 

3  Hatnoshi  Cladosporium tenuissimum Cooke  

4  Kari  Ephelis sp. 

5  Sakhar  Sporisorium sp. 

6  Adwali  Alternaria longissima Deighton & Mc Garvie

7  Vinzar  Volutella sp. 

8  Dapode  Drechslera sp. 

Bryophytes in rice field

The ecological distribution of many bryophytes is highly sensitive to factors that

affect plant water status. Many bryophyte species have evolved morphological structures

that enhance water uptake, water storage and/or limit water losses. This unique feature

plays significant role in natural ecological balance. Rice field ecosystem is a monoculture

agro-ecosystem. It is observed that Bryophytes play significant role in water conservation

during progressive growth of rice crop. Mosses play a role as inhibitors of soil erosion due

to their trample-resistant structure and their regenerative ability. Minerals supplied by rain

by leaching of the canopy and by through –fall are trapped by bryophytes, Thus bryophytes

play an important role in retaining minerals that might otherwise be quickly leached from

the soil. Bryophytes also play an important role in iron-ore deposition. Bryophytes are non-

vascular, thalloid plants adapted uniquely in their organization and physiology. Water is

primary requirement to keep these plants in correct and default physical and metabolic

state. Histological and physiological adaptations in thallus allows plants to absorb water

and food from remote sources such as rainwater, dews and leachates, soil surface, etc.

(Olarinmoye, 1974).

Rice fields are apparent habitats for bryophytes which grow on variety of the

substrates such as soil, rocks, trenches and walls around the fields, etc. Since bryophytes

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assist in maintaining moisture for a long time, their association with rice became an

important aspect to study. Maharashtra state has rich diversity of rice due to variation in

soil, climate and choice of local people. The association of bryophytes and the water

retention is known but such studies are not reported from the areas under study. The Bhor

and Velhe taluka of Pune district, Maharashtra are well known for the rice production. The

observations from this region revealed the presence of 7 bryophytes which contributes

towards water retention phenomenon (Gaikwad et al., 2012)

The rice field undergoes three major ecological phases - aquatic, semi-aquatic and

terrestrial dry, during a single paddy cultivation cycle. The aquatic phase of the rice field

represents the vegetative and the initiation of reproductive phase of the rice plant. While

the semi-aquatic and terrestrial dry phase correspond to the grain ripening stage of the rice

crop (Fernando, 1995). Thus, rice fields are dynamic environment that undergo rapid

temporal variation in relation to the growth of the rice plant and the hydrological status.

Therefore, as a whole, the ecology of rice fields is dominated by rapid physical, chemical

and biological changes.

The surveys were undertaken during years (2009-2011) from Bhor and Velhe

region during different stages of rice crop. The documentation of life cycle of Bryophyte

and rice crop was made periodically. The mature plant samples were collected in clean

plastic bags to establish their identity. The samples were brought in laboratory, dust was

removed and clean samples were identified using literature (Biradar & Shirke, 1992). The

specimens were prepared and mounted with standard technique (Azuelo, 2010). The

specimens were deposited in Agharkar Herbarium at Maharashtra Association for

Cultivation of Sciences, Pune (AHMA). Photodocumentation was carried out using

‘Olympus C7070’ digital camera. The observations were recorded systematically.

Descriptions of the specimens:

• Family: Ricciaceae

• Riccia cruciata Kash - Thalli monoecious, spongy cruciform when young, in

rosettes, 5 to 7 x 2 to 3 mm. dorsal surface light green at maturity, usually two

times dichotomously branched, obovate to oblong, dorsal groove shallow.

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Epidermis thin, scales are small, delicate purple weak, semi lunar. Rhizoids mostly

smooth, few, tuberculate. Antheridia globular with inconspicuous

papillae.Archegonia in a row, papillae prominent.

Locality: Bhor-Bare; Velhe-Dapode

• Riccia melanospora Kash - Plants are monoecious, small, greyish, dichotomously

brownish in overlapping patches. Thalli are 2.5 to 4.5 X 1 to 2 mm. Segments

linear, ciliate, cilia along the margins near the apical portion, dorsal groove deep &

narrow near the apex or slightly concave towards the posterior side; scales

prominent, overlapping, deep purple. Rhizoids usually smooth, sometimes

tuberculate. Antheridia are in mid dorsal row, with conspicuous papillae,

sporophytes embedded in thallus seen prominent on the dorsal surface.

Locality – Velhe – Ambavane, Margasani

• Riccia gangetica Ahmad - Plants are monoecious, dorsal surface bluish green,

margin purple when young, whitish at maturity. Thalli 1.5 to 2.5 X 1 to 1.5 mm,

one to two times dichotomously branched. Thalli linear-obovate, dorsal grove

conspicuous. Ventral scales prominent purple. Rhizoiids many, smooth &

tuberculated, Antheridia and archegonia are in 1-3 rows along with mid dorsal

groove. Antheridial papillae prominent hyaline projecting above the surface of the

thallus. Archegonial neck prominent, hyaline protruding above thallus surface.

Locality – Velhe- Margasani

• Riccia billardieri Mont. & Nees ex Gottsche, Lindenb. & Nees- Thalli

monoecious, once or twice forked overlapping or in incomplete rosette. Thalli

usually 4 to 12 X 2 to 3 mm. dorsal groove narrow anteriorly, which gradually

becomes flat on posterior side, epidermal cells hyaline and oval. Ventral scales

prominent, purple, semilunar rhizoids many, ascending from mid ventral side, 8 to

12 mm long, both the types simple & tuberculate. Cross section of thallus 4 to 6

times as broad as high, Antheridia in 1 to 3 rows ostioles conspicuous, projecting

above the surface of thallus, mostly on midrib, archegonia in rows, projecting

above thallus.

Locality – Bhor- Hirdoshi, Venwadi, Nandgaon

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• Riccia crystallina L. - Plants monoecious forming rosettes 12 to 20 mm in

diameter, May be overlapping, Generally yellowish green or reddish, Thalli spongy

6 to 12 mm long, dichotomously branched dorsal surface of matured thalli pitted

due to this organization of the epidermis, pits visible, dorsal grove inconspicuous

except at the apex, rhizoids mostly smooth few tuberculate.

Locality – Velhe- Dhanep

2) Family: Targioniaceae

a. Cyathodium tuberosum Kash - Individual plant is dioecious, 8 to 10 mm long &

the rosette is about same diameter; thallus membranous, fan shaped with entire

margin, dichotomous branching resulting into lobes. On the ventral side, smooth

rhizoids are evenly spread all along the surface of the thallus except margin, 8 mm

long; hyaline lines are more distinctly seen on ventral side except margin & tip.

Antheridial chambers are marginal in position to the distal end of the thallus on

dorsal side while the archegonia develop on ventral side individually capsules

develop in the fold of thallus appear black in colour at maturity.

Locality - Bhor- Ambeghar; Velhe- Dapode.

3) Family: Anthocerotaceae

• Anthoceros erectus Kash - Plants dioeciously erect grouped into tufts thick fleshy

spongy with a basal stalk expanding at the top into an open palm like structure,

female thalli are large 10 X 7 mm, stalk is shorter as compared to that of the male

thallus, male plants intermixed with the females, the inflated portion is cup like in

male thalli while it is funnel shaped in female thalli, margin of both male and

female thalli are highly dissected to form finger like projecting lobes.

Locality – Velhe - Ambavane

Bryophytes found to be grown on wet rocks, soil, vertical walls retaining the

moisture even after drying of the soil. R. gangetica mostly found to be associated

with R. billarclieri, C. tuberosum grows forming dense mats. These mats on

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vertical walls as well as ground field inhabits only in shade portions. All the species

showed their sporophytic phases at the time of harvest of the crop.

These species commonly shows strong association with Algal communities. The

association of Nostoc spp. with Anthoceros spp. enhances the formation of

hormogonia in Nostoc spp. (Campbell & Meeks, 1989). It also been reported that

the symbiotic association of Cyanobacteria helps in nitrogen fixation (Saxena,

1981). This helps to improve the quality and quantity of Rice.

During present studies, seven species belonging to three genera and three families

were recorded. Out of these, family Ricciaceae includes one genus and five species,

Targioniaceae and Anthocerotaceae includes one genus and one species each.

Floristic analysis has been complied in Table 1.

The association of Bryophytes retains the water and enriches soil of the rice fields. This is

a first report of Bryophytes from the region of Maharashtra in rice fields.

Table 4.20: Floristic analysis of Bryophytes

No.  Family  Species  1. 

Ricciaceae 

Riccia cruciata Kash  2.  Riccia melanospora Kash 3.  Riccia gangetica Ahmad  4.  Riccia billardieri Mont. & Nees ex Gottsche, Lindenb. & Nees5.  Riccia crystallina L. 6.  Targioniaceae  Cyathodium tuberosum Kash  7.  Anthocerotaceae  Anthoceros erectus Kash  

Pteridophytes:

Pteridophytes are vascular cryptogams and form a neglected group of plants in

biodiversity as far as their economic value is concerned. It can safely be assumed that they

were the first land plants with a vascular system. Their food and medicinal values are

reported in earlier literature. The Ayurvedic systems of medicine referred by Sushruta (ca

100 AD) and Charka (ca 100 AD) recommended the medicinal uses of some ferns in their

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Samhitas. Ferns are also used by the physicians in Unani system of medicine (Uddin et al.,

1998).

Rice fields are apparent habitats for pteridophytes which grow on variety of the

substrates such as soil, rocks, trenches and walls around the field. Since pteridophytes

support in maintaining moisture for longtime, their association with rice become an

important aspect to study.

In the present study survey of pteridophyte in the rice fields from Bhor and Velhe

tauka including its identification, determination of micro habitat in rice fields and their

association was carried out. The documentation of life cycle of pteridophyte and rice crop

was made periodically. The plants were identified using literature. The specimens were

prepared and mounted with standard technique

The specimens were deposited in Agharkar Herbarium at Maharashtra Association for

Cultivation of Sciences, Pune (AHMA).

During field survey following specimens were recorded

1] Adiantum lunulatum Burm. f.

Family :- Adiantaceae

Roots are very thin, fibrous, about 10-15 cm long, reddish-black in colour, soft and

branched. Rhizome - Long, upto 2 mm thick, glabrous, prostrate or erect, dark reddish-

brown or a black in colour.

Frond - Rachis shiny black, simply pinnate, pinna roughly lunulate, subdimidiate, lower

edge nearly in line and oblique with its black shiny petiole, upper edge bluntly rounded

and more or less lobed, a few sori in a continuous line on the under surface along the edge,

with a false indusium

2] Marselia quadrifolia Linn.

Family : Marseliaceae

Aquatic fern bearing 4 parted leaf resembling '4-leaf clover' (Trifolium). Leaves

floating in deep water or erect in shallow water or on land. Leaflets obdeltoid, to 3/4" long,

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glaucous, petioles to 8" long; Sporocarp (ferns) ellipsoid, to 3/16" long, dark brown, on

stalks to 3/4" long, attached to base of petioles.

3] Selaginella miniatospora (Dalz.) Bak.

Family : Selaginellaceae

Stems elongated, with numerous small leaves; creeping, and rooting from characteristic,

leafless, root-bearing branches (rhizophores); overtly dichotomising vegetatively;

dorsiventrally organized, with four ranks of leaves (with two rows of leaves dorsally, and a

row of larger leaves down each side); with strongly flattened branches; without secondary

thickening. Leaves ligulate; 4-ranked on the branches; of two kinds: those of the two ranks

on the upper side of the stem appressed and directed towards the stem apex, and those of

the two lower ranks larger and spreading laterally; about 1 mm long (on the upper side of

the stem), or 2 mm long (on the sides); appressed and spreading.

Heterosporous.Sporophylls ovate, cuspidate, keeled; aggregated into well defined terminal

cones. Cones sessile at the tips of the normal shoots (short, 4-sided). The sporangia basal

and subsessile on the adaxial surfaces of the sporophylls, non-septate.

4] Adiantum capillus - veneris L.

Family : Adiantaceae

The Adiantum capillus-veneris fern grows from 6 to 12 in (15 to 30 cm) in height;

its fronds arising in clusters from creeping rhizomes 8 to 27.5 in (20 to 70 cm) tall, with

very delicate, light green fronds much subdivided into pinnae 0.2 to 0.4 in (5 to 10 mm)

long and broad; the frond rachis is black and wiry.

5] Actinopteris dichotoma Kuhn.

Family : Adiantaceae

This species is found in hot, dry habitats, growing at the base of rocks and in

crevices, but also on deep soil in shady places in rice fields. The distinctive fronds issue

from a prostrate rhizome and are somewhat dimorphic. The laminae are 15-45 mm long

and fan-shaped with a spread of some 180 degrees, the tip of each segment bearing 2-5

teeth.

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Pammel (1911) compiled a manual of poisonous plants from Eastern North

America with brief notes on economic and medicinal plants which included some

medicinal pteridophytes. In Chinese system of medicine, many ferns are also prescribed by

local doctors (Kimura and Noro, 1965). Later on modern biological and pharmaceutical

studies were carried out on pteridophytes by different scholars. Banerjee and Sen (1980)

conducted the only extensive survey of antibiotic activity among the ferns and reported

about a hundred species having such property. Dixit and Vohra (1984) reported edible and

medicinally important pteridophytic species from India. Kaushik (1998) emphasized on the

ethno-botanical importance of ferns of Rajasthan, India. The ethno-botanical uses of this

unique group are of immense importance (Singh et al., 1989; Dhiman, 1998). The most

important studies on food and medicinal values of pteridophytes were conducted by Nayar

(1957), Hodge (1973) and Dixit (1974, 1975). Recently, Ghosh et al. (2004) reported some

edible pteridophytes as vegetables and medicines.

Exotic invasive weeds in rice fields:

General definition of weeds as unwanted plants grown in cultivated crop and

compete with storage of food material in soils. Classification of weeds is based on

indigenous weeds and exotic weeds established since long time and naturalized in our

soils. These exotic weeds introduced from other countries known as invasive weeds. These

weeds plants have escaped from its original ecosystem and are established in regional

floras. They are spread in the rice fields and displace native biota and threaten valued

agricultural crops and environment due to profuse growth.

During the weed ecological survey 44 invasive weeds were recorded. Exotic

invasive weeds are 44 occurred in Bhor and Velhe regions. They belong to 17 species, 39

genera, 44 species. All are herbs native to Brazil, Central, south America and few from

Africa, East Indies and Asia. Maximum weeds are from Asteraceae, Ameranthaceae,

Euphrobiaceae, Poaceae, Cyperaceae, Fabaceae families. Aches or seed of Asteraceae

family are easily dispersed from one region to another by wind or transportation of food

grains. .

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INVASIVE MEDICINAL WEEDS FROM RICE FIELD:

The data generated from 20 local farmers using weeds as medicines. In this respect

19 weeds were recorded having properties like anthelmintic, anti-inflammatory, anti-

tumour activity diuretic, astringent, tonic etc. Local people use some plant parts for kidney

stone, jaundice, skin diseases, cough and cold, piles, fever, burns, urinary disorders, etc.

Ageratum conyzoides L .

Family - Asteraceae Habit - Herb Local name - Sahdevi, Osadi

Part used - whole plant

Medicinal use - Root juice is anthelminthic.

A paste of root mixed with the bark of Schinus wallichi L. is applied on dislocated

bones.

The leaves are styptic. The dried leaves powder is applied on cuts sores and

ruftures caused by leprosy. A past of leaves is used as poultice to remove thorn

from skin.

The juice of flower heads is used externally to treat scabies.

Alternanthera tenellacolla

Family - Amaranthaceae Habit - Herb

Part used - Whole plant

Medicinal use - Aqueous extract of A. tenella is used for anti-inflammatory activity

& anti-tumour activity

It has antimicrobial & antiviral properties.

Argemone mexicana L.

Family - Papaveraceae Habit - Herb Local name - Pivala dhotra

Part Used - Root bark and stem latex

Medicinal use - Fresh bark or dried bark with betel leaves for jaundice & latex for

skin

disease& eye diseases .

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Bidens pilosa L.

Family - Asteracee

Part used - Leaves Habit - Herb

Medicinal use - The leaf extract is used as diuretic, inflammation of digestive tract

and anti-diarrhoeal. Also leaves are used astringent.

Cassia uniflora Miller

Family - Caesalpinaceae Habit - Herb Local name - Takala

Part Used - Leaves &Seeds

Medicinal uses - Seed power is used for wound healing. Leaf juice is used to cure

dysentery & diarrhoea.

Celosia argentea Linn.

Family - Amarantheceae Habit - Herb Local name - Kurdu

Parts Used - Whole plant

Medicinal Use - Plant extract is used to cure dysentery, diarrhea, acute abdominal

pains, inflamed stomach and treatment of tuberculosis. Leaf juice is applied on

wounds, sores, ulcers, skin eruptions, ulcers of mouth & relieves itching. Root juice

is given early in the morning for cure kidney stone. Drops of leaf juice are used for

tired eyes to clear vision.

Chrozophora rottleri (Geis.) Spreng.

Family - Euphorbiaceae Habit - Herb

Part used - Fruits

Medicinal Uses - The juice of fruit is used to cure cough &cold.

Croton bonplandianum Baill.

Family - Euphorbiaceae Habit - Herb

Part used - Seeds

Medicinal use - Seeds are purgative.

Dinebra retroflexa (Vahl.) Panz.

Family - Poaceae Habit - Herb Local name - Meshkathi

Part used - Root & culm

Medicinal use - The culm is used in form of aqueous paste for face care and

decotion of root is used externally as well as internally in treatment of piles.

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Echinochola crusgalli (L.) Beauv.

Family - Poaceae Habit - Herb

Part used - whole plant

Medicinal use -The root powder is applied as a styptic to wounds. The leaf extract

is used as tonic & treating carbuticles sores spleen troubles cancer & wounds.

Eclipta prostrata (L.) Mant.

Family - Asteraceae Habit - Herb Local name - Maka

Part used - Leaf

Medicinal use - Leaf extract is applied on hair for strengthening them, making

black and prevents falling off. The farmers and tribal apply the leaf juice on the

cuts and wounds. There is popular opinion that the herb taken orally and applied

externally will turn the hair black. The plant extract is applied to the body in fevers.

Euphorbia hirta Linn.

Family - Euphorbiaceae Habit - Herb Local name - Dudhani

Parts used - Whole plant

Medicinal use - Plant decoction is given orally in the treatment of asthma,

bronchitis, coughs bowel complaints stomach ache & dysentery. Leaf paste is

applied in burns and scorpion stings. Leaf extract is consumed orally in gonorrhoea

and snake bites. A drop of latex is poured in eye for eye complaints and is applied

on eczema and scabies.

Leaf juice is given for the treatment of acute abdominal pains, infection of the

mouth and to Stops local bleeding.

Euphorbia indica (Lam.) Croizt.

Family -Euphorbiaceae

Oxalis corniculata L.

Family - Oxilidaceae Habit - Herb Local name - Ambushi

Part used - whole plant

Medicinal use - The whole plant is anthlelmintic, anti-phlogistic, astringent,

depurative, diuretic, emmenagogue, febrifuge, lithontripic, stomachic & styptic. It

is used in treatment of influenza, fever urinary tract infection enteritis diarrhoea

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boils & pimples. The leaf juice applied for insect bites burns and skin piton 9t has

an antibacterial activity

Parthenium hysterophorous L.

Family - Asteraceae Habit - Herb Local name - Congress, Gajargavat

Part used - whole plant

Medicinal use - Root decoction is used in dysentery and used as tonic febrifuqe &

emmeriagogue. Parthenium is also reported as promising remedy against hepatic a

mebiosis. Parthnenin a toxin of Parthenium is found pharmacologically active

against neuralgia and certain types of rheumatism.

Portulaca oleracea Linn.

Family - Portulacaceae Habit - Herb Local name - Ghol

Part used -Whole plant.

Medicinal Use - Plant juice is taken orally to cure Liver kidney, bladder problems

& scurvy. It is also useful to increases milk flow in nursing mothers, good for

painful or difficult urination, relieves dry coughs, shortness of breath &

immoderate thirst cures inflamed eyes mouth sores. It is applied on swollen gums

fasten loose teeth. Leaf juice is applied for relieves pain from burns, healing of

wounds, promotes flow of urine menstrual flow, help in digestion & expulsion of

intestinal worms.

Sida acuta Burm.f.

Family - Malvaceae Habit - Herb

Part used - Root

Medicinal use - Root extract is used as rejuvenating nerve tonic & degenerative

disorders.

Tribulus terrestris L.

Family - Zygophyllaceae Habit - Herb Local name - Sarata, Gokharu

Part used - fruits, leaves and roots.

Medicinal uses - Fruits are used as tonic. Fruits power along with wheat flour is

given orally backaches and other complaints to ladies after delivery. Fruit powder

is also given orally in urinary diseases. Leaf juices are referred orally in stomaches.

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Tridax procumbens Linn.

Family - Asteraceae Habit - Herb Local name - Kurmudi

Part used - Leaves

Medicinal Use-Leaf juice is applied to check the hemorrhage of wounds and

Bronchial catarrh which is an inflammation of the mucous membrane with thick

phlegm .Leaf juice is given to cure dysentery & diarrhoea.

Xanthium strumarium L.

Family - Asteraceae Habit - Herb Local name - Landaga

Part used - Leaves, roots flowers and fruits

Medicinal uses - Plant is given to domestic animals for promoting milk yielding

capacity. Decoction of plant is given orally in leucorrhoea, menorrhagia and in

malarial fevers. Fruits extract is applied on small pox and are having cooling effect.

Flower tincture is useful in toothaches.

List of insects, vertebrates and invertebrates in rice field:

In case of rice fields many insects feed at different stages of growth. Grasshoppers belong

to the super family Acridoidea and Pyrgomorphoidea of the order Orthoptera and suborder

Caelifera. Acridoidea shows maximum diversity and constitutes only one family i.e.,

Acrididae whereas pyrgomorphoidea also has only one family i.e., Pyrgomorphidae,

widely distributed in India. Grasshoppers are of great economic importance, because they

constitute an important group of pests and pose a constant threat to cereal crops, pulses,

vegetables, orchards, grassland and forest plantations all over the world. Grasshoppers

cause significant damage to tree seedlings and agricultural crops (Joshi et al., 1999), hence

considered as oligophagous and according to host preferences classified as graminivorous,

forbivorous and ambivorous or mixed feeders (Mulkern 1967). Grasshopper can damage

rice in all stage of crop growth. Both nymphs and adults can feed on leaves by cutting the

edges of leaves. When found in greater number can feed even midribs and total leaves and

cause extensive defoliations.

However in present study some invertebrate pests attack on rice crop during the initial

germination and establishment phase of growth.

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1. Long horned grass hopper (adult one)

2. Robber flies

3. Meloid bettle (Mylabris sp)

4. Apis cerana (honey bees)

5. Wasp (unknown sp)

6. Cockroches (Pycnoscleus sp)

7. Earwia (probably Anrsolabis)

8. Chrysomelidae beetle (leaf eating bettle)

9. Homopteran bugs

10. Fruit fly (unknown sp)

11. Fishes

12. Earthwarms (Eukerria saltensis)

13. Aquatic snails (Isidorella newcombi): Grazing on young seedling of rice.)

14. Rodents.: These are serious pest for rice crops

15. Crabs & Frogs