8/3/2019 Schneider Et Al 2010_ Cat Ba Langur

1/9

FciIgl {ll{ {|{}ocgofhl {k`ihovhs t|fhc{`cga o{ og cg`lvlg}hs kihhofivo}cul lg}lvtvc{l kigglk}cga o|}`iv{ gigtviec} t|fhc{`lv{ okomljck cg{}c}|}cig{ vl{lovk`

hcfvovcl{ ogm vl{lovk` e|gmlv{ cg }`l kijjig aioh ie jopcjcrcga okkl{{ }i kvc}ckoh vl{lovk`'

Fl`oucivoh If{lvuo}cig{ ogm Gi}l{ ig }`l Ulv}ckoh Vogacga To}}lvg ie

}`l Kvc}ckohhs Lgmogalvlm Ko} Fo Hoga|v (]vok`stc}`lk|{ Tihciklt`oh|{

Tihciklt`oh|{# cg Ucl}goj

O|}`iv({# 2C{oflhh [k`glcmlv Cgal @'J' ]clhlg Bi`oggo Viml Tcl}lv Hlulhcgn ogm Mogclho [k`v|mml

[i|vkl2 Tvcjo}l Kig{lvuo}cig (6

8/3/2019 Schneider Et Al 2010_ Cat Ba Langur

2/9???

Tvcjo}l Kig{lvuo}cig 64?4 (6 Uog Gaik ]`cg` l} oh' 64?4#' @ilulv

o eo{} mlulhitcga lkigijs kijfcglm c}` chhlaoh chmhcel

}voml `ofc}o} mlavomo}cig ogm `|g}cga `o{ hlm }i tit|ho}cig

mlkhcgl{ cg ohh {tlkcl{ (Hcttihm ogm U| Gaik ]`og` ?330#'

ovl ei|gm cg Ucl}goj (Jc}}lvjlclvl} oh' 6443#'

cg Ucl}goj ([k`v|mml l} oh' 6443# c{ }`l aihmlg*`lomlm ivKo} Fo hoga|v ]vok`stc}`lk|{ tihciklt`oh|{ tihciklt`oh|{

(]vi|l{{ov} ?3??# `ck` ikk|v{ ighs ig Ko} Fo C{hogm cg

giv}`lvg Ucl}goj ([k`v|mml l} oh' 6443#' C} flhiga{ }i }`l

{|feojchs ie }`l hloe lo}cga jignls{ (Kihifcgol# ogm o{ }`l

{johhl{} mc{}vcf|}cig ie ogs ie }`l hoga|v {tlkcl{ (Gomhlvl} oh'

ig }`l C\KG Vlm Hc{} (C\KG 6443# o{ o vl{|h} ie chhlaoh |g}*

cga `ck` `o{ hlm }i {johh c{iho}lm {|ftit|ho}cig{ ogm hi

tit|ho}cig g|jflv{' Tiok`cga `o{ vl{|h}lm cg o {lvci|{ tit|*

ho}cig mlkhcgl evij og l{}cjo}lm 6=446144 cgmcucm|oh{ cg

cg 6444 ([}lgnl 644;#' Kigklvglm ofi|} }`l mlkhcgcga g|j*

flv{ cg 6444 Jg{}lv Rii ogm }`l Riihiackoh [ikcl}s eiv }`l

Kig{lvuo}cig ie [tlkcl{ ogm Tit|ho}cig{ (RAOT# ei|gmlm

}`l Ko} Fo Hoga|v Kig{lvuo}cig Tviblk} (KFHKT#' ]`l tvib*

Hoga|v A|ovmcga Tviavoj ogm }`l l{}ofhc{`jlg} ie o hoga|v

{ogk}|ovs `o{ hlm }i og cgkvlo{l cg tit|ho}cig g|jflv{ }i

5414 Ko} Fo hoga|v{ o} tvl{lg} ([k`v|mml l} oh' 6443#' ]`l

vljocgcga cgmcucm|oh{ o} }`l }cjl ie vc}cga ovl evoajlg}lm

cg}i {cp {|ftit|ho}cig{ ie `ck` {ijl ovl ohh*eljohl avi|t{

`chl i}`lv{ ovl c{iho}lm ig {johh iee{`ivl c{hogm{'Hc}}hl cgeivjo}cig c{ ouochofhl kigklvgcga }`l lkihias

ogm fcihias ie }`l Ko} Fo hoga|v' Vl{lovk` ig mochs fl`ou*

civ to}}lvg{ l{tlkcohhs kog tviucml kv|kcoh mo}o eiv }`l kig*

{lvuo}cig jogoaljlg} ie }`l Ko} Fo hoga|v c}` vl{tlk} }i

tit|ho}cig jigc}ivcga' ]`l Ko} Fo hoga|v {`i{ {}vck} fl`ou*

civoh omot}o}cig{ }i c}{ tvlelvvlm hcjl{}igl nov{} `chh `ofc}o}'

]`vi|a`i|} }`l slov eiv lpojthl c} {hllt{ cg koul{ (Gomhlv

ogm Higa 6444#' ]`l Ko} Fo hoga|v c{ igl ie o g|jflv ie

}opo ie }`l ]' evogkic{c avi|t (Aviul{ 644? 644=> [k`v|mml

8/3/2019 Schneider Et Al 2010_ Cat Ba Langur

3/9

[k`glcmlvl} oh'

??6

l} oh' 6443# ie `ck` }`l `c}l*`lomlm hoga|v (]vok`stc}`l*

k|{ t' hl|kiklt`oh|{# cg K`cgo c{ c}{ khi{l{} vlho}cul' Hc}}hl

c{ ngig ofi|} }`l lkihias ogm fl`ouciv ie }`l ]' evogkic{c

avi|t `ck` jonl{ vlelvlgkl jo}lvcoh {kovkl' Cgul{}cao}cga

}`l fl`oucivoh omot}o}cig ie }`c{ {tlkcl{ }i c}{ `ofc}o} ki|hm

vlq|cvljlg}{' E|v}`lvjivl mo}o ofi|} }`l ulv}ckoh vogacga

fl`ouciv ie o {tlkcl{ kog tviucml cjtiv}og} cgeivjo}cig eiv

kig{lvuo}cig ogm jogoaljlg} tviavoj{ (Khljjig{ ogm

F|k``ihr ?331> Kovi ?330#' ]`c{ totlv ml{kvcfl{ }`l mochs

}cjl ohhiko}cig }i mceelvlg} ok}cuc}cl{ ogm }`l ulv}ckoh vogacga

fl`ouciv ie }i chm avi|t{ ie Ko} Fo hoga|v{'

Jl}`im{

[}|ms ovlo

Eclhmivn o{ kigm|k}lm ig Ko} Fo C{hogm cg @oc T`iga

Tviucgkl giv}`lvg Ucl}goj (64=6!64

8/3/2019 Schneider Et Al 2010_ Cat Ba Langur

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Fl`oucivoh if{lvuo}cig{ ie }`l Ko} Fo hoga|v

??;

ie eiim ogm {lovk`cga eiv eiim c}lj{ (lpojcgcga ulal}o}cig#'

[ikcoh fl`ouciv cgkh|mlm aviijcga ok}cuc}cl{ ogm thoscga

Cgmcucm|oh{ }`o} ki|hm gi} fl {llg m|vcga }`l {kog{ lvl

{ca`} m|vcga }`l if{lvuo}cig }`l if{lvulv{ oc}lm eiv }`lj

okkivmcga }i {crl ogm e|v kihiv2 cgeog}{ (kijthl}lhs ivogal#

b|ulgchl{ (aihmlg*slhhi `lom {`i|hmlv{ ogm lp}vljc}cl{#

ogm om|h}{ (aihmlg*slhhi }i kvlojs*`c}l lom{#'

]`lcv |{l ie }`l mceelvlg} lhluo}cig{ ig }`l khcee{ ogm chh{

o{ vlkivmlm o} lok` {kog' Ei|v hlulh{ lvl mc{}cga|c{`lm' O}

lok` {kog l gi}lm `lvl }`l jobivc}s ie }`l hoga|v{ lvl }i

fl ei|gm2 }it (|ttlv }`cvm ie }`l `chh iv khcee# jcmmhl (jcmmhl

}`cvm# fi}}ij (hil{} }`cvm# iv uohhls (hihogm#'

Eiv }`l mo}o ogohs{c{ l kohk|ho}lm }i}oh ki|g}{ ie lok`

fl`ouciv' Eiv }`l ogohs{c{ ie }`l ok}cuc}s to}}lvg l tiihlm

fl`ouciv vlkivm{ cg}i {cp }cjl*cg}lvuoh{ lok` ie }i `i|v{

{}ov}cga o} 45244 ogm lgmcga o} ?0244' ]`l }cjl }`l hoga|v{

{tlg} cg lok` fl`ouciv o{ }`lg kohk|ho}lm o{ o tlvklg}oal ie

}`l }i}oh ki|g} ie ohh fl`ouciv{ o# tlv oal kho{{ (}cjl f|mal}#

f# tlv }cjl cg}lvuoh (ok}cuc}s to}}lvg# ogm k# tlv lhluo}cig hlulh(vogacga to}}lvg#' ]`l }cjl }`l hoga|v avi|t{ lvl if{lvulm

cg lok` }cjl cg}lvuoh (}cjl {llg# o{ kohk|ho}lm o{ o tlvklg}*

oal ie }`l }i}oh if{lvuo}cig }cjl (

8/3/2019 Schneider Et Al 2010_ Cat Ba Langur

5/9

[k`glcmlvl} oh'

??=

]`ls ighs {tlg} 6'6, ie }`l }cjl cg }`l uohhls' Alglvohhs }`l

hoga|v{ jiulm evij igl lhluo}cig }i ogi}`lv ?'< }cjl{ tlv

`i|v' ]`l hoga|v{ {tlg} jivl }cjl vl{}cga ig }`l `chh {hitl{

ogm khcee{ ogm l{tlkcohhs glov }`l }it }`og lh{l`lvl' ]`l

}cjl {tlg} cg {ikcoh fl`ouciv mlkvlo{lm o{ }`ls ml{klgmlm

ogm {ikcoh fl`ouciv o{ gi} if{lvulm o} ohh cg }`l uohhls' Eiv*

oacga ok}cuc}s cgkvlo{lm kig{cmlvofhs cg }`l hilv lhluo}cig{

ogm o{ tvlmijcgog} cg }`l uohhls{ (Eca' [}ogeivm

?33?> Ehloahl ?333# ogm }`c{ c{ lucmlg}hs }`l to}}lvg eiv }`l

}i Ko} Fo hoga|v avi|t{ }`o} l `oul {}|mclm' Cgeog}{ ogm

b|ulgchl{ {tlg} kig{cmlvofhs jivl }cjl }`og }`l om|h}{ cg

{ikcoh fl`ouciv aviijcga ogm thoscga' ]`l {ojl c{ }v|l eiv

}`l `c}l*`lomlm hoga|v (Hc ogm Vialv{ 644=# ogm c{ o{{i*

kco}lm c}` }`lcv t`s{ckoh ogm fl`oucivoh mlulhitjlg} ogm

{ikcohcro}cig (Ticvclvl} oh' ?310#'

]`l hoga|v{ {tlg} j|k` ie }`lcv mos vl{}cga l{tlkcohhs

cg }`l jivgcga `i|v{ ogm ovi|gm jcm*mos |t }i ?=244 (jivl}`og 14, ie lok` 6*`i|v tlvcim#' I}`lv hoga|v{ {|k` o{

l} oh' 6441# }`l S|ggog {g|f*gi{lm

jignls (V`cgitc}`lk|{ fcl}c# (Mcga ogm R`oi 644=# ogm }`l

`c}l*`lomlm hoga|v (@|oga l} oh' 644;# ovl {cjchov cg }`c{

vl{tlk}' ]`l }i avi|t{ {tlg} hl{{ }cjl vl{}cga cg }`l oe}lv*

giig (

8/3/2019 Schneider Et Al 2010_ Cat Ba Langur

6/9

Fl`oucivoh if{lvuo}cig{ ie }`l Ko} Fo hoga|v

??