Biological Assessment of Arctic Char (Sa/velinus a/pinus L.) Stocks and Su~mary of the Atlantic

. Salmon (Sa/rna sa/ar L.) Fishery In Northern Labrador

J. Brian Dempson

i

Research and Resource Services Directorate Fisheries and Marine Service Department of Fisheries and the Environment P. O. Box 5667 St. John's, Newfoundland A1C 5X1

O(ftober 1978 J

Fisheries & Marine Service Technical Report No. 817

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Fisheries and Marine Service

Technical Report 817

Oc tober 1978

BIOLOGICAL ASSESSMENT OF ARCTIC CHAR (SALVELINUS ALPINUS L.)

STOCKS AND SUMMARY OF THE ATLANTIC SALMON (SALMO SALAR L.)

FIS~ERY IN NORTHERN LABRADOR

by

J. Brian Dempson

Research and Resource Services Directorate

Fisheries and Marine Service

Department of Fisheries and the Environment

P.O. Box 5667

St. John's, Newfoundland A1C 5Xl

This is the fifty-second Technical Report from the

Research and Resource Services Directorate, St. John's

i i

•

(c) Minister of Supply and Services Canada 1978

Cat. no. Fs 97-6/ 52 ISSN 0701-7626

iii

CONTENTS

Abstract iv

Introduction 1 l"1aterials and methods 1

Commercial landings 1

Commercial sampling 4 Mortality 4

Growth 5

Yield per recruit 5 Results and discussion 6

. Commercial landings 6

Size breakdown and weekly landings of char and salmon 6

Landings of Arctic char and Atlantic salmon by area 6

Catch and effort 16

Commercial sampling 17

Age and length composition 17

Mortality 20

Growth 23

Yield per recruit 26

Summary 30

Acknowledgments 31

References 31 Appendix 35 Appendix II 43 Appendix III 50 Appendix IV 51

Appendix V 54

iv

ABSTRACT

Dempson, J. Brian. 1978. Biological assessment of Arctic char (SaZveZinus alpinus L.) stocks and summary of the Atlantic salmon (SaZmo saZar L.) fishery in northern Labrador. Fish. Mar. Servo Tech. Rep. 817: v + 54 p.

The status of Arctic char stocks in northern Labrador is evaluated using the Beverton and Holt (1957) yield per recruit model. Von Bertalanffy growth curves, fitted to age and length data from Voisey, Anaktalik, Tikkoatokak, Napartok and Hebron areas, supplied parameters for this assessment. Estimated levels of F (FO. l ) ranged from 0.31 to 0.39.opt Total instantaneous mortality rates, ranging from 0.38 at Hebron to 0.85 at Tikkoatokak Bay, are derived from catch curves and indicate that present rates of fishing mortality exceed optimal levels in Voisey and Anaktalik bays, while surpassing the maximum value (F ) in Tikkoatokak max Bay. Arctic char landings increased substantially in 1977 to 210,406 kg. Total effort rose by 52% while the catch per unit of effort diminished by 20%.

The intensity and distribution of fishing effort directed toward Arctic char is partially related to the abundance of Atlantic salmon on the northern Labrador coast. Therefore, the salmon fishery, which totalled 41,618 kg, is briefly discussed and a summary is provided for landings of Arctic char and salmon by area, size category and by week.

Key words: Arctic char, catch and effort, growth, mortality, yield per recruit, Labrador.

, , RESUME

Dempson, J. Brian. 1978. Biological assessment of Arctic char (SaZveZinus aZ~inus L.) stocks and summary of the Atlantic salmon (SaZmo saZar L.) fishery in northern Labrador. Fish. Mar. Servo Tech. Rep. 817: v + 54 p.

Le modele de rendement par recrutement, de Beverton et Holt (1957), sert a evaluer 1 letat des stocks d'Omble chevalier du nord du Labrador. Les courbes de croissance de Von Bertalanffy, mises en rapport avec les donnees sur l'age et sur la longueur des poissons des regions de Voisey, Anaktalik, Tikkoatokak, Napartok et Hebron, ont fourni les parametres de cette evaluation. Les niveaux evalues de (FO,l) ont varie deFopt 0,31 a 0,39. Ce sont les courbes des prises qui ont fourni les coefficients instantanes de mortalite totale, variant de 0,38, pour Hebron, a 0,85, pour la baie Tikkoatokak; ces derniers indiquent que les coefficients actuels de mortalite due a la peche depassent les niveaux optimaux calcules pour les baies Voisey et Anaktalik, ainsi que la valeur maximale (F ) de la baie Tikkoatokak. Les debarquements d'Omble chevalier ontmax augmente considerablement en 1977, atteignant 210 406 kg. L'effort total a connu une hausse de 52% tandis que la prise par unite d'effort a diminue de 20%.

v

L'intensite et la distribution de 1'effort de peche de 1 'Omble chevalier sont liees en partie ~ 1'abondance du Saumon de 1 'Atlantique sur la cote nord du Labrador. L'article traite donc brievement de la peche du Saumon, qui a atteint 41 618 kg, et donne un aper~u des debarquements d'Omble chevalier et de Saumon, par region, par categorie de taille et par semaine.

Mots-cles: Omble chevalier, prise et effort, croissance, mortalite, rendement par recrutement, Labrador.

INTRODUCTI ON

Relatively little information is available pertaining to the biology of Arctic char in Labrador. Andrews and Lear (1956) examined age, length and weight composition, described feeding and parasitism and examined meristic variation in commercial samples from various populations along the coast. Peet (1968, 1971) collected age, length and weight data and also assessed meristic variations with northern Labrador populations from reconnaissance survey sampling and counting fence operations on several rivers in southeast Labrador. Similarly Murphy (1972, 1974) collected basic biological data from counting fence operations on Northwest Tributary of Sand Hill River and from the commercial fishery conducted in that area of southeast Labrador. Coady and Best (1976) described growth characteristics, estimated mortality rates and originated a detailed account of char landings on an area-to-area, stock-by-stock basis. In a continuation of these latter studies, Dempson and Best (1978) analyzed results of the 1976 Arctic char fishery, and by utilizing data from Coady and Best (1976) verified over-exploitation by using a combined assessment of various stocks in northern Labrador.

I /

The purpose of this paper is to present a more complete assessment of the status of northern Labrador Arctic char stocks utilizing the Beverton and Holt yield per recruit model. It is based on more substantial data accumulated during 1977 from six primary char fishing areas. The results of the commercial char and Atlantic salmon fishery, which ranged from Anton1s in the south to Hebron in the north (Fig. 1), are also presented.

MATERIALS AND METHODS

COMMERCIAL LANDINGS

Information on the commercial landings of char and salmon were obtained from the Economics and Intelligence Branch (E &I) of Fisheries and Marine Service. Purchase slips, prepared by E & I, were issued to buyers and were filled out at the time of catch receipt. Information requested consisted of the name of the fisherman, license number, area where fish were caught (Fig. 2), date, number of nets used, species and the number of pounds of fish in each size category. In addition, numbers of fish caught were added to catch records during 1977.

2

."1

r::'·~· LABRADOR ,fI,

\.'1 i " l.". / (~._._._._._,_._._"_._._"_._"_

-\....,.

580

ATLANTIC OCEAN

80 0 80 160 I!!hEl!!--!E3.....!!!!!I;;;=~~! ===:===:==1'

SCALE (Kill

..... ;,',

/..... .i \

/" '. ~ ,..

QUEBEC

, /', " .J \'

'.,J , r

! \.

Fig. 1. Extent of the major Arctic char fisheries in Labrador during 1977.

3

56° 56°

~ -x­

~

THE DOMES

CUTTHROAT

HEBRON

NAPARTOK

TIKKOATOKAK

;=*MSERRIVER -

NAIN DOG ISLAND ANAKTALIK

~

25 0 25 h ...... ---! !

SCALE (KMl

63° 62° 61°

Fig. 2. Coastal breakdown of Nain commercial fishing areas.

~

4

Analysis of these data provided weekly breakdowns of landings of char and salmon by size category and by area. Weights listed are in kilograms and have been converted to round condition by appropriate conversion factors: Arctic char, gutted, head-on to round fresh weight, x 1.24; Atlantic salmon, gutted, head-off to round fresh weight, x 1.25. Calculated mean weights were used in estimating total numbers of fish caught by area and by week. Catch per unit effort estimates were derived following the method initiated by Coady and Best (1976) in terms of man-weeks and net-weeks fished.

and are expressed

they Landings of salted char were identifiable as to were omitted from the catch and effort analysis.

area only. Thus,

COMMERCIAL SAMPLING

The Nain fish plant served as the centre for all sampling activities during 1977. Char were off-loaded at the plant by fishermen operating in adjacent areas or by collection vessels which gathered char from distant areas along the Labrador coast. Individual catches, identifiable as to origin, were sorted on the basis of size category and quality by plant personnel. Sampling data used in the assessment were then randomly chosen in the proper portion from respective size categories. Sampling from Voisey, Anaktalik, Tikkoatokak, Okak, Napartok and Hebron areas continued throughout the summer in order to ensure adequate data were collected from each area for the duration of the fishery.

In addition to recording fork length (cm) and gutted head-on weight (kg), both otoliths (sagitta) were extracted for subsequent age determination according to the method of Nordeng (1961). Length and age composition were determined for stocks representing the six areas sampled. Comparisons were made with Voisey, Tikkoatokak and Okak areas from available 1974 data. Percentage length frequencies were plotted using 2-cm intervals, and percentage age compositions were represented in the form of histograms.

MORTALITY

Estimates of total mortality (Z) were calculated from least squares regressions fitted to the descending limb of catch curves, based on the number caught in each fully recruited age class. Normally age compositions are combined for successive years in order to eliminate fluctuations resulting from variable recruitment of fish to the exploited phase of the population (Beverton and Holt 1956~ Ricker 1975). Owing to lack of consecutive sampling data, this was not possible for the various areas with the exception of Hebron, where 1976 and 1977 sampling data were combined.

Comparisons of variation in mortality rates over a 3-yr period were feasible using sampling data from Coady and Best (1976). In particular, variations noted in older age compositions were examined with respect to significant changes in total mortality coefficients.

A second estimate for total mortality was calculated by applying information obtained in 1974 with that from 1977. This followed the method of Paloheimo (1961) as outlined by Ricker (1975) where age composition, catch and estimates of total effort were known. Rates of exploitation

5

(u) were calculated applying both estimates of the instantaneous rate of total mortality (Z).

In a prior assessment for the combined fishery, a value of 0.20 was assumed and used as (M), the natural mortality coefficient (Dempson and Best 1978). However, similar to Jensen and Berg (1977), natural mortality rate for this assessment was assumed at 0.17 based on an estimate by Moore (1975) for an unexploited anadromous char population in the Cumberland Sound area of Baffin Island. This is also in close agreement to the value of 0.153 derived by Hunter (1976) for Arctic char in the Sylvia Grinnell River, Frobisher Bay.

GROWTH

Growth curves were fitted to age and length data from Voisey Bay, Anaktalik, Tikkoatokak, Okak, Napartok and Hebron areas by the least squares method of Allen (1966) according to the von Bertalanffy (1938) equation:

lt = (l_e-K(t-to))Loo where lt length of fish (cm) at age t (years)

Loo = theoretical maximum length K constant determining rate of change in length increments

to hypothetical age at which lt = 0

A semi-logarithmic plot of dl against t (Sandeman 1969) of the dt

equation dl = K Loo e-K(t-to) dt

was used in comparing rate of instantaneous growth with change in age. This was accomplished by graphically collating growth data for various areas sampled during 1977.

Least squares regression analyses were calculated on logarithmic transformed length-weight data from all areas according to the form:

log w = log a + b (log 1), where w = gutted, head-on weight (kg) and

1 = fork length (cm)

Regressions were compared by analysis of covariance, and were employed to convert theoretical maximum length (Loo) to theoretical maximum weight (Woo) for the yield per recruit assessment .

•

YIELD PER RECRUIT

The Beverton and Holt (1957) yield per recruit model was applied to assess exploited Labrador Arctic char stocks following a method similar to that of Bowering (1977) and Bowering and Pitt (1978) for witch flounder, Clark et al. (1977) for pollock and Borrmann (1977) for roundnose grenadier. Von Bertalanffy growth equations supplied several parameters for the assessment. I~ean selection length, used in determining mean selection age (T ,) was determined from ascending left limbs of length frequency curvesPfollowing the method of Hodder (1964). By rearranging the von

6

Bertalanffy growth equation, mean selection length was then converted to mean age at first capture. Levels of fishing mortality (F) up to 2.5 were used in computing yield per recruit values.

Optimum fishing levels, FO. l (ICNAF Redbook 1972) as described by

Gulland and Boerema (1973) were calculated and compared to catches corresponding to fishing at the level of F and present levels of max fishing mortality as calculated for 1977.

RESULTS AND DISCUSSION

COMMERCIAL LANDINGS

Size b~eakdown and weekly landings of cha~ and salmon

The size breakdown of the 1977 Arctic char landings is outlined in Fig. 3 along with that of the previous 7 yr for comparison. Approximately 75% of total coastal landings were under 2.3 kg (gutted head-on weight); however, a decreasing trend is apparent in landings of this size category since 1974. Comparisons between areas revealed considerable variation. Char landings under 2.3 kg ranged from 91% in Dog Island, 85% in Nain Bay, 80% in Tikkoatokak, to only 62% for Anaktalik.

Atlantic salmon landings were similarly broken down into size classes. During 1977, 78% of landings in northern Labrador exceeded 2.72 kg (gutted head-off weight). For 1976, 82% of salmon landed at Nain were over 2.72 kg, whereas large salmon in southern Labrador represented 71-78% of the total catch (Reddin 1978).

The close relationship between the char and salmon fisheries, elucidated by Coady (1974) and Coady and Best (1976), was again evident in 1977 (Table 1, Fig. 4). In comparison with previous years, fishing effort during the initial part of the season was devoted entirely to char and was concentrated in areas adjacent to and immediately south of Nain (Fig. 5). With the improvement in coastal ice conditions and subsequent appearance of migrating salmon along the outer islands, effort was diverted north to other char and salmon berths along the coast. Salmon landings increased substantially during week 4 and peaked in week 5. However, the availability of salmon diminished rapidly as evidenced by the decline in catch per unit effort (Table 1). During 1976, both catch and catch per effort for salmon remained high throughout weeks 5-8, with char declining in both categories. Conversely, in 1977 attention again focused on Arctic char during this period as it became apparent that salmon were not as abundant as in previous years.

Landings of A~ctic cha~ and Atlantic salmon by a~ea

The Labrador coast north of Hamilton Inlet is divided into two major fishing areas (Fig. 1). The Nain fishery handles char and salmon caught from Anton's Point and northward (Fig. 2), while the Makkovik fishery encompasses all areas from Anton's Point southward to the area of Rigolet. Landings of Arctic char at Nain totalled 186,201 kg

7

85 rr---r---r-----,-------,----r---r--~-~

80 >­a: ot3 70 ~ ~ u w60 N

(/)

~ 50 ~

~40 u a: w Q..

>- 30 co (/)

~20 Ci z ::; 10

1 ,I,I,

/-,-----__ 1 ,

1 " UNDER 2·3 KG. 1 " I , I

I 'I I

'72 '73 '74 '75 '76 '77 YEAR

Fig. 3. Relative changes in the size composition of char landings at Nain, 1970-77.

30

.. d ~20 Cl z: ::::l o a: 15 I

(/) c> z Ci 10 z ~ .....J

5

00

~CHAR

I,,1 , , 1 ' 1 " 1 ,

," SALMON '\,,'" 1 '

.... " 2 3 4 5 6 7 8 9 to II 12

WEEK

Fig. 4. Summary by week of Nain char and salmon landings.

8

Table 1. Weekly landings of char and salmon at the Nain fish plant, 1977.

Fishing week

Landings (kg round)

Char Salmon

Man-weeks fished

Char Salmon

Catch/man-week (kg round

Char Salmon

1 2 3 4 5 6 7 8 9

10 11 12

June 26-July 2 July 3-9 July 10-16 July 17-23 July 24-30 July 31-Aug. 6 Aug. 7-13 Aug. 14-20 Aug. 21-27 Aug.28-Sept. 3 Sept. 4-10 Sept. 11-17

9,611 29,835 23,235 15,630 14,373 25,006 18,975 9,260

10,554 5,799 2,335

477

4 35

1,489 12,515 9,201 6,817 3,236 3,794 2,149 1,548

830

39 66 83 59 79

122 118

75 68 39 23 3

1 8

36 74

108 104

70 65 47 35 17

246 452 280 265 182 205 161 123 155 149 102 159

4 4

41 169

85 66 46 58 46 44 49

Summary 165,090 41,618 774 565 213 74

9

(round fresh weight) and represented 88.5% of total char caught along the coast. The remainder, 24,205 kg, were landed at Makkovik. Total Labrador catch for 1977 exceeded landings in 1976 by approximately 40% and contributed in excess of 50% of total Canadian production (R. Peet, B. Wong, pers. comm.).

Approximately 75% of the char caught in northern Labrador originated from the Voisey Bay, Anaktalik, Tikkoatokak, Okak and Napartok areas. For 1977, most areas showed substantial increases when compared to the average landings for the period 1974-76 (Fig. 6). For the third consecutive year the Tikkoatokak area recorded the highest catch. Upwards of 100,000 kg have been removed from this bay since 1975. This is one area in particular in which the char stocks are in a rather precarious state. Evidence from this study indicates that these stocks are being severely overfished and thus the potential exists for the fishery in this area to collapse.

Atlantic salmon landed at Nain totalled 41,618 kg (round fresh weight), a decrease of 41% from 1976. Preliminary results for salmon caught in the Makkovik fishery indicate an increase of 12% to 82,796 kg.

The geographical separation between char and salmon berths was again evident during 1977. The majority of salmon landings originated along the outer bays and islands. Dog Island, Black Island, Kiglapaits and the Cutthroat areas produced 76% of the salmon landed at Nain, with the latter area contributing in excess of 40% of the total. Previous records indicate the importance of these areas with respect to the Nain salmon fishery (Coady and Best 1976; Dernpson and Best 1978). With the exception of Cutthroat, char landings are generally low in these regions.

Char and salmon landings are summarized by area in Tables 2 and 4 respectively. Appendices 1 and 2 provide a more detailed account of the landings by area, size category, and by week. For comparative purposes, char data from previous years were converted into a standard form (kg-round) and are summarized in Appendix 3. Table 3 lists final char landings by area including those salted landings which were identifiable as to origin only. Not included, however, are 186 kg unaccounted for by area. ;[

Arctic char are generally considered to be a localized species remaining in immediate esyuaries and bays, close to their home rivers (Yessipov 1935; Scott aud' Crossman 1973). Several studies have supported this assumption. Nieli9h (1961), in determining the migrations and homing instinct of char from Greenland, found that the maximum distance travelled by a tagged individual was 62 km; however, 86.4% were recaptured less than 30 km from the locality in which they were tagged. Movements of char tagged in the George River, Quebec, were within 40 km of their release site (Le Jeune 1967), while in Norway, Jensen and Berg (1977) reported most tag recaptures within 25 km from the initial tagging location; however, a few anadromous char did travel much farther. In the Cumberland Sound area of Baffin Island, average maximum distances travelled by anadromous char were 40-50 km (Moore 1975). Studies in northern Labrador indicate that of char tagged in the Fraser River, northwest of Nain, 86% of the recaptures have been from either Nain Bay or from fish returning to the Fraser River. The remaining recaptures

10

were caught in adjacent Tikkoatokak Bay (Fig. 2) but within 40-50 km from the mouth of the Fraser River. In southern Labrador, char were tagged at the counting fence facility on Northwest Tributary of Sand Hill River. Recaptures of 96% from nearby Sand Hill Cove and remaining recaptures within 25 km of Sand Hill River confirm the localized nature of Arctic char movements.

In view of the relatively slow growth and maturation of the char coupled with its localized nature, it is, therefore, obvious why these stocks have to be carefully managed. A population more or less geographically isolated cannot withstand long periods of intense exploitation (Bigelow 1963). A commercial fishery should be diffuse in order to avoid damaging stocks from a concentration of effort in specific localities (Dunbar and Hildebrand 1952). As Dunbar (1970) points out, the Danes in Greenland have long realized that, in order to conserve commercially fished Arctic char stocks, each area must be carefully monitored. The idea of rotational fishing, as traditionally carried out in the far north (Hunter 1976; Johnson 1976) may have merit if it could be adapted to fit in with the present large commercial char fishing operation in Labrador. The collapse of the char stocks at Nachvak, Labrador (Andrews and Lear 1956), Frobisher Bay (Hunter 1976), Tree River, Northwest Territories (Robertson and Dowler 1973) and the documentation of overfishing in several rivers in northern Quebec, in particular the Georges River (Le Jeune 1968), should serve as ample reminder of the vulnerability of Arctic char and what can eas~ly

happen if catches are unregulated.

II

Table 2. Summary by area of Nain char landings, 1977.

Total catch Man-weeks Catch per man-week Net weeks

Catch per net-week

(kg round) fished (kg) fished (kg)

Anton's Point 2,010 19 106 32 63 Voisey Bay Anaktalik

22,490 21,604

58 74

388 292

112 138

201 157

Dog Island Nain Bay Tikkoatokak

2,038 8,463

39,489

51 31 97

40 273 407

108 36

177

19 235 223

Webb Bay Black Island

2,516 3,391

20 60

126 57

36 191

70 18

Kiglapaits Okak

4,320 19,099

43 70

100 273

89 126

49 152

Cutthroat 13,215 111 119 210 63 Mugford Napartok Hebron

1,376 19,124 5,955

9 93 38

153 206 157

12 165 66

115 116 90

SUlTiTlary 165,090 774 213 1,498 110

•

12

Table 3. Total Arctic char landings for 1977, including pickled production.

Area Kg round Kg round-salted Total

Anton's Point Voisey Bay Anaktalik

2,010.322,490.2 21,604.4

101. 7 2,112.0 22,490.2 21,604.4

Dog Island Nain Bay Tikkoatokak

2,037.7 8,462.6

39,489.4

2,037.7 8,462.6

39,489.4 Webb Bay Black Island

2,516.3 3,390.5

2,516.3 3,390.5

Kiglapaits Okak

4,319.8 19,099.1

1,119.4 8,499.0

5,439.2 27,598.1

Cutthroat 13,215.5 2,282.3 15,497.8 Mugford Napartok Hebron

1,375.5 19,123.7 5,594.9

8,923.2 1,375.5

28,046.9 5,594.9

Tota1 Kgs (round) 165,089.9 20,925.6 186,015.5

13

Table 4. Area breakdown of Nain salmon landings, 1977.

Total catch Man-weeks Catch per man-week Net weeks

Catch per net-week

Area (kg round) fished (kg) fished (kg)

Anton's Point 126 10 13 17 7 Voisey Bay Anaktalik

83 27

13 5

6 5

25 11

3 2

Dog Island Nain Bay Tikkoatokak

6,000

22

81

5

74

4

166

14

36

2 Webb Bay Black Island

236 8,662

17 93

14 93

31 271

8 32

Kiglapaits 5,358 44 122 90 60 Okak 842 51 17 92 9 Cutthroat 17,657 136 130 254 70 Mugford Napartok Hebron

2,175 430

79 31

28 14

150 53

15 8

Surrmary 41,618 565 74 1,174 35

14

10 12

WEEK

4

NAPARTOK

KIGLAPAITS

... 010

1 ~ P'II /, I2 4 6 8 10 12

10 BLACK ISLAND

~ ~ .&. 0,7 0-03

~

50

30

10

YO

10

MUGFORD

g t Jh---,I1""--'1Ir-T""OI1 - a 2 4 6 8 10 12 >< ~50 CUTTHROAT ¥ -49:;; 40 '" ~30 C> z:5 20

10

af-r--.--l'''''''' a

,-----\ ~:e.:u >< 2 4 6 8 10 12

SCALE (KM)

25 0 HnM

~""""'\?rL~~~~:__~_-"""~~~ 2 4 6 8 10 I

WEEK

8 10 12

VOISEY BAY

FRASER TESSISOAK LAKE

:?:~N:;~R-­20

10

a 2 4 6 8 10 12

14

10

6

4

WEE

60

40

20

o 2 4 6 8 10 12

-140 o ~IOO x ~ 60 >< - 20 '" '" z o z ""-'

Fig. 5. Arctic char landings summarized by area and by week.

570

15

~ cD I' ­a't QllO~~nl'l

I I' ­~ I' ­

a'tI' ­~. 1VOllH.unJ

<[

w a:: <[

0 LO 0 LO 0 LO 0 LO LO 0 LO ~ ~ I""l NI""l N

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16

Total fishing effort directed towards Arctic char in 1977 rose to 774 man-weeks fished, an increase of 52% over the previous year. However, the level of success for the 1977 fishery, as indicated by the catch per unit of effort, diminished by approximately 20% in comparison with 1976. This decline appeared to be a direct result of increased fishing. A succession of weak year-classes may also contribute to a decreased abundance; however, the reduced mean age of char in Voisey and Tikkoatokak bays is characteristic of the former (Gulland 1968).

Despite the expansion of the commercial fishery into the Napartok and Hebron areas in 1976 and 1977 respectively, effort has continued to increase in the immediate vicinity of Nain. This increase amounted to 44% in Voisey, Anaktalik, Nain and Tikkoatokak bays. Presently, there are few restrictions governing the northern char fishery in particular with respect to the number of entrants into the fishery each year. Prior to 1970, numbers of fishermen fluctuated annually with 66 fishing in 1962 (Coady and Best 1976) and 19 in 1966 (Dyke 1967). In 1976 there were approximately 100 commercial licenses granted while the figure for 1977 approached 130. What is definitely needed is a more even distribution of fishing effort if the present rate of fishing continues to increase in the absence of regulatory measures being imposed to offset localized overfishing in the Nain area, and certainly continued monitoring of catch and effort is imperative.

With respect to the Atlantic salmon fishery at Nain, the catch per unit of effort and consequently the availability of the species (Gulland 1964) was 53% lower in 1977 than in the previous year. Effort, however, rose by 26%. Dog Island, Black Island, Kiglapaits, and Cutthroat continued to produce the majority of salmon catches with fishing effort similarly concentrated in these areas (Table 4).

In trying to identify reasons for the decline in total catch per unit of effort, fishing success was examined with respect to char and salmon fishing berths (Table 5). In comparison with 1976, the abundance of char decreased slightly in predominant char areas but increased in salmon berths. Availability of salmon, however, declined in both char and salmon fishing regions.

Table 5. Catch per unit effort (kg per man-week) in relation to primary char and salmon fishing areas.

Char berths Salmon berths char salmon char salmon

1976 298 60 70 168 1977 281 19 89 106

17

The migratory pattern of Atlantic salmon differs from that of the Arctic char. Water temperature and other related environmental conditions, for example ice, may greatly influence salmon movements. It appears that heavy ice along the northern Labrador coast in 1977 altered salmon distribution and thus was at least partially responsible for the decline in catch per unit of effort. Undoubtedly effort directed towards Arctic char would be substantially augmented should the salmon fishery fail which is not inconceivable according to Coady and Best (1976), considering they were virtually absent in the Nain area prior to 1969.

COMMERCIAL SAMPLING

Age and length composition

A summary of numbers at age, mean length and weight, with range and standard error is provided in Appendix IV. As a result of numbers of fish caught being recorded on the majority of catch records, average weights were determined and used in calculating total numbers of char caught by area. When this information was combined with age composition data, total numbers at age were estimated for areas sampled during 1977 (Appendix V).

Comparison of age and length distributions indicate some variation has occurred in the composition of commercial catches since 1974. Younger fish are contributing more to the fishery in areas contiguous to Nain (Fig. 7). In Tikkoatokak Bay, 7- and 8-yr-old fish made up the majority of catch during 1977 (mean age, 8.1 yr), whereas 8-, 9- and 10­yr-old char were the most significant contributors in 1974 (mean age, 9.4 yr). The shift to younger ages was less noticeable in Voisey Bay with 7-, 8- and 9-yr-old char representing the greatest proportion (mean age, 8.6 yr), as opposed to 8-, 9- and 10-yr-olds in 1974 (mean age, 9.4 yr). Apart from the predominance of 12-yr-old Arctic char in the Okak samp~es, the majority of the catch was still composed of 10-14-yr-olds. Mean age for 1977 was 12.0 yr compared with 11.7 yr for Okak Bay in 1974. Reduced effort in the Okak area since 1974 is a contributing factor.

The progressive increase in fishing effort directed towards Tikkoatokak Bay stocks since 1974 (Appendix 3, Table 2) has resulted in diminishing the percentage of Arctic char over 10 yr of age as determined from commercial sampling. The changes in percentage composition of char in older age classes for Voisey, Tikkoatokak and Okak bays were as follows:

age(yr) 1974 1977

Voisey Bay over 10 39.4 23.0 Ti kkoatokak Bay over 10 42.7 14.3 Okak Bay over 14 17.6 21. 2

With respect to remaining areas, Anaktalik had a mean age of 9.3 yr, with 8-, 9- and lO-yr-old char contributing 72% of the catch Fig. 8). The oldest char, 21 yr, originated from Napartok Bay. Mean age for this area was 12.3 yr, with 12-yr-old char in excess of 30% of the landings. The lower mean age of char at Hebron (10.6 yr) may be partially explained in relation to the date of first landings from the area. These were during week 6 (July 31-August 6). Arctic char in northern fiords are reported to have an earlier upstream migration

18

30 VOISEY BAY

1974

N: 170

VOISEY BAY 1977

N: 313

30

20

10 t ­Z

~ 0a::: w Q..

30

20

10

o

30

TIKKOATOKAK 1974

TIKKOATOKAK 1977

N: 441

OKAK 1974

30

20

10

O~~~L.J

OKAK 1977

N: IS6

6 7 8 9 10 " 12 13 14 IS 16 AGE (YEARS)

17 18 19 20 21 29 39 49 59 69 LENGTH FREQUENCY

12 eM INTERVALS)

79

Fig. 7. Comparison of age and length compositions Tikkoatokak and Okak bays.

of Arctic char from Voisey,

19

ANAKTALIK 1977

N 201

NAPARTOK 30 1977

to-z N 285~20 a:: I.LJ

Q.. 10

0

HEBRON 30 1976-77

N 23020

6 7 8 9 10 II 12 13 14 15 16 17 18 19 20 21 29 39 49 59 69 79 AGE (YEARS) LENGTH FREOUENCY

12CM INTERVALS)

Fig. 8. Age and length composition of char from Anaktalik, Napartok and Hebron areas.

20

than those in more southern bays (Brice-Bennett 1977), and continuing studies on the Fraser River, Labrador, have suggested that the average size of returning migrations decreased as the run progressed. Hence it is possible that larger char from Hebron were not truly represented in the landings and thus effected a reduced mean age for the population.

Length composition generally followed closely with that of age (Fig. 7 and 8). There was a pronounced shift to smaller char in Tikkoatokak Bay since 1974, however, there has been a significant increase in the mean length of char from the Okak area during the same period (P = 0.01).

Mortality

Estimates of survival rates and total mortality (Z), as derived from catch curves, are usually those prevalent at the time when fish are recruited into the fishery and not of those rates occurring during the year of capture (Ricker 1958). This is important to consider when comparisons are made between years. Instantaneous total mortality, as derived from catch curves, are presented in Fig. 9 and 10 for all areas commercially sampled.

The decline in older age classes in Tikkoatokak Bay was reflected in the increased total mortality rate over the period 1974-77 (Table 6). The Okak area, where effort has been lower in contrast with 1974, had an increase in the proportion of older char and this was similarly reflected in a decreased total mortality. Total annual mortalities ranged from 31.6% in Hebron to 57.3% in the Tikkoatokak area (Fig. 9 and 10) .

Table 6. Comparison of instantaneous total mortality rates (Z).

Ricker catch curve mortality

1974 1977 Paloheimo mortality

Voisey Anaktalik

0.74 0.59 0.80

0.70

Tikkoatokak 0.70 0.85 1.03 Okak 0.51 0.44 0.38 Napartok Hebron

0.47 0.38

0.45

• • • • • • •

I -l--r

1974

A:52·3% AGES 9-13

•

1974

A: 50'3%

AGES 9-14

A: 40·0% AGES 11-16

• • •

21

Fig. 9. Catch curves of age distribution of Arctic char taken by gil1nets in Voisey, Tikkoatokak and Okak bays.

6

5

4

3

2

0

5

4

z ... 3

c.=> 0 -oJ

2

0

5

4

3

2

0

-~ ­I

VOISEY BAY VOISEY BAY 1977

\ Z: 0·59 A:44·6%Z=0·74 • r : 0·99 AGES 8 -14 • r : 0·99 "'. .,

\ '\

•

\.• \•

• TIKKOATOKAK 1977T1KKOATOKAK \ • Z : 0·85 A: 57'3%Z : 0·70 \ •• r : 0·99 AGES 8-14r : 0·98 • \ •

•

•

-I

OKAK 1977OKAK 1974

Z : 0·44 A: 35'6% Z: 0·51 r: 0·95 AGES 12-16

•• .""• • ~7•

~ • •

• •

12 14 16 18 20

22

4-­

31­

2-­

II- ­

0

•

•

• •

•

.\

ANAKTALIK 1977

z: 0-80 A: 55·1 % r :0-99 AGES 10 -14

-

-

-

-

z

51­

41­

31-­

o

2

4

I-­

II­

I­

•

•

•

NAPARTOK 1977 -

• • ~ Z:0-47 A: 37·5 % -

• r: 0·99 AGES 12-15 • "­• -

-

." ~.

-

"'. - .­-

.~ HEBRON 1976 -1977

2

3 -

l­ •

• •"'­ • z: 0-38

r : 0-95

A: 31-6%

AGES 10-14

-

_

- . • -

o • • 6 8 10 12 14 16 18 20 22

AGE (YEARS)

Fig. 10. Catch curves for Arctic char in gillnet catches from Anaktalik, Napartok and Hebron areas.

23

A comparison of Paloheimo and Ricker catch curve mortalities has been provided (Table 6). The only substantial difference noted was for the Tikkoatokak area where total mortality rate was estimated at 1.03. Total annual mortalities, as calculated ·from the Paloheimo method, were 50.3% for Voisey Bay, 64.3% for Tikkoatokak, 31.6% and 36.2% for Okak and Napartok regions respectively.

Lowest rates of exploitation were 0.17 at Hebron and 0.22 of Okak. Tikkoatokak had the highest rate at 0.46. Estimates for remaining areas were 0.32 at Voisey Bay, 0.44 at Anaktalik and 0.24 for Napartok. Paloheimo mortalities generally yielded higher exploitation rates with the severest rate occurring in Tikkoatokak Bay at 0.54. Other values were 0.38, 0.17,0.23 for Voisey Bay, Okak and Napartok respectively.

GY'Owth

The growth pattern for Arctic char from Voisey, Anaktalik, Tikkoatokak, Napartok and Hebron was described by the von Bertalanffy growth equation (Allen 1966). K-values ranged between 0.14 and 0.34, the latter for Napartok char (Table 7). Loo- values ranged between 63.3 and 73.6 em, and generally agreed with maximum empirical values for respective areas with the exception of char from Napartok Bay. Von Bertalanffy growth convergence was not obtained for Okak Bay data. Plotted values, as determined from the von Bertalanffy equation, corresponded closely with empirical data of commercial samples (Fig. 11). A less accurate curve below fitted values may be explained by the size selectivity of fishing gear in which smaller fish are often disproportionately represented (Grainger 1953; Ricker 1975).

24

Table 7. von Bertalanffy growth parameters with standard error and 95% confidence limits for Arctic char from 5 Labrador areas, 1977.

Variable Fi na 1

estimate Standard error

Confidence limits Lower limit Upper 1imit

Vo;sey (N = 313)

K to Loo

0.18 0.18

70.92

0.07 1.60 5.79

0.05 -2.95 59.58

0.32 3.31

82.27

Anaktalik (N = 209)

K to Loo

0.16 -0.24 73.55

0.12 3.27

10.91

-0.06 -6.65 52.17

0.39 6.18

94.94

Tikkoatokak (N = 441)

K to Loo

0.18 1.20

71.84

0.06 1.02 7.10

0.00 -0.81 57.91

0.30 3.21

85.77

Napartok (N = 285)

K to Loo

0.34 3.14

63.25

0.16 2.36 1. 76

0.04 -1.49 59.80

0.65 7.78

66.69

Hebron (N = 230)

K to Loo

0.14 -0.04 66.46

0.08 2.65 9.30

-0.02 -5.24 48.23

0.31 5.16

84.68

- -

25

I I I I I I

80

0_...:-r-­..... .."..-.-.-...... 0

060

(~ • 0

.A"."

:/.'"

40 VOISEY 1977 NAPARTOK 1977

20

-

o I I I I , I I I I I

80

•.>-~ .-:----~i"60

/'~ .... .......... ..... 0u

•",...,. .'" ­

ANAKTALIK 1977 TIKKOATOKAK 1977 _

20 / -

I Io

80 I­

____0­60

~.-. • 0 ........ . 0

40 /.~. HEBRON 1977

20

2 4 6 8 10 12 14 16 18 20 22 2 4 6 8 10 12 14 16 18 20 AGE (YEARS)

Fig. 11. Growth curves of Arctic char from areas sampled in 1977. Open circles represent samples of less than 5.

26

The high K-value for Napartok char indicated that the rate of change in length increments (Ricker 1975) and therefore the rate at which L~ was approached was comparatively fast (Fig. 12). This appears reasonable as the theoretical maximum length calculated for Napartok char (Loo = 63.3 cm) was substantially lower in comparison with other areas. Previous studies, however, suggest a decrease in the value of K with an increase in latitude (Coady and Best 1976) implying that the rate of approach to L~ is slower in more northerly areas. An improved sample size may have yielded a more realistically higher value for L~

in conjunction with a lower Brody growth coefficient.

Growth comparisons between male and female Arctic char were not possible with 1977 data. Coady and Best (1976), however, reported no significant difference in growth rate between sexes in char from Nain Bay. Males grew to a large size but their rate of approach to L~ was slower than for female char. The latter was similarly found in char from northern Norway by Jensen and Berg (1977) who employed a different approach in determining von Bertalanffy growth parameters.

Results of the length-weight regression analyses have been provided (Table 8). Covariance analyses indicated that slopes (Fl, 1541 = 11.44) and intercepts (Fl, 1546 = 86.21) were significantly different among areas at less than the 1% level.

Table 8. Length-weight (gutted head-on) regression coefficients, from log W= log a + b (log 1), for Arctic char sampled from various coastal Labrador areas.

Area No. of fish log a b S.D. of b

Voisey 322 -4.90 2.96 0.14 Anaktalik 213 -4.68 2.83 0.13 Tikkoatokak 458 -5.25 3.15 0.19 Okak 162 -4.04 2.47 0.11 Napartok 306 -4.89 2.92 0.13 Hebron 238 -4.38 2.64 0.17

Yield per recruit

Parameters used in the Beverton and Holt yield per recruit analyses have been provided (Table 9).

The yield curve for Napartok Bay was essentially flat-topped indicating no definitive F up to F = 2.5 (Fig. 13). Landings of Arctic char for max 1977 in relation to catches corresponding to F and FO. l are summarizedmax in Table 10. Present rates of fishing mortality exceeded optimum levels in Voisey and Anaktalik bays, while surpassing the maximum value in

----

27

20·0 ,--r---r--.---r----r--,---.----.---,---,--,---,

10·0

8·0

6·0

4·0-~I~

::I: ..... ~ 2·0 0:: t.:)

V>

0 ~

l.LI

~ (-0 ..... ~ 0·8 ..... V>

~0'6

0·4

0·2

3

2 I 4 5

I. VOISEY 2. T1KKOATOKAK 3. NAPARTOK 4. ANAKTALIK 5. HEBRON

0·1 '---~-~--7----::---:-----l=---l--...l.--..l.---L--L--.J 12 13 14 15

Fig. 12. Growth rate plotted against age for char from five Labrador areas.

Tikkoatokak Bay (Fig. 13). Based upon the calculated values, overfishing was as high as 86% in Tikkoatokak Bay, while only 12% in the Voisey area.

Analysis of the 1976 char fishery revealed the need for a 36% reduction in total landings (Dempson and Best 1978). Results of the 1977 analyses suggest a 33% reduction in landings from Voisey, Anaktalik and Tikkoatokak.This was determined by comparing actual landings from these areas with that indicated to be the optimal catch obtained by fishing at the FO. l rate of fishing mortality. The assessment also

implies that Arctic char stocks in the northern areas of Napartok and Hebron are presently underexploited and consequently effort could be increased in these regions.

Table 9. Parameters used in the yield per recruit assessment.

Areas Parameter Vo;sey Anaktalik Ti kkoatokak Napartok Hebron

w~ - asymptotic weight (kg) 4.76 5.13 4.79 2.89 3.38 K - growth coefficient to - theoretical age for It=O tp - age at recruitment t p ,­ mean selection age

age at last significantt -contribution to fishery

0.1810 0.1815 6.0 6.42

14

0.1653 -0.2372 6.0 6.70

14

0.1774 1.1964 6.0 6.09

14

0.3436 3.1433 8.0 8.51

19

0.1433 -0.0432 6.0 7.20

17

Table 10. Landings of Arctic char for 1977 with respect to catches corresponding to Fmax and FO' I ' Bracketed figures based on Paloheimo mortality rates.

Vo;sey Anaktalik Tikkoatokak Napartok Hebron

Present F 0.42 (0.53)

0.63 0.68 (0.86)

0.30 (0.28)

0.21

Landings (kg) 22,490 21,604 39,489 28,047 5,955

FO' 1 0.37 0.39 0.31 0.39 0.31

Catch at FO.I(kg) 20,061 (16,870)

14,767 21,210 (18,105)

35,384 (37,120)

8,495

F max 1.06 1.38 0.58 1.14

Catch at Fmax(kg) 42,790 (35,984)

34,683 35,151 (30,005)

21,925

0-4

1-4

29

1·6

/x -Fmal'·4 x-­ -FIlIGl-" -FO'IFo·,1·2

1·0

0·8

0·6

VOISEY BAY ANAKTALIK0·4

0·2

0·0

~=1-2

~ )·0 _---x::> a:: FO'I Fmal

t:lO-B a:: -,...- -Fmal ffiO·6 (001a..

TIKKOATOKAK HEBRON=: 0·4 w

>= 0·2

0·0

1·4 ~ "-Fx 0·1

O-B

NAPARTOK

0·2

0-0 ~;:-'-;;----;::L:--::"'-:-::-'-::-----:-'-~:L=---:-L:-----:-'-~:L=-~~::...L..----:--I.,,----,~-L.----l_...L--L.----l.-...l.-----L----l_..LJ 0·2 0·4 0·6 0·8 1·0 1·2 '·4 1·6 '·8 2·0 2·2 0·2 0·4 0·6 0-8 1·0 1·2 '·4 1·6 1·8 2·0 2·2

FISHING MORTALITY

Fig. 13. Yield per recruit curves indicating present rate of fishing mortality (X). FO. l and F fat' char from northern Labrador areas. max

30

SUMMARY

1. Labrador landings of Arctic char totalled 210,406 kg in 1977 and was one of the highest catches on record.

2. Average size of char landed at Nain has increased slightly since 1974; however, approximately 75% of the landings are still under 2.3 kg (gutted head-on weight).

3. Approximately 75% of Arctic char caught in northern Labrador originated from the five areas, Voisey Bay, Anaktalik, Tikkoatokak, Okak and Napa rtok.

4. Tikkoatokak Bay recorded the highest catch for the third consecutive year, with in excess of 100,000 kg being removed since 1975.

5. Total effort rose by 52% in comparison with 1976; however, the catch per unit of effort decreased by 20%.

6. Majority of the effort is still concentrated in the immediate Nain area despite expansion of the commercial fishery into the Napartok and Hebron areas in 1976 and 1977 respectively.

7. Salmon landings at Nain diminished by 41% to 4·1,618 kg.

8. Catch per unit effort for salmon was 53% lower than in 1976. The decreased availability appears related to environmental conditions altering salmon migrations.

9. A total of 1690 Arctic char from six locations were sampled for age, length and weight composition.

10. Comparisons with 1974 data indicated a reduction in the mean age of commercial catches from the Nain region. Mean age for Tikkoatokak Bay char dropped 1.3 yr to 8.1. Napartok Bay had the highest mean age at 12.3 yr.

11. Total annual mortalities ranged from 31.6% at Hebron to 57.3% for the Tikkoatokak area. Similarly, exploitation rates varied from 0.17 to 0.46 for Hebron and Tikkoatokak respectively.

12. Age and length data were employed in fitting von Bertalanffy growth curves.

13. Significant differences were found in length and weight characteristics of char from the various areas sampled.

14. Beverton and Holt yield per recruit model was used to assess the status of northern Labrador char stocks from parameters derived from the von Bertalanffy growth equation.

15. Optimal management of the fishery was assumed at the FO. l level of fishing mortality. Present levels of fishing mortality exceeded optimal levels in Voisey and Anaktalik bays, while surpassing the maximum value in Tikkoatokak Bay where overfishing was as high as 86%.

31

16. Tagging studies in Labrador supported the assumption on the localized nature of Arctic char. It is essential that exploited stocks be carefully assessed and managed because of its slow growth and maturation. Tikkoatokak Bay stocks in particular appear to be severely overfished and continued monitoring is imperative.

17. Limited data on individual stocks prevent more detailed assessments using precise analytical models from being employed. Thus, certain methods and assumptions adopted in this assessment were the only ones available.

ACKNOWL EDGMENTS

Appreciation is extended to personnel at the Nain Fish Plant, in particular to Mr. Jim Webb, Manager, his wife Sue and Sarah Webb, for their continued interest and cooperation. Mr. Terry Nicholls and Herb Mullett prepared the figures while Reg Tucker provided computer assistance. Finally I wish to thank biologists Dave Reddin, Bob Wiseman, Henry Lear and Dr. John Pippy, Program Head, for their helpful suggestions concerning the text. Mrs. Terry Hutchings kindly typed several drafts of the report.

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1957. On the dynamics of exploited fish populations. U.K. Min. Agric. Fish., Fish. Invest. (Ser. 2) 19: 533 p.

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32

Bowering, W. R., and T. K. Pitt. 1978. An evaluation of the status of witch flounder, Glyptocephalus cynoglossus, from ICNAF Divisions 2J, 3K and 3L. ICNAF Selected Papers No.3: 23-28.

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1968. Appraisal of a fishery. In Methods for assessment of fish production in freshwaters, 236-245, W. E. Ricker [Ed.], IBP Handbook No.3, Blackwell Scientific Publications, Oxford and Edinburgh.

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33

ICNAF. 1972. Report of Assessments Subcommittee. Int. Comm. Northwest Atl. Fish. Redb. Part I, p. 41.

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Peet, R. F. 1968. A report on a reconnaissance survey and counting fence operation conducted in south-east Labrador during 1966. Prog. Rep. No. 48. Res. Dev. Branch, Newfoundland Region, 153 p.

1971. A report on the counting trap and reconnaissance surveys conducted in central coastal Labrador during 1967. Prog. Rep. No. 68. Res. Dev. Branch, Newfoundland Region, 286 p.

Reddin, D. G. 1978. The Newfoundland and Labrador 1976 Atlantic salmon commercial catch data. Fish. Mar. Servo Tech. Rep. in Progress.

Ricker, W. E. 1958. Handbook of computations for biological statistics of fish populations. Bull. Fish. Res. Board Can. 119: 300 p.

34

1975. Computation and interpretation of biological statistics of fish populations. Bull. Fish. Res. Board Can. 191: 382 p.

Robertson, M. R., and D. H. Dowler. 1973. The history of exploitation of the Arctic char of Tree River, N.W.T. Paper presented at CCFFR, Halifax, 10 p. (manuscript).

Sandeman, E. J. 1969. Age determination and growth rate of redfish, Sebastes sp., from selected areas around Newfoundland. Int. Comm. Northwest Atl. Fish. Res. Bull. 6: 79-106.

Scott, W. B., and E. J. Crossman. 1973. Freshwater fishes of Canada. Bull. Fish. Res. Board Can. 184: 966 p.

Yessipov, W. 1935. Materials on the life history and fishery of the char of Novaya Zemlya (Salvelinus alpinus L.). Trans. Arctic Inst. USSR. 17: 5-70. (Russian with English summary).

35

A[)pendix 1. Summary of Arctic char landings (kg round) by area, size catcqory and by week, 1977.

l. Anton's Point

2. Voi sey Bay

3. Anaktal i k

4. Dog Island

5. Nain Bay

6. Tikkoatokak

7. Webb Bay

8. Black Island

9. Kiglapaits

10. Tasiuyak - no char

ll. Okak

12. Cutthroat

13. Mugford

14. Napartok

15. Hebron

36

Appendix I. (1) Anton's Point.

Size category Under 2.3 2.3 - 4.5 Over 4.5

Week kg % kg % kg % Weekly tota 1

1 2 3 508.9 75.8 162.3 24.2 671.2 4 243.0 85.5 41.1 14.5 284.1 5 200.9 69.6 87.8 30.4 288.7 6 222.8 73.1 81.9 26.9 304.7 7 346.3 75.0 115.3 25.0 461.6 8 9

10 11 12

Total 1521. 9 75.7 488.4 24.3 2010.3

Appendix 1. (2) Voisey Bay.

Week Under 2.3

kg %

Size category 2.3 - 4.5

kg % Over 4.5

kg % Weekly total

1 2 3 4 5 6 7 8 9

10 11 12

1106.9 5915.2 3640.4 1777.0 973.9 857.1 428.1

54.9 64.8 62.4 75.0 74.7 68.3 73.2

910.3 3213.0 2192.6 591.0 330.3 398.0 156.4

45,1 35.2 37.6 25.0 25.3 31.7 26.8

2017.2 9128.2 5833.0 2368.0 1304.2 1255.1 584.5

Total 14698.6 115.4 7791.6 34.6 22490.2 --­ -' _. -_.~. _.--- "._- .... _.. - ­ . ... - - - _.-.­

37

Appendix I. (3) Anaktalik.

Size category Under 2.3 2.3 - 4.5 Over 4.5

Week kg % kg % kg % Weekly total

1 2 3 4 5 6 7 8 9

10 11 12

1513.1 4497.1 5123.4 1354.4 657.2 160.1 151.3

61.1 56.0 65.4 70.2 70.9 76.2 80.3

964.5 3538.8 2711.8 575.2 270.2 50.1 37.2

38.9 44.0 34.6 29.8 29.1 23.8 19.7

2477.6 8035.9 7835.2 1929.6 927.4 210.2 188.5

Total 13456.6 62.3 8147.8 37.7 21604.4 -,----­ - ... -­ -­

Appendix I. (4) Dog Island

Week Under 2.3

kg %

Size category 2.3 - 4.5

kg 0'70

Over 4.5 kg % Weekly total

1 2 3 4 5 6 7 8 9

10 11 12

135.8 266.9 332.2 176.2 21.9 57.7

396.9 58.8

416.3

93.5 89.9 92.6 92.6 85.5 85.5 94.0

100.0 88.1

9.5 29.9 26.4 14.0 3.7 9.8

25.4

56.3

6.5 10.1 7.4 7.4

14.5 14.5 6.0

11.9

145.3 296.8 358.6 190.2 25.6 67.5

422.3 58.8

472.6

Total 1862.7 91. 4 175.0 8.6 2037.7 .­ - ­ .. '. --.

38

Appendix 1. (5) Nain Bay.

Si ze ca tegory Under 2.3 2.3 - 4.5 Over 4.5

Week kg % kg % kg % Weekly total

1 304.4 76.3 94.5 23.7 398.9 2 1107.1 63.4 638.6 36.6 1745.7 3 473.7 82.1 103.0 17.9 576.7 4 783.7 86.6 120.7 13 .4 904.4 5 2011. 6 91.3 190.9 8.7 2202.5 6 2195.6 93.6 151.1 6.4 2346.7 7 277 .6 96.5 10.1 3.5 287.7 8 9

10 11 12

Total 7153.7 84.5 1308.9 15.5 8462.6

Appendix I. (6) Tikkoatokak.

Week Under 2.3

kg %

Si ze 2.3

kg

category - 4.5

% Over 4.5

kg % l~eekly total

1 2 3 4 5 6 7 8 9

10 11 12

3864.9 8497.1 6161. 4 41191. 0 3236.4­2891. 5 2451. 9

81.9 77 .8 74.5 82.4 82.0 84.8 88.6

852.4 2427.8

·2110.3 958.4 712.5 518.4 315.4

18.1 22.2 25.5 17.6 18.0 15.2 11.4

4717.3 10924.9 8271. 7 5449.4 3948.9 3409.9 2767.3

Total 31594.2 80.0 7895.2 20.0 39489.4 .... - . -

39

Append i x I. (7) vJebb Bay.

Size category Under 2.3 2.3 - 4.5 Over 4.5

vieek kg % kg % kg % Weekly total

1 2 3 29.3 62.3 17.7 37.7 47.0 4 553.0 79.8 140.1 20.2 693.1 5 813.6 78.3 225.4 21.7 1039.0 6 7 226.5 84.3 4·2.2 15.7 268.7 8 312.8 88.8 39.4 11.2 352.2 9 103.5 89.0 12.8 11. 0 116.3

10 11 12

Tota 1 2038.7 81.0 477 .6 19.0 2516.3 ... --------­

Appendix I. (8) Black Island.

Size category Under 2.3 2.3 - 4.5 Over 4.5

Week kg % kg % kg % Weekly total

1 2 3 4 27.8 100.0 27.8 5 640.1 90.8 64.6 9.2 704.7 6 870.1 91.7 78.7 8.3 948.8 7 553.0 85.5 93.7 14.5 646.7 8 610.4 88.5 79.6 11.5 690.0 9 284.7 92.4 23.3 7.6 308.0

10 45.9 82.0 10.1 18.0 56.0 11 7.3 100.0 7.3 12 1.2 100.0 1.2

Tota 1 3040.5 89.7 350.0 10.3 3390.5

40

Appendix I. (9) Kiglapaits.

Size category Under 2.3 2.3 - 4.5 Over 4.5

Week kg % kg % kg % Weekly total

1 2 3 4 37.4 67.8 17.8 32.2 55.2 5 2.4 10.1 21.4 89.9 23.8 6 973.2 68.1 456.0 31.9 1429.2 7 1779.8 82.1 387.6 17.9 2167.4 8 175.1 75.0 58.3 25.0 233.4 9 240.8 71.2 97.2 28.8 338.0

10 46.1 73.5 16.6 26.5 62.7 11 10.1 100.0 10.1 12

Total 3254.8 75.3 1065.0 24.7 4319.8

Appendix I. (11) Okak.

Size category Under 2.3 2.3 - 4.5 Over 4.5

Week kg % kg % kg % Weekly total

1 2 3 4 1941. 6 59.4 1326.4 40.6 3268.0 5 717.0 54.9 590.0 45.1 1307.0 6 250.9 63.8 142.1 36.2 393.0 7 508.8 74.1 177.5 25.9 686.3 8 1864.8 77 .8 532.4 22.2 2397.2 9 5258.0 79.4 1366.5 20.6 6624.5

10 2060.0 83.4 409.9 16.6 2469.9 11 1544.9 79.2 404.9 20.8 1949.8 12 3.4 100.0 3.4

Tota 1 14149.4 74.1 4949.7 25.9 19099.1

41

Appendix 1. (12) Cutthroat.

Size category Under 2.3 2.3 - 4.5 Over 4.5

Week kg % kg %­ kg . % Weekly total

1 2 3 4 105.2 81.1 24.5 18.9 129.7 5 532.0 86.2 85.3 13.8 617.3 6 2885.7 74.3 1000.6 25.7 3886.3 7 3565.3 72.2 1375.9 27.8 4941.2 8 1752.1 76.9 526.3 23.1 2278.4 9 793.8 75.6 256.9 24.5 1050.7

10 153.3 71.5 61.0 28.5 214.3 11 90.7 92.9 6.9 7.1 97.6 12

Tota 1 9878.1 74.7 3337.4 25.3 13215.5

Append i x 1. (13) Mugford.

Week Under 2.3

kg %

Size 2.3

kg

ca tegory - 4.5

% Over 4.5

kg % Weekly total

1 2 3 4 5 6 7 8 9

10 11 12

280.7 602.8

52.2 71.9

256.9 235.1

47.8 28.1

537.6 837.9

Tota 1 883.5 64.2 492.0 35.8 1375.5

42

Appendi x I. (14) Napartok.

Size category Under2.3 2.3 - 4.5 Over 4.5

Week kg 'f" kg rx kg OJ, Week ly total

1 2 3 4 344.3 91.7 31.2 8.3 375.5 5 891. 5 75.9 283.6 24.1 1175.1 6 5988.7 73.3 2178.1 26.7 8166.8 7 3394.9 76.6 1036.3 23.4 4431. 2 8 883.2 84.5 162.4 15.5 1045.6 9 1777.6 90.9 178.1 9.1 1955.7

10 1508.9 85.6 253.3 14.4 1762.2 11 104.1 49.2 107.5 50.8 211. 6 12

Tota 1 14893.2 77 .9 4230.5 22.1 19123.7 - -.- . -_.

Appendix 1. (15) Hebron.

Size category Under 2.3 2.3 - 4.5 Over 4.5

Week kg % kg % kg % Week ly total

1 2 3 4 5 6 1137.8 78.0 320.8 22.0 1458.6 7 1127.6 83.3 225.9 16.7 1353.5 8 1939.7 86.7 298.4 13.3 2238.1 9 67.5 72 .3 25.9 27.7 93.4

10 707.9 87.3 103.4 12.7 811.3 11 12

Total 4980.5 83.6 974.4 16.4 5954.9

43

Appendix II. Summary of Atlantic salmon landings (kg round) by area, size category and by week, 1977 .

l. Anton's Point

2. Voisey Bay

3. Anaktalik

4. Dog Island

5. Nain Bay - No salmon

6. Tikkoatokak

7. Webb Bay

8. Black Island

9. Kiglapaits

10. Tasiuyak - No salmon

11. Okak

12. Cutthroat

13. Mugford - No salmon

14. Napartok

15. Hebron

44

Appendix II. (1) Anton's Point.

Size category Under 2.72 kg Over 2.72 ~

\\Teek kg % kg % ~'Jeekly total

1 2 -3 0.0 0.0 22.1 100 22.1 4 0.0 0.0 13.4 100 13.4 5 40.2 58.9 28.1 41.1 68.3 6 0.0 0.0 18.2 100 18.2 7 0.0 0.0 4.3 100 4.3 8 9

10 11 12

Summary 40.2 31. 8 86.1 68.2 126.3

Appendix II. (2 ) Voisey Bay.

~7eek

Under kg

2.72 Size kg

%

category Over kg

2.72 kg % ~7eekly total

1 2 3 4

.5 6 7 8 9

10 11 12

0.0 1.1 2.2 0.0 2.3 0.0

0 12.5 7.1 0.0

19.3 0.0

4.3 7.7

28.9 19.3

9.6 7.8

100 87.5 92.9

100.0 80.7

100.0

4.3 8.8

31.1 19.3 11. 9

7.8

SUIT1n1ary 5.6 6.7 77.6 93.3 83.2

45

Appendix II. ( 3) f\nakta1ik.

\'1eek Under kg

2.72 Size kg

%

category Over kg

2.72 kg % Weekly total

1 2 3 4 5 6 7 8 9

10 11 12

Summary

0.0 0.0 0.0

0.0

4.0 4.0

19.1

27.1

100.0 100.0 100.0

100.0

4.0 4.0

19.1

27.1

Appendix II. (4 ) Dog Island

v-7eek Under kg

2.72 Size kg

%

category Over kg

2.72 kg % Weekly total

1 2 3 -4 70.9 11. 4 551. 0 88.6 621. 9 5 442.4 22.6 1517.1 77.4 1959.5 6 332.4 38.0 542.2 62.0 874.6 7 410.1 44.2 517.4 55.8 927.5 8 137.5 34.7 258.9 65.3 396.4 9 159.4 35.1 295.4 64.9 454.8

10 67.1 18.4 296.8 81.6 363.9 11 19.7 10.2 173.0 89.8 192.7 12 24.4 11. 7 184.9 88.3 209.3

Surr[1~C'.ry 1663.9 27.7 4336.7 72.3 6000.6

46

-­ - ­ -.-- ­

Appendix II. (6 ) Tikkoatokak.

\'7eek Under kg

2.72 Size kg

%

category Over kg

2.72 kg % ~'Teekly total

1 2 3 4 5 6 7 8 9

10 11 12

0.0 0.0 0.0

0.0 0.0 0.0

7.9 9.1 5.1

100.0 100.0 100.0

7.9 9.1 5.1

SUTnmary 0.0 0.0 22.1 100.0 22.1

Appendix II. (7 ) vTebb Bay.

~veek

Under kg

2.72 Size category kg Over

% kg 2.72 kg

% ~'1eekly total

1 2 3 4 5 6 7 8 9

10 11 12

Summary

3.0 15.4 19.6

24.8 9.1

71.9

25.4 1~ .1 47.9

40.8 67.9

30.5

8.8 93.8 21.3

36.0 4.3

1601.2

74.6 85.9 52.1

59.2 32.1

69.5

11.8 109.2

40.9

60.8 13.4

236.1

47

Appendix II. (8 ) Black Island.

v-7eek Under kg

2.72 Size kg

%

category Over kg

2.72 kg % Weekly total

1 2 3 4 5 6 7 8 9

10 11 12

Surrmary

1.7 224.8 410.3 439.0 412.8 384.5 216.5 151. 4

88.6

2329.6

12.7 18.2 32.5 34.0 32.5 27.9 31. 3 20.9 15.3

26.9

1.1. 7 1011.2

851.8 853.5 857.4 991. 6 690.9 573.2 490.9

6332.2

87.3 81.8 67.5 66.0 67.5 72.1 68.7 79.1 84.7

73.1

13.4 1236.0 1262.1 1292.5 1270.2 1376.1

907.4 724.6 579.5

8661. 8

Appendix II. (9 ) Kiolapaits.

Week Under kg

2.72 Size kg

%

category Over kg

2.72 kg % Weekly total

•

1 2 3 4 5 6 7 8 9

10 11 12

11. 3 24.8

418.1 527.6 80.9

212.4 105.5

16.2

17.6 7.5

23.5 31. 3 34.7 25.4 28.4 27.9

52.8 307.4

1358.7 1160.0

152.6 622.7 265.4

41.9

82.4 92.5 76.5 68.7 65.3 74.6 71.6 72.1

64.1 332.2

1776.8 1687.6

233.5 835.1 370.9 58.1

SUTIunary 1396.8 26.1 3961. 5 73.9 5358.3

48

Appendix II. (11) Okak.

c'L,1.2C ca tcgorJ":

v-7eek Under kg

2.72 kg %

Over kg

2.72 kg "% \<Jeekly total

1 2 3 4 5 6 7 8 9

10 11 12

1.7 3.1

15.3 17.5 13.3 31.1 41.3 79.1

17.9 1~.0

21.0 22.9 21. 3 20.8 24.2 28.5

7. 8 19.0 57.6 59.0 49.3

118.8 129.6 198.8

82.1 86.0 79.0 77.1 78.7 79.2 75.8 71.5

9.5 22.1 72.9 76.5 62.6

149.9 170.9 277.9

Surr~ary 202.4 24.0 639.9 76.0 842.3

Appendix II. (12) Cutthroat.

\'1eek Under kg

2.72 Size kg

%

category Over kg

2.72 kg % Weekly total

1 2 3 4 41.5 5.9 663.9 94.1 705.4 5 731. 2 8.5 7854.9 91.5 8586.1 6 960.7 21.4 3533.5 78.6 4494.2 7 548.4 26.5 1518.0 73.5. 2066.4 8 283.5 36.1 484.8 63.9 768.3 9 257.5 34.7 484.5 65.3 742.0

10 54.5 31.6 118.1 68.4 172.6 11 18.7 23.1 62.4 76.9 81.1 12 4.3 10.5 36.6 89.5 40.9

Summary 2900.3 16.4 14756.7 83.6 17057.0

49

Appendix II. (14 ) Napartok.

\'leek Under kg

2.72 Size category kg Over

% kg 2.72 kg

% h'eek1y total

1 2 3 4 5 6 7 8 9

10 11 12

0.0 12.6

126.1 113.9

81.4 46.8

7.4 34.1

0.0 8.2

19.0 21. 3 32.0 21. 0 6.0

15.9

7.1 141. 5 537.1 420.5 173.3 176.1 116.4 180.2

100.0 91.8 81.0 78.7 68.0 79.0 94.0 84.1

7.1 154.1 663.2 534.4 254.7 222.9 123.8 214.3

Summary 422.3 19.4 1752.2 80.6 2174.5

Appendix II. (15 ) Hebron.

\'7eek Under kg

2.72 Size kg

%

category Over kg

2.72 kg % Feek1y total

1 2 3

•

"= 5 6 7 8 9

10 11 12

17.4 36.5 20.7

18.9

79.8 20.4 10.9

47.9

4.4 142.9 168.9

20.6

20.2 79.6 89.1

52.1

21.8 179.4 189.6

39.5

SUiTfl~ary 93.5 21.7 336.8 78.3 L!30.3

-- ------ ­ ~~--- - - --- ­

Appendix III. Arctic char landings by area and effort, for the period 1974-1976.

Total catch

Area 1974

(Round weight) (kg) 1975 1976

Man-weeks fished 1974 1975 1976

Catch per man-weeks (kg)

1974 1975 1976

Napartok Mugford Cutthroat

--

12,641

--

2,703

28,972 1,970 7,526

--

95

--

47

54 15 73

--

133

--

58

537 131 103

Okak 34,250 2,354 17,812 105 15 43 326 157 414 Tasiuyak Kiglapaits Black Island

1,467 5,131 4,264

-1,504 2,101

281 6,089 2,725

15 26 60

-32 62

2 59 48

98 197

71

-47 34

141 103

57 Webb Bay Tikkoatokak

580 9,960

833 27,695

4,550 31,568

1 28

5 76

15 81

580 356

167 364

303 390

Dog Island Nain Bay Anakta 1i k

2,659 12,461 7,821

653 -

2,548

212 3,119

14,670

38 37 28

40 -

10

11 10 45

70 337 279

16 -

255

19 312 326

(Jl

0

Voisey Bay Anton's Point

20,045 9,135

238 3,489

12,232 3,172

64 34

2 20

45 6

313 269

119 174

272 529

Summary 120,414 44,118 134,898 531 309 507 227 143 266

•

51

Appendix IV. (i) Age, length and weight data for Arctic char from Voisey Bay, 1977 .

•

Age Number

Mean Lgh. (em)

s.e. of

mean Range

Mean Wt. (kg)

s.e. of

mean Ranae

6 7 8 9

10 11 12 13 14

9 59

114 59 35 17 11 6 3

47.5 50.1 53.3 57.0 59.4 60.1 64.5 62.7 68.2

1. 42 0.51 0.36 0.58 0.96 1. 29 1.04 2.69 3.23

42.8-54.2 44.4-60.5 M.9-63.0 47.8-69.3 46.2-70.3 50.5-70.0 56.1-68.3 52.1-69.3 64.2-74.6

1.57 1. 75 2.06 2.58 2.91 2.84 3.65 3.42 4.10

0.16 0.06 0.05 0.10 0.17 0.16 0.22 0.46 0.47

1. 14-2.36 1.12-2.94 1.17-3.82 1. 30-4. 86 1. 28-5.46 1. 75-4.22 2.70-4.81 1.61-4.97 3.42-5.01

Summary 313 55.0 0.34 42.8-74.6 2.33 0.05 1. 12-5.46

Appendix IV. (ii) Age, length and weight data for Arctic char from Anaktalik, 1977 .

Age Number

Mean Lgh. (cm)

s.e. of

mean Ranqe

Mean Wt. (kg)

s.e. of

mean Range

•

•

6 7 8 9

10 11 12 13 14

2 12 60 42 43 27 9 4 2

50.3 51.8 54.3 57.2 60.3 63.1 62.6 62.4 61. 2

1. 90 1. 29 0.4·l 0.73 1.01 1.10 1.6·1 1.42 8.35

48.4-52.2 44.8-60.0 44.9-63.0 44.3-69.5 44.5-72.6 49.0-76.2 55.7-69.9 60.0-65.7 52.8-69.5

1. 79 1.98 2.19 2.55 3.11 3.39 3.32 3.16 3.32

0.17 0.14 0.05 0.11 0.17 0.23 0.25 0.27 0.31

1.61-1.97 1.41-3.06 1. 36-3.25 1. 44-4.66 1.49-6.60 1. 31-6.42 2.21-4.19 2.47-3.61 2.01-4.65

Summary 201 57.8 0.43 44.3-76.2 2.69 0.06 1.31-6.60

52

Appendix IV. (i i i) Aqe~ length and weight data for Arctic char from Tikkoatokak,1977.

- - _._-- ----­

Mean s.e. Mean s. e.

Age Number Lgh. (cm)

of mean Range

Wt. (kg)

of mean Range

6 7 8 9

10 11 12 13

29 134 136

79 39 14 7 2

41.7 46.1 49.8 54.2 57.8 58.1 59.4 68.7

0.71 0.36 0.39 0.61 0.82 0.85 1. 39 5.15

32.1-48.4 36.0-62.8 40.0-59.8 42.1-73.8 44.2-66.2 53.4-64.3 56.4-67.1 63.5-73.8

0.87 1.26 1.59 2.02 2.43 2.48 2.49 3.94

0.05 0.04 0.05 0.07 0.11 0.20 0.20 0.88

0.40-1.36 0.47-2.91 0.69-3.05 0.68-3.97 1.13-3.78 1.61-3.26 1. 95-3.46 3.06-4.84

14 1 55.3 1. 91

Summary 441 50.1 0.31 32.1-73.8 1.65 0.04 0.40-4.84

Appendix IV. (iv) Age, length and weight data for Arctic char from Okak Bay, 1977 .

Mean s.e. Mean s. e.

Age Number Lgh. (cm)

of mean Range

Wt. (kg)

of mean Range

6 1 49.3 0.00 49.3-49.3 1.65 0.00 1. 65-1. 65 7 1 48.2 0.00 48.2-~8.2 1.19 0.00 1.19-1. 19 8 3 51. 2 1. 31 49.5-53.8 1.96 0.35 1. 61-2.65 9 9 57.8 0.87 54.6-62.5 2.54 0.15 1. 96-3.08

10 11

22 26

60.4 57.3

1.03 0.86

5().9-66.9 51.0-67.2

2.81 2.53

0.14 0.11

1. 74-4 .10 1.77-4.53

12 43 60.3 0.7f! 50.6-74.6 2.77 0.10 1. 74-4.63 13 18 60.1 0.76 53.7-68.0 2.72 0.12 1.71-3.97 14 13 58.1 1.16 47.9-65.0 2.68 0.17 1.38-3.67 15 11 61.7 1.20 55.4-69.0 3.15 0.25 1.91-4.61 16 6 61.2 2.92 . 51.4-71.0 3.25 0.33 1. 84-3.87 17 1 64.4 3.72 18 1 68.4 3.66 19 1 70.8 5.08

Summary 156 59.4 47.9-74.6 2.75 0.05 1. 19-5.08

53

Appendix IV. (v) Age, length and weight data for Arctic char from Napartok, 1977 .

Mean s.e. Mean s.e.

Age Number Lgh. (cm)

of mean Range

Wt. (kg)

of mean Ranqe

8 3 53.3 2.99 47.5-57.5 1.84 0.27 1. 28-2.11 9 10 54.5 1.49 48.5-62.6 2.10 0.19 1.62-3.22

10 27 57.2 0.82 47.7-65.7 2.19 0.10 1.36-3.31 11 50 59.1 0.74 48.6-73.6 2.47 0.10 1. 36-5 .18 12 86 60.2 0.52 50.8-73.2 2.54 0.r7 1.46-5.17 13 44 60.7 0.69 52.8-70.8 2.63 0.10 1.50-3.99 14 33 02.7 1.00 52.7-75.1 2.83 0.15 1. 66-5.59 15 20 61.0 1.41 50.6-70.2 2.69 0.17 1. 55-4. fiO 16 5 59.3 2.09 54.2-64.6 2.32 0.29 1. 56-2.96 17 3 58.9 0.52 58.0-59.8 2.47 0.12 l.26-2.69 18 2 65.9 4.10 61.8-70.0 3.01 0.46 2.57-3.47 19 1 59.2 2.16 20 21 1 52.9 1.71

Surrunary 285 59.9 0.31 47.5-75.1 2.53 0.04 1. 28-5.59

Appendix IV. (vi) Age, length and weight data for Arctic char from Hebron, 1976-1977 .

Mean s.e. Mean s.e.

Age Number Lgh. (cm)

of mean Ran<le

Wt. (kg)

of mean Range

6 7 8 9

10 11 12 13 14 15 16

3 7

22 38 50 38 37 19 11 3 1

40.8 41.8 44.4 48.7 50.4 51.9 55.8 57.2 54.1 53.6 49.0

2.62 2.20 0.79 0.85 0.87 0.85 1. 05 1.83 1.77 1.86

37.5-46.0 30.1-47.5 33.9-51.4 42.4-62.2 32.7-63.1 37.7-66.2 44.5-71.1 39.5-69.3 39.3-60.4 50.0-56.2

0.98 1.07 1.23 1. 59 1. 74 1. 76 2.23 2.41 1.82 1.86 1. 56

0.19 0.14 0.06 0.07 0.09 0.10 0.14 0.24 0.16 0.27

0.72-1.35 0.33-1.38 0.69-1. 92 1.05-2.78 0.45-3.32 0.67-4.24 1. 26-4.50 0.55-5.00 0.69-2.63 1. 34-2.211

17 1 61.7 2.59

Summary 230 51.1 0.46 30.1-71.1 1. 79 0.05 0.33-5.00

54

Appendix V. lstimated numbers at age for Arctic char caught in various areas during 1977.

Area Voisey Bay Anaktalik Tikkoatokak Okak Napartok Hebron

Total* catch kg-round

22,490 21,604 39,489 27,598 28,047 5,955

Age

6 7 8 9

10 11 12 13 14 15 16 17 18 19 20 21

290 1,902 3,642 1,902 1,128

580 354 194

97

94 562

2,799 1,957 2,004 1,254

412 187 94

1,454 6,498 6,541 3,826 1,881

684 342 107

43

78 78

249 759

1,845 2,185 3,610 1,517 1,086

929 510

78 78 78

132 461

1,251 2,304 3,976 2,028 1,527

922 237 132 92 53

53

36 87

272 454 613 468 454 221 137

36 11 11

Total 10,089 9,363 21,376 13 ,080 13,168 2,800

*includes pickled landings for Okak and Napartok.