18

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

† Coelacanthus sharjah Khalaf, 2013 : A New

Coelacanth Fish Fossil Species from Sharjah

Natural History and Botanical Museum, Sharjah,

Emirate of Sharjah, United Arab Emirates

ةفحوور سكك: نوع جديد أل 3102س شارجة خلف ، سيالكانثو

الُجكبيز )سيالكانث( كن كتفحف الشارقة للتاريخ الطبيعي والنباتي ،

دولة اإلكارات العربية الكتفحد إكار الشارقة ، الشارقة ،

By: Prof. Dr. Sc. Norman Ali Bassam Ali Taher Khalaf-

Sakerfalke von Jaffa

Abstract: A new fossil species of Coelacanth Fish of the genus

Coelacanthus (Class Sarcopterygii, Subclass Crossopterygii, Order Coelacanthiformes, Family Coelacanthidae) from Sharjah Natural History and Botanical Museum in Sharjah, United Arab Emirates is described. The origin of the specimen is unknown. The new fossil species is distinguished from the other ten Coelacanthus species by its slightly different skeletal, skull and fin features. It is morphologically distinct from the other ten species. The new species was named †

Coelacanthus sharjah Khalaf, 2013.

The new Coelacanth fish fossil † Coelacanthus sharjah Khalaf, 2013 at Sharjah Natural

History and Botanical Museum, Sharjah, United Arab Emirates. 26.09.2013.

http://www.flickr.com/photos/50022881@N00/9962145603/

19

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

Keywords: Sarcopterygii, Coelacanthiformes, Coelacanthidae,

Coelacanthus sharjah, Coelacanth, Sharjah Coelacanth, Arabian Coelacanth, Emirati Coelacanth, Fossil, New species, Sharjah Natural History and Botanical Museum, Sharjah, Emirate of Sharjah, United Arab Emirates, Near East, Middle East.

The anterior part of the new Coelacanth fish fossil † Coelacanthus sharjah Khalaf, 2013

at Sharjah Natural History and Botanical Museum, Sharjah, United Arab Emirates.

26.09.2013. http://www.flickr.com/photos/50022881@N00/9967289834/

Introduction

During a visit to Sharjah Natural History and Botanical Museum in Sharjah, Emirate of Sharjah, United Arab Emirates on Thursday 26.09.2013, I examined and took measurements of a Coelacanth Fish Fossil.

After examining the Coelacanthus fish at Sharjah Natural History Museum, I began comparing between the different Coelacanthus fossil species.

There are ten Coelacanthus recognized species: † Coelacanthus banffensis Lambe, 1916; † Coelacanthus evolutus Beltan, 1980; † Coelacanthus gracilis Agassiz, 1844; †

20

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

Coelacanthus granulatus Agassiz, 1836; † Coelacanthus harlemensis Winkler, 1871; † Coelacanthus lunzensis Reis, 1900; † Coelacanthus madagascariensis Woodward, 1910; † Coelacanthus minor Agassiz, 1844; † Coelacanthus welleri Eastman, 1908; † Coelacanthus whitea Lehman, 1952.

The new fossil species † Coelacanthus sharjah Khalaf, 2013 is distinguished from the other ten Coelacanthus species by its slightly different skeletal, skull and fin features. It is morphologically distinct from the other ten species.

The middle part of the new Coelacanth fish fossil † Coelacanthus sharjah Khalaf, 2013

at Sharjah Natural History and Botanical Museum, Sharjah, United Arab Emirates.

26.09.2013. http://www.flickr.com/photos/50022881@N00/9967468845/

Description

Coelacanthus ("Hollow Spine") is a genus of extinct coelacanths that first appeared during the Late Permian period, 251 million years ago. In fact, this was the first genus of coelacanths ever described by Louis Agassiz in 1836, as the order Coelacanthiformes is named after it.

21

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

They bear a superficial similarity to the living Latimeria, though they were smaller, and had more elongated heads. Individuals grew up to 3 feet in length, and had small lobed fins, suggesting that Coelacanthus were open-water predators (Wikipedia).

Coelacanthus was a long-lived genus with a worldwide distribution. They survived the Permian–Triassic extinction event, and eventually died out during the end of the Jurassic, 145.5 million years ago.

The posterior part of the new Coelacanth fish fossil † Coelacanthus sharjah Khalaf, 2013

at Sharjah Natural History and Botanical Museum, Sharjah, United Arab Emirates.

26.09.2013. http://www.flickr.com/photos/50022881@N00/9967697023/

Measurements of the Sharjah Coelacanth Fossil

Total length: 35.00 cm; Head from mouth tip to gill cover: 8.00 cm; Tail length:

9.00 cm; Head width at the gill cover: 9.00 cm; Body width in the middle (dorsal-

ventral): 9 cm, tail base width: 6.5 cm; Anterior dorsal fin: 5.50 cm; Posterior

dorsal fin: 4.50 cm; Pectoral fin: 6.50 cm; Pelvic fin: 5.50 cm; Anal fin: 5.00 cm;

Tail rays: 6.00 cm.

22

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

Etymology / Derivation of the Scientific Name Coelacanthus in New Latin literally: hollow spine; ―Coel” indicating a cavity within a body or a hollow organ or part, from Greek ―koilos” hollow + Greek ―akanthos” spined, adj. derivative of ―ákantha” spine, referring to the hollow caudal fin rays of the first fossil specimen described and named by Louis Agassiz in 1836. The Arabic species name ―sharjah‖ refers to Sharjah, Emirate of Sharjah, United Arab Emirates where the fossil specimen is deposited at the Sharjah Natural History and Botanical Museum.

Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa is taking measurements of the new

Coelacanth fish fossil † Coelacanthus sharjah Khalaf, 2013 at Sharjah Natural History

and Botanical Museum, Sharjah, Emirate of Sharjah, United Arab Emirates. 26.09.2013.

http://www.flickr.com/photos/50022881@N00/9967928753/

Conclusion:

After studying, examining and measuring the Coelacanthus fish

fossil specimen at Sharjah Natural History and Botanical Museum,

and comparing with the different Coelacanthus species, and

referring to many zoological references, and searching the Internet,

23

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

I came finally to a conclusion that we are in front of a new

Coelacanth fossil species.

I gave it the scientific name † Coelacanthus sharjah, new fossil

species. The species name “sharjah” is for Sharjah, Emirate of

Sharjah, United Arab Emirates.

† Coelacanthus sharjah, new fossil species:

Scientific Binomial name: † Coelacanthus sharjah Khalaf, 2013

Authority: Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa.

Common Names: Sharjah Coelacanth, Arabian Coelacanth, Emirati

Coelacanth.

Holotype Fossil: CSNHM-1, 35.00 cm, Sharjah Natural History and

Botanical Museum, Sharjah, Emirate of Sharjah, United Arab

Emirates.

Origin: Unknown.

Date of Study: 26th September, 2013.

Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa is showing the new Coelacanth fish

fossil † Coelacanthus sharjah Khalaf, 2013 at Sharjah Natural History and Botanical

Museum, Sharjah, Emirate of Sharjah, United Arab Emirates. 26.09.2013.

http://www.flickr.com/photos/50022881@N00/9968030076/

24

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

References and Internet Websites

Animals. Wikia. Indonesian Coelacanth. http://animals.wikia.com/wiki/Indonesian_Coelacanth Bath Royal Literary and Scientific Institution. Coelacanthus lepturus Agassiz. http://www.brlsi.org/museum-collections/online-museum/fossils/23760 BioLib. Genus Coelacanthus Agassiz, 1836 †. http://www.biolib.cz/en/taxontree/id706087/ Bruton, M.N.; Cabral, A.J.P.; Fricke, H. (1992). First capture of a coelacanth, Latimeria chalumnae (Pisces, Latimeriidae), off Mozambique. S. Afr. J. Sci. 88. pp. 225—227. Butler, Carolyn. Bilder: Laurent Ballesta. Der Quastenflosser: Ein Fossil taucht auf. http://www.nationalgeographic.de/reportagen/der-quastenflosser-ein-fossil-taucht-auf Coelacanthus, Quastenflosser aus NW-Deutschland. www.fossilweb.de/fossilien/quastenflosser/ Courtenay-Latimer, M. (1979). My story of the first coelacanth. Occas. Pap. Calif. Acad. Sci. 134. pp. 6-10. Culture Grid. Coelacanthus granulatus. http://www.culturegrid.org.uk/search/2747678.html Egerton, P. (1850). New fish descriptions. A Monograph of the Permian Fossils of England, 233-236. † Coelacanthus caudalis Egerton 1850 (lobe-finned fish). http://paleodb.org/?a=basicTaxonInfo&taxon_no=149738 Erdmann, M.V.; Caldwell, R.L.; Moosa, M.K. (1998). Indonesian `King of the sea' discovered. Nature 395: 335. Forey, P.L. (1998). History of the Coelacanth Fishes. Chapman & Hall, Londres. 365 p. Freepedia. Quastenflosser. http://de.freepedia.org/Quastenflosser.html Fricke, H. (1997). Living coelacanths: values, eco-ethics and human responsibility. Marine Ecology Progress Series 166, 231-236. Fricke, H.; Hissmann, K. (1994). Home range and migrations of the living coelacanth Latimeria chalumnae. Marine Biology 120, 171-180. Fricke, H.; Hissmann, K. (2000). Feeding ecology and evolutionary survival of the living coelacanth (Latimeria chalumnae). Marine Biology 136, 379-386. Fricke, H.; K. Hissmann; J. Schauer; M. Erdmann; M.K. Moosa; R. Plante (2000). Biogeography of the Indonesian coelacanths. Nature 403, 38. Fricke, H.; K. Hissmann; J. Schauer; R. Plante (1995). Yet more danger for coelacanths. Nature 374, 314. Fricke, H.; K. Hissmann; J. Schauer; O. Reinicke; L. Kasang; R. Plante (1991). Habitat and population size of the coelacanth Latimeria chalumnae at Grande Comore. Environmental Biology of Fishes 32, 287-300. Fricke, H.; O. Reinicke; H. Hofer; W. Nachtigall (1987). Locomotion of the coelacanth Latimeria chalumnae in its natural environment. Nature 329, 331-333.

25

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

Hissmann, K.; H. Fricke; J. Schauer (1998). Population monitoring of the coelacanth (Latimeria chalumnae). Conservation Biology 12, 1-8. Hissmann, K.; H. Fricke; J. Schauer (2000). Patterns of time and space utilisation in coelacanths (Latimeria chalumnae), determined by ultrasonic telemetry. Marine Biology 136: 943-952. Huxley, T.H. (1861). Preliminary essay upon the systematic arrangement of the fishes of the Devonian epoch. Figures and descriptions illustrative of British organic remains. Mem. Geol. Surv. UK Dec. 10 (1861). pp. 1—40. Kappeler, Markus (1999). Komoren-Quastenflosser Latimeria chalumnae. Markus Kappeler / Groth AG. (erschienen in der WWF Conservation Stamp Collection, Groth AG, Unteraegeri). www.markuskappeler.ch/tex/texs/quastenflosser.html

Prof. Dr. Sc. Norman Ali Bassam Khalaf with a Coelacanth (Latimeria chalumnae)

stuffed specimen at the Educational Science Museum in Kuwait, State of Kuwait on

22.09.2013. Photo by my daughter: Nora Norman Ali Khalaf.

http://www.flickr.com/photos/50022881@N00/9896090416/

Khalaf, Norman Ali Bassam (1980). Tabie‘t Al-Talawon fi Al-Haywanat (The Colouration of Animals). Al-Biology Bulletin. Number 1. January 1980, Safar 1401. Biological Society, Kuwait University, State of Kuwait. pp. 4-5. (In Arabic). Khalaf, Norman Ali Bassam (1981). Fawa'ed Alasmak. (The Benefits of Fishes).

26

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

Al-Biology Magazine, Biological Society, Kuwait University, State of Kuwait. Number 1. Sunday 7.6.1981, 5. Sha'ban 1401. pp. 54-55. (In Arabic).

Sketch and measurements of the Sharjah Coelacanth Fossil † Coelacanthus sharjah

Khalaf, 2013. Done by: Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa.

http://www.flickr.com/photos/50022881@N00/10001840855/

27

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

Khalaf, Norman Ali Bassam (1982). Samak Al-Coelacanth (The Coelacanth Fish). Al-Biology Magazine. Number 2. February 1982. Biological Society, Kuwait University, State of Kuwait. pp. 14-15. (In Arabic). Khalaf, Norman (Translator) (1982). Al-Miah Al-Mulawatha Tohaded Al-Asmak Bi‘ilinqiraad (Water Pollution threatens the Fish Fauna with Extinction). Al-Biology Bulletin. Number 18, Third Year, First Semester, Saturday 6.11.1982. Biological Society, Kuwait University, State of Kuwait. pp. 7. (Translation from German into Arabic). Khalaf, Norman Ali (1983). Al-Samaka Al-‗Auljumiyah Al-Naqaqa fi Al-Khaleej Al-Arabi [The Toad Fish (Batrachus grunniens) in the Arabian Gulf]. Bulletin of the Biological Studies Club, Kuwait University, State of Kuwait. First Year, Number 3, 23 November 1983. pp. 10-11. (In Arabic).

Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa is taking measurements of the new

Coelacanth fish fossil † Coelacanthus sharjah Khalaf, 2013 at Sharjah Natural History

and Botanical Museum, Sharjah, Emirate of Sharjah, United Arab Emirates. 26.09.2013.

http://www.flickr.com/photos/50022881@N00/10002030764/

Khalaf, Norman Ali (Translator) (1983). Al-Tasjeel Al-Hay Al-Awal li-Samaket Kozat Al-Snobar (Monocentris japonicus, Houttuyn) min Al-Bahr Al-Ahmar [The Pinecone Fish (Monocentris japonicus, Houttuyn), A First Live Record from the Red Sea] by: Chaim Kropach. Bulletin of the Biological Studies Club, Kuwait University, State of Kuwait. First Year, Number 4, 7.12.1983. pp. 6-8. (In Arabic). Khalaf, Norman Ali B. (1986). The Schooling of Fishes. Gazelle: The Palestinian

28

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

Biological Bulletin. Number 9. Fourth Year. Ramadan 1406. May 1986. Department of Zoology, University of Durham, Durham, United Kingdom. pp. 1-13. Khalaf, Norman Ali B. (1986). The Fish Fauna in Van Mildert Pond, Durham City, North East England. Gazelle: The Palestinian Biological Bulletin. Number 9. Fourth Year. Ramadan 1406. May 1986. Department of Zoology, University of Durham, Durham, United Kingdom. pp. 14-20. Khalaf, N.A.B. (1986). The Schooling of Sumatra Barbs (Barbus tetrazona tetrazona) and Minnows (Phoxinus phoxinus). Dissertation, Master of Science in Ecology, Departments of Zoology and Botany, University of Durham, England. September 1986. pps. 59 + iv. Khalaf, Norman Ali B. (1987). The Coelacanth (Latimeria chalumnae) in the Science and Natural History Museum, State of Kuwait. Gazelle: The Palestinian Biological Bulletin. Number 15. Fifth Year. July 1987. Rilchingen-Hanweiler, Germany. pp. 1-8. Khalaf, Norman Ali Bassam (1987). On a Collection of Devon Period Animal Fossils from the Saarland, in the Geologische Museum Saarberg in Saarbrücken, Germany. Gazelle: The Palestinian Biological Bulletin. Rilchingen-Hanweiler, Federal Republic of Germany. Number 15, Fifth Year, Thul Qi‘dah 1407 AH, July 1987 AD. pp. 9-10. Khalaf, Norman Ali B. (1987). The Great White Shark (Carcharodon carcharias) from the State of Kuwait, Arabian Gulf. Gazelle: The Palestinian Biological Bulletin. Number 16. Fifth Year. Safar 1408 AH. September 1987 AD. Rilchingen-Hanweiler, Federal Republic of Germany. pp. 1-7. Khalaf, Norman Ali (1989). Qa‘ema li-ba‘d Asmak Al-Kuwait fi Al-Mathaf Al-‗Ilmi Bi-Dawlat Al-Kuwait (A List of some Kuwaiti Fishes from the Science & Natural History Museum, State of Kuwait). Gazelle: The Palestinian Biological Bulletin. Number 19. Seventh Year. December 1989. Bonn 2-Bad Godesberg, Federal Republic of Germany. pp. 3. (In Arabic and English). Khalaf-von Jaffa, Norman Ali Bassam (1991). A Trip to Zoo Budapest, Hungary. Gazelle: The Palestinian Biological Bulletin. Bonn-Bad Godesberg, Federal Republic of Germany. Number 21, Ninth Year, January 1991. pp. 1-4. Khalaf-von Jaffa, Norman Ali Bassam (1991). The Gulf War and its effect on the Arabian Ecosystem (Part One). Gazelle: The Palestinian Biological Bulletin. Bonn-Bad Godesberg, Federal Republic of Germany. Number 23, Ninth Year, July 1991. pp. 1-12. Khalaf-von Jaffa, Norman Ali Bassam (1991). The Gulf War and its effect on the Arabian Ecosystem (Part Two). Gazelle: The Palestinian Biological Bulletin. Bonn-Bad Godesberg, Federal Republic of Germany. Number 24, Ninth Year, August 1991. pp. 1-10. Khalaf-von Jaffa, Norman Ali Bassam (1991). The Gulf War and its effect on the Arabian Ecosystem (Part Three). Gazelle: The Palestinian Biological Bulletin. Bonn-Bad Godesberg, Federal Republic of Germany. Number 25, Ninth Year,

29

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

September 1991. pp. 1-7. Khalaf, Norman Ali Bassam (1992). Notes on the Biological Ecology of the Marshes in Southern Iraq. Gazelle: The Palestinian Biological Bulletin. Bonn-Bad Godesberg, Federal Republic of Germany. Number 29, Tenth Year, September 1992. pp. 1-9. (In Arabic).

The new Coelacanth fish fossil † Coelacanthus sharjah Khalaf, 2013 at Sharjah Natural

History and Botanical Museum, Sharjah, United Arab Emirates. 26.09.2013.

http://www.flickr.com/photos/50022881@N00/10002464264/

Khalaf, Norman Ali Bassam (1992). The United Nations Ecological Report confirms: The Regime of Saddam is destroying the Marshes (Al-Ahwar) Ecosystem. Sawt Al-Kuwait International Newspaper. Saturday 17 October 1992, 21 Rabi‘e Al-Thani 1412. pp. 15. (In Arabic). Khalaf-von Jaffa, Norman Ali Bassam (1992). An Introduction to the Animal Life in Palestine. Gazelle. Gazelle: The Palestinian Biological Bulletin. Bonn-Bad Godesberg, Federal Republic of Germany. Number 30, Tenth Year, October 1992. pp. 1-7. (In Arabic). Khalaf-von Jaffa, Norman Ali B. (1993). Al-Mushkilatan Al-Ma‘eyah wa Al-Bi‘eyah fi Al-Dafah Al-Gharbiyah wa Qita‘ Ghaza Al-Muhtalain (Ka-Juzu‘ min Al-Sharq Al-Awsat) [The Water and the Ecological Problems in the Occupied West Bank and Gaza Strip (As Part of the Middle East)]. Gazelle: The Palestinian Biological Bulletin. Number 31. Eleventh Year. December 1993. Bonn, Federal Republic of Germany. pp. 1- 29. (In Arabic). Khalaf-von Jaffa, Norman Ali Bassam (1994). An Introduction to the Animal Life in Palestine. Shqae‘q Al-Nouma‘n (Anemone coronaria). A Quarterly Magazine Issued by the Program EAI (Education for Awareness and for Involvement). Environmental Education / Children for Nature Protection. In Cooperation with Dept. of General and Higher Education. P.L.O., Palestine. Number 4. Huzairan (June) 1994. pp. 16-21. (In Arabic).

30

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

Acquaintance Card: Majallet Al-Ghazzal (Gazelle Magazine): The Palestinian Biological Bulletin. Bonn, Germany. Shqae‘q Al-Nouma‘n (Anemone coronaria). A Quarterly Magazine Issued by the Program EAI (Education for Awareness and for Involvement). Environmental Education / Children for Nature Protection. In Cooperation with Dept. of General and Higher Education. P.L.O., Palestine. Number 4. Huzairan (June) 1994. pp. 51-52. (In Arabic). Khalaf-von Jaffa, Norman Ali B.(1995). Alasmak fi Filistin (Die Fische von Palästina / The Fishes of Palestine). Gazelle: The Palestinian Biological Bulletin. Number 33. Thirteenth Year. December 1995. Bonn, Germany. pp.1-35. (In Arabic). Khalaf, Ali Bassam (1997). Amir Al-Bahar Al-Arabi (The Arabian Sea Prince) Shihab Al-Deen Ahmad Bin Majed. Magazin der Akademie. Nummer 1. Zu Elke‘da 1417 H, Maerz 1997. Koenig Fahad Akademie – Bonn, Bonn-Bad Godesberg, Deutschland. pp. 23-24. (in Arabic). Khalaf, Norman Ali Bassam (2001). The Extinct and Endangered Animals in Palestine. In: Gazelle: The Palestinian Biological Bulletin Home Page. Extinct and Endangered Animals and Reintroduction. http://gazelle.8m.net/photo3.html Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (Gründer) (seit Juni 2001). Yahoo! Deutschland Group: Wale und Delphine. http://de.groups.yahoo.com/group/Wale_und_Delphine/ Khalaf-von Jaffa, Norman Ali (2004). Gazelle: Das Palästinensische Biologische Bulletin. Eine Wissenschaftliche Reise in Palästina, Arabien und Europa zwischen 1983 – 2004. / Gazelle: The Palestinian Biological Bulletin. A Scientific Journey in Palestine, Arabia and Europe between 1983 – 2004. Erste Auflage / First Edition, Juli 2004: 452 Seiten / Pages. Zweite erweiterte Auflage (Second Extended Edition), August 2004: 460 Seiten / Pages. Norman Ali Khalaf, Bonn-Bad Godesberg, Germany. http://dr-norman-ali-khalaf-books.webs.com/ Khalaf-von Jaffa, Norman Ali (2005). Der Komoren-Quastenflosser (Latimeria chalumnae) und der Manado-Quastenflosser (Latimeria menadoensis). Gazelle: The Palestinian Biological Bulletin. Number 38. Twenty Third Year. February 2005. Sharjah, United Arab Emirates. pp. 1-8. http://quastenflosser.webs.com/ Khalaf-von Jaffa, Norman Ali (2005). The Story of Prophet Musa (Moses) and the Fish. Gazelle: The Palestinian Biological Bulletin. Sharjah, United Arab Emirates. Number 38, Twenty-third Year, February 2005. pp. 14-15. Khalaf-von Jaffa, Norman Ali (2005). Moses Perch (Lutjanus russelli, Bleeker 1849). Gazelle: The Palestinian Biological Bulletin. Sharjah, United Arab Emirates. Number 38, Twenty-third Year, February 2005. pp. 15. Khalaf-von Jaffa, Norman Ali (2005). The Fish of Musa (Samak Musa). Gazelle: The Palestinian Biological Bulletin. Sharjah, United Arab Emirates. Number 38, Twenty-third Year, February 2005. pp. 16. Khalaf-von Jaffa, Norman Ali (2005). Samak Al-Luchs (Al-Hamoor) or the Orange-Spotted Grouper (Epinephelus coioides) in Palestine (Mediterranean Sea)

31

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

and the United Arab Emirates (Arabian Gulf). Gazelle: The Palestinian Biological Bulletin. Sharjah, United Arab Emirates. Number 39, Twenty-third Year, March 2005. pp. 1-6. Khalaf-von Jaffa, Norman Ali (2005). Jaffa (Yaffa): The History of an Old Palestinian Arab City on the Mediterranean Sea. Gazelle: The Palestinian Biological Bulletin. Sharjah, United Arab Emirates. Number 39, Twenty-third Year, March 2005. pp. 7-8. Khalaf-von Jaffa, Norman Ali (2005). The Andromeda Sea Monster of Jaffa. Gazelle: The Palestinian Biological Bulletin. Sharjah, United Arab Emirates. Number 39, Twenty-third Year, March 2005. pp. 8. Khalaf-von Jaffa, Norman Ali (2005). The Jewfish (Epinephelus itajara) / Der Riesenzackenbarsch oder Judenfisch (Epinephelus itajara). Gazelle: The Palestinian Biological Bulletin. Sharjah, United Arab Emirates. Number 39, Twenty-third Year, March 2005. pp. 9-12. Khalaf-von Jaffa, Norman Ali (2005). The Arabian Freshwater Fishes in the Arabia‘s Wildlife Centre, Sharjah Desert Park, Sharjah, United Arab Emirates. Gazelle: The Palestinian Biological Bulletin. Number 40, Twenty-third Year, April 2005. pp. 1-9. Sharjah, United Arab Emirates. http://www.geocities.com/jaffacity/Arabian_Freshwater_Fish.html Khalaf-von Jaffa, Norman Ali (Gründer) (seit Juni 2005). Der Quastenflosser: Coelacanth Latimeria Yahoo! Deutschland Group. http://de.groups.yahoo.com/group/Quastenflosser/ Khalaf-von Jaffa, Norman Ali (2005). The Koran Angelfish (Pomacanthus semicirculatus, Cuvier, 1831). Gazelle: The Palestinian Biological Bulletin. Number 44. Twenty-third Year. August 2005. Jamada Alakhira 1426. Sharjah, United Arab Emirates. pp. 1-8. http://koran-angelfish.webs.com/ Khalaf-von Jaffa, Norman Ali (2005). Aquatica Arabica. An Aquatic Scientific Journey in Palestine, Arabia and Europe between 1980 - 2005 / Aquatica Arabica. Eine Aquatische Wissenschaftliche Reise in Palästina, Arabien und Europa zwischen 1980 - 2005. Erste Auflage / First Edition, August 2005: 376 Seiten / Pages. Norman Ali Khalaf, Rilchingen-Hanweiler, Bundesrepublik Deutschland & Sharjah, United Arab Emirates. http://dr-norman-ali-khalaf-books.webs.com/aquaticaarabica.htm Khalaf, N.A.B. (2005). The Schooling of Sumatra Barbs (Barbus tetrazona tetrazona) and Minnows (Phoxinus phoxinus). [M.Sc. Dissertation in Ecology, Departments of Zoology and Botany, University of Durham, England. September 1986. pps. 59 + iv]. In: Aquatica Arabica. An Aquatic Scientific Journey in Palestine, Arabia and Europe between 1980 - 2005. Erste Auflage, August 2005. Norman Ali Khalaf, Rilchingen-Hanweiler, Bundesrepublik Deutschland & Sharjah, United Arab Emirates. pp. 28-93. Khalaf-von Jaffa, Norman Ali (2005). The Rafah Zoo in the Rafah Refugee Camp, Gaza Strip, Palestine : A Story of Destruction by the Israeli Occupation Army. Gazelle: The Palestinian Biological Bulletin. Number 46, Twenty-third Year,

32

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

October 2005, Ramadan 1426. pp. 1-11. Sharjah, United Arab Emirates. (In Arabic). Khalaf-von Jaffa, Norman Ali Bassam (2005). The Qalqilia Zoo and the Natural History Museum in the City of Qalqilia, West Bank, Occupied Palestine. Gazelle: The Palestinian Biological Bulletin. Number 47, Twenty-third Year, November 2005, Shawal 1426. pp. 1-10. Sharjah, United Arab Emirates. (In Arabic). Khalaf-von Jaffa, Norman Ali Bassam (Member of PALESTA) (2005). Palestinian Scientists and Technologists Abroad (PALESTA). Gazelle: The Palestinian Biological Bulletin. Number 47, Twenty-third Year, November 2005, Shawal 1426. pp. 11-12. Sharjah, United Arab Emirates. (In Arabic). Khalaf, Norman Ali Bassam Ali Taher (2006). Eine Persönlichkeit aus Jaffa, Palästina / A Personality from Jaffa, Palestine: Bassam Ali Taher Khalaf (Abu Ali) (1938-2006). Gazelle: The Palestinian Biological Bulletin. Number 56, Twenty-fourth Year, August 2006. pp. 8-19. Sharjah, United Arab Emirates. http://bassam-ali-taher-khalaf.webs.com/ Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (2006). Ein Besuch im Neunkircher Zoo, Neunkirchen, Saarland, Deutschland / A Visit to Neunkirchen Zoo, Neunkirchen, Saarland, Germany. Gazelle: The Palestinian Biological Bulletin. Number 59, November 2006. pp. 1-25. Sharjah, United Arab Emirates. (in Arabisch / Arabic). http://khalaf.homepage24.de/text_88839638_85658724_59480041_deutsch.html Khalaf, Norman Ali Bassam Ali Taher (2007). Zum 1. Todestag : Eine Persönlichkeit aus Jaffa, Palästina / The First Death Anniversary : A Personality from Jaffa, Palestine : Bassam Ali Taher Khalaf (Abu Ali) (1938-2006). Gazelle: Das Palaestinensische Biologische Bulletin. Nummer 62, Februar 2007, Muharram 1428 AH. Seite 11. Sharjah, Vereinigte Arabische Emirate. http://bassam-ali-taher-khalaf.webs.com/ Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (Gründer) (seit Juni 2007). Yahoo! Deutschland Group: Fauna Palaestina. http://de.groups.yahoo.com/group/Fauna_Palaestina/ Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (Gründer) (seit August 2007). Haie – Sharks Yahoo! Deutschland Group. http://de.groups.yahoo.com/group/Haie_Sharks/ Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (Gründer) (seit September 2007). Yahoo! Deutschland Group: Fauna Arabica. http://de.groups.yahoo.com/group/Fauna_Arabica/ Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (2007). Haywanat Filistin (Fauna of Palestine). Wikipedia, Al-Mawsu'a Al-Hurra (The Free Encyclopedia). Gazelle: The Palestinian Biological Bulletin. Number 69, Twenty-fifth Year, September 2007 CE, Sha‘ban 1428 AH. pp. 1-4. (in Arabic). http://ar.wikipedia.org/wiki/%D8%AD%D9%8A%D9%88%D8%A7%D9%86%D8%A7%D8%AA_%D9%81%D9%84%D8%B3%D8%B7%D9%8A%D9%86 Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2007). A

33

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

Whale Shark (Rhincodon typus, Smith 1828) caught off the Kuwaiti Coast: The Second Record from the State of Kuwait, Arabian / Persian Gulf. Gazelle: The Palestinian Biological Bulletin. Number 71, November 2007. pp. 1-20. Sharjah, United Arab Emirates. (Abstracts in English and Arabic). http://whale-shark.webs.com/whalesharkinkuwait.htm

Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa at Sharjah Natural History and

Botanical Museum, Sharjah, Emirate of Sharjah, United Arab Emirates. 26.09.2013.

http://www.flickr.com/photos/50022881@N00/10003070683/

Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2007). Rhiniodon typus Smith, 1828 or Rhincodon typus Smith, 1829: The Story of a Scientific Name. Gazelle: The Palestinian Biological Bulletin. Number 71, November 2007. pp. 21. Sharjah, United Arab Emirates. http://whale-shark.webs.com/rhiniodontypus.htm Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2007). Whale Sharks in Palestinian Waters: A Whale Shark (Rhincodon typus, Smith 1828)

34

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

rescued near the Tantura Beach, Carmel Coast, North Palestine: The First Record from the Palestinian Mediterranean Coast. Gazelle: The Palestinian Biological Bulletin. Number 71, November 2007. pp. 22-23. Sharjah, United Arab Emirates. (Abstracts in English and Arabic). http://whale-shark.webs.com/ Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2007). Whale Sharks in Palestinian Waters: Whale Sharks (Rhincodon typus, Smith 1828) near Um Al-Rashrash (Eilat) Beach, Gulf of Aqaba, South Palestine: First Records from the Palestinian Red Sea Coast. Gazelle: The Palestinian Biological Bulletin. Number 71, November 2007. pp. 23-26. Sharjah, United Arab Emirates. (Abstract in English and Arabic). http://whale-shark.webs.com/ Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2007). An Ocean Sunfish or Common Mola (Mola mola, Linnaeus 1758) caught off the coast of Gaza: The First Record from Palestine, East Mediterranean Sea. Gazelle: The Palestinian Biological Bulletin. Number 72, December 2007, pp. 1-16. (Abstracts in English and Arabic). http://www.geocities.com/jaffacity/Gaza_Ocean_Sunfish.html Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2008). Cetacea Palaestina: The Whales and Dolphins in Palestinian Waters. Cetacean Species Guide for Palestine. Gazelle: The Palestinian Biological Bulletin. Number 83, November 2008, Thu Al-Qi‘ada 1429 AH. pp. 1-14. Sharjah, United Arab Emirates. http://cetacea-palaestina.webs.com/ Khalaf, Dr. Sc. Norman Ali Bassam Ali Taher (2009). Zum 3. Todestag : Eine Persönlichkeit aus Jaffa, Palästina / The Third Death Anniversary : A Personality from Jaffa, Palestine : Bassam Ali Taher Khalaf (Abu Ali) (1938-2006). Gazelle: Das Palästinensische Biologische Bulletin. Nummer 86, Februar 2009, Safar 1430 AH. Seite 21. Sharjah, Vereinigte Arabische Emirate. http://bassam-ali-taher-khalaf.webs.com/ Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2009). A Longcomb Sawfish (Pristis zijsron Bleeker, 1851) caught off the coast of Dibba, United Arab Emirates, Gulf of Oman. Gazelle: The Palestinian Biological Bulletin. Number 88, April 2009, Rabi‘e Al Thani 1430 AH. pp. 1-14. http://dibba-sawfish.webs.com/ Khalaf-Sakerfalke von Jaffa, Dr.Sc. Norman Ali Bassam Ali Taher (2009). Garra barreimiae wurayahi Khalaf, 2009 : A New Blind Cave Fish Subspecies from Wadi Al Wurayah Pools, Emirate of Fujairah, United Arab Emirates. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 90, June 2009, Jumada Al-Akhera 1430 AH. pp. 1-15. Sharjah, United Arab Emirates. http://emirati-blind-cave-fish.webs.com/ Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2009). Flora and Fauna in Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 91, July 2009, Rajab 1430 AH. pp. 1-31. Sharjah, United Arab Emirates. http://flora-fauna-palestine.webs.com/ Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2009).

35

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

Oreochromis mossambicus bassamkhalafi Khalaf, 2009 : A New Mozambique Tilapia Subspecies from Wadi Al Wurayah Pools, Emirate of Fujairah, United Arab Emirates. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 92, August 2009, Sha‘ban 1430 AH. pp. 1-25. Sharjah, United Arab Emirates. http://emirati-tilapia.webs.com/ Khalaf-von Jaffa, Dr. Norman Ali Bassam (2009). Fauna Palaestina – Part One. A Zoological Journey in Palestine, Arabia and Europe between 1983 – 2006 / Fauna Palaestina – Teil Eins. Eine Zoologische Reise in Palästina, Arabien und Europa zwischen 1983 – 2006. ISBN 978-9948-03-865-8. Erste Auflage/First Edition, September 2009: 412 Seiten/Pages. Self Publisher: Dr. Norman Ali Bassam Khalaf-von Jaffa, Sharjah, United Arab Emirates & Rilchingen-Hanweiler, Bundesrepublik Deutschland. http://dr-norman-ali-khalaf-books.webs.com/faunapalaestinapart1.htm Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2009). Bowmouth Guitarfish (Rhina ancylostoma Bloch & Schneider, 1801) at Sharjah Aquarium, Sharjah, United Arab Emirates. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Twenty-seventh Year, Number 93, September 2009, Ramadan 1430 AH. pp. 1-18. http://bowmouth-guitarfish-emirates.webs.com/bowmouthguitarfishuae.htm Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2009). Whale Shark (Rhincodon typus, Smith 1828) Records from the United Arab Emirates between 1989 - 2009. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Twenty-seventh Year, Number 94, October 2009, Shawal 1430 AH. pp. 1-28. Sharjah, United Arab Emirates. http://whale-shark.webs.com/whalesharkinemirates.htm Dr. Sc. Norman Ali Bassam Ali Taher Khalaf-Sakerfalke von Jaffa, Zoologist, Ecologist and Geologist : The Scientific References (1980-2009). http://dr-norman-ali-khalaf-references.webs.com/ Khalaf-von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2010). Fauna Emiratus - Part One. Zoological Studies in the United Arab Emirates between 2004 - 2009. / Fauna Emiratus – Teil Eins. Zoologische Studien in die Vereinigten Arabischen Emirate zwischen 2004 - 2009. ISBN 978-9948-15-462-4. Erste Auflage/First Edition, November 2010: 350 Seiten / Pages. Self Publisher: Dr. Norman Ali Bassam Khalaf-von Jaffa, Dubai and Sharjah, United Arab Emirates & Rilchingen-Hanweiler, Bundesrepublik Deutschland. http://dr-norman-ali-khalaf-books.webs.com/faunaemiratuspart1.htm Khalaf-von Jaffa, Dr. Norman Ali Bassam (2012). Fauna Palaestina – Part Two. Zoological Studies in Palestine between 1983 – 2009 / Fauna Palaestina - Teil Zwei. Zoologische Studien in Palästina zwischen 1983 – 2009. ISBN 978-9948-16-667-2. 1. Auflage / First Edition : July 2012, Shaaban 1433 H. 208 Seiten / Pages (Arabic Part 120 Pages and the English Part 88 Pages). Publisher: Dar Al Jundi Publishing House, Jerusalem, Palestine. http://dr-norman-ali-khalaf-books.webs.com/faunapalaestinapart2.htm

36

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

Khalaf-von Jaffa, Dr. Norman Ali Bassam (2013). Fauna Palaestina – Part Three. Zoological Studies in Palestine between 2005 – 2012 / Fauna Palaestina - Teil Drei. Zoologische Studien in Palästina zwischen 2005 – 2012. ISBN 978-9950-383-35-7. Erste Auflage / First Edition : July 2013, Shaaban 1434 H. 364 pages (English Part 350 Pages and the Arabic Part 14 Pages). Publisher: Dar Al Jundi Publishing House, Jerusalem, State of Palestine. http://dr-norman-ali-khalaf-books.webs.com/faunapalaestinapart3.htm Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2013). Garra rufa wadiqana Khalaf, 2013: A New Freshwater Doctor Fish Subspecies from Wadi Qana Nature Reserve, Salfit Governorate, State of Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 103, July 2013, Ramadan 1434 AH. pp. 1-25. Dubai and Sharjah, United Arab Emirates. http://palestine-fishes.webs.com/palestine-doctor-fish

Khalaf-von Jaffa, Prof. Dr. Sc. Norman Ali Bassam (2013). Taxon Profile: Subspecies Garra rufa wadiqana Khalaf, 2013. BioLib.cz. Biological Library. http://www.biolib.cz/en/taxon/id1059609/

Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2013). Palestine Doctor Fish (Garra rufa wadiqana Khalaf, 2013). EOL. Encyclopedia of Life. http://eol.org/collections/80813

Taxon profile << Go back one level - Coelacanthus

species

Sharjah Coelacanth

Coelacanthus sharjah Khalaf, 2013 †

kingdom Animalia - animals » phylum Chordata -chordates » class Sarcopterygii » order Coelacanthiformes -coelacanth » family Coelacanthidae

Other names

= Arabian Coelacanth

= Emirati Coelacanth

More >>

Taxon Profile: species Sharjah Coelacanth Coelacanthus sharjah Khalaf, 2013. BioLib.

Biological Library. http://www.biolib.cz/en/taxon/id1068520/

Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2013). †

Coelacanthus sharjah Khalaf, 2013 : A New Coelacanth Fish Fossil Species from Sharjah Natural History and Botanical Museum, Sharjah, Emirate of Sharjah, United Arab Emirates. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 -

37

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

6288. Number 106, October 2013, Thu Al Hijja 1434 AH. pp. 18–38. Dubai and Sharjah, United Arab Emirates.

http://quastenflosser.webs.com/coelacanthussharjah.htm Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2013). Picture of the new Coelacanth fish fossil † Coelacanthus sharjah Khalaf, 2013 at Sharjah Natural History and Botanical Museum, Sharjah, United Arab Emirates. 26.09.2013. Flickr. http://www.flickr.com/photos/50022881@N00/9962145603/ Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2013). Taxon Profile: Species Sharjah Coelacanth Coelacanthus sharjah Khalaf, 2013. BioLib. Biological Library. http://www.biolib.cz/en/taxon/id1068520/ Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2013). Sharjah Coelacanth († Coelacanthus sharjah Khalaf, 2013). EOL. Encyclopedia of Life. http://eol.org/collections/95987/

genus

Coelacanthus Agassiz, 1836 †

kingdom Animalia - animals » phylum Chordata - chordates » class Sarcopterygii » order Coelacanthiformes -coelacanth

Temporal range: Upper Permian - Upper Jurassic

[No associated text for selected language]

Subordinated taxa

Number of records: 11

species Coelacanthus banffensis Lambe, 1916 † species Coelacanthus evolutus Beltan, 1980 † species Coelacanthus gracilis Agassiz, 1844 †

species Coelacanthus granulatus Agassiz, 1836 † species Coelacanthus harlemensis Winkler, 1871 † species Coelacanthus lunzensis Reis, 1900 † species Coelacanthus madagascariensis Woodward, 1910 † species Coelacanthus minor Agassiz, 1844 †

species Coelacanthus sharjah Khalaf, 2013 - Sharjah Coelacanth †

species Coelacanthus welleri Eastman, 1908 † species Coelacanthus whitea Lehman, 1952 †

Taxon profile: genus Coelacanthus Agassiz, 1836. BioLib. Biological Library.

http://www.biolib.cz/en/taxon/id706087/

Lambers, Paul H. (1991).The identity of the type specimen of Coelacanthus harlemensis Winkler (Pisces, Actinistia) from the lithographic limestone of Solnhofen (Tithonian), Bavaria. Paläontologische Zeitschrift, June 1991, Volume 65, Issue 1-2, pp 173-189. http://link.springer.com/article/10.1007/BF02985782 Moy-Thomas, J. A. and T. S. Westoll (1935). On the Permian Coelacanth, Coelacanthus granulatus, Ag. Geological Magazine / Volume 72 / Issue 10 / October 1935, pp 446-457.

38

Gazelle : The Palestinian Biological Bulletin – Number 106 – October 2013

http://journals.cambridge.org/action/displayAbstract;jsessionid=EC3D0054E699FBFFBDFC5B1B21A66DB1.journals?fromPage=online&aid=4904620 Moy-Thomas, J. A. & E. I. White (1934). The Pectoral Fin of Coelacanthus tingleyensis. Nature 133, 499-499 (31 March 1934) | doi:10.1038/133499b0. http://www.nature.com/nature/journal/v133/n3361/abs/133499b0.html Nature. Ancient Coelacanths: Coelacanthus banffensis and whitea sp. Amazing Story! http://nature.ca/discover/treasures/foss/tr2/coe_e.cfm Pouyaud, Laurent; Wirjoatmodjoc, Soetikno; Rachmatikac, Ike; Tjakrawidjajac, Agus; Hadiatyc, Renny; Hadied, Wartono (1999). Une nouvelle espèce de coelacanthe. Preuves génétiques et morphologiques. A new species of coelacanth. Comptes rendus de l'Académie des sciences - Série III - Sciences de la vie / Life Sciences - 1999, 322, 261-267. www.elsevier.fr/html/news/cras3mars99/pouyaud.html Schauer, Juergen; Fricke, Hans. The Biology of the Coelacanth Latimeria chalumnae. www.orn.mpg.de/~hissmann/coela.html Schliewen, U.; Fricke, H.; Schartl, M.; Epplen, J.T.; Paeaebo, S. (1993). Which home for coelacanth?. Nature 363: 405. Smith, J.L.B. (1939). A living fish of mesozoic type. Nature 143. pp. 455— 456. Smith, J.L.B. (1956). Old Fourlegs: The Story of the Coelacanth. Longmans, Londres. The Fish out of Time. Coelacanth Information. www.dinofish.com/ The Fossil Forum. Coelacanthus granulatus, a Permian Coelacanth from Germany. 2011. http://www.thefossilforum.com/index.php?/gallery/image/18163-coelacanthus-granulatus-a-permian-coelacanth-from-germany/ Wikipedia. Coelacanth. http://en.wikipedia.org/wiki/Coelacanth Wikpedia. Coelacanthus. http://en.wikipedia.org/wiki/Coelacanthus Wikipedia. Quastenflosser. http://de.wikipedia.org/?title=Quastenflosser

The new Coelacanth fish fossil † Coelacanthus sharjah Khalaf, 2013 at Sharjah Natural

History and Botanical Museum, Sharjah, United Arab Emirates. 26.09.2013.

http://www.flickr.com/photos/50022881@N00/10002920596/