MISCELLAN OUS PUBLICAT ION OCCASIONAL PAPER NO , 80

Records of the Zoological Survey of ndia

STUDY ON INDIAN PENTATOMOIDEA (HETEROPTERA : INSECTA)

by

B. DATT,A, L. K. GHOSH AND MAYA DHAR

Issued by the Director Zoological 5ULvey of India, Calcutta

RECORDS OF THE

ZOOLOGICAL SURVEY OF INDIA

MISCELLANEOUS PUBLICATION OCCASIONAL PAPER No. 80

STUDY ON INDIAN PENTATOMOIDEA (HETE.ROPTERA : I NSECfA)

By

B. DATTA, L. K. GHOSH

and

MAYA DHAR

ZcoloRicai Survey of India, CaiCllf1a

~~i11f 51~~

Edited by the Director, Zoological Survey of India

1985

@ Copyright, Go~rnmei1t ~f India, 1985

Publi:slted August, 1985

PRICE: lalaDd : Rs. 36.00 ForeilD : ') £ 4.50 $ 7.00

Printed at The Pooran Preas, 21, Balaram Ohase Street, Calcutta-100 004 and Published by the DirectQr, Zoological Survey of India, Calcutla~

RECORDS

OF rrHE

ZOOLOGICAL SURVEY OF INDIA MISCELLANEOUS PUBLICATION

OCCASIONAL PAPER

No. 80

INTRODUCTION

SYSTEMATIC ACCOUNT

Family ASOPIDAE

CYoNIDAE

DIN IOORIDAE

GRAPHOSOMATIDAE

PEN TATOM IDAE

PLATASPIDAE

PBYLIOCEPHALIDAE

SCUTELLARIDAE

TESSAROTO MATIDAE

UROSTYLIDAE

SUMMARY

REFERENCES

1985

CONTENTS

Pages: 1-43

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26

STUDY ON INDIAN PENTATOMOIDEA (HETEROP"fERA INSE,CT A)

By

B. DAITA, L .. K. GHOSH and MAYA DHAR

Zoological Survey of India" 34, Chittaranjan Avenue,

Calcutta-12.

INTRODUCTION

Th'e, pentatomid bugs sustain their identities from rest of hemipteran species in having singular triangulate scutel1ar structure. They are comprised of 51,,1000 species of which 589 Orienta~ spec;~ are delineated by Distant (1902, 1908, 1918) on taxonomic aspects. The research workers have record'ed the convergence of pentatomid bugsl from Neotropica1, Indomalayasian and Ethiopian r.egions with brilliant shades of conlposite COIOT pigmentation as demar,cating fade~s of temperate forms from l~ropical ones.

The prelsent authors in conformity with the line of approacll adopted by Schouteden (1909), Leston (1952, 1954), Ruckes (1963), McDona1d (1966) and Ahmad and Kamaluddin (1982) have charac­terised the structuralJ' disslimj)ariti~6 of the ar/matures of the paramc'f\!S of sixty one spech~s belonging to forty four genera and observed thlet most re]~abl,e significant key traits as ,s[ufficien:t to pTeserve th;e distinctiveness of the individual sp,ecies. The 'e~onomic importance of the resp:octive species is further noted here. The authors have followed the exis,tin !!I nomen.c1atural status of the families.

~,

The authors are, grateful to Dr. B. K. Tikader:o D'irector, Zoological Survey of India, Calcutta for providing th'e fadilitie:s and encouragement.

2 REC. ZOOL. SURV. INDIA, Occ. PAPER No. 80

SYSTEMATIC ACCOUNT

Family A SOP IDAE

G~.nus Asopus Burmeister

1834. Asopus Burmeister, Nov. Act. A cad. Leop., 16 : 29·2-

1. Asopus maIabaricus (Fabricius) (Fjg. 1)

1775. Cimex nlalabaricus Fabricius, Syst. Ent. : 718

Body densely punctate, reddish; head with a spot at base; pronotum with 3 black spots;, two at anter-ior margin;. scuteHum spotted at basal angles·; forewing membrane black; abdomen black~ ventraHy with bands.

Length : 13 mnl. Genitalia Clasper (fig.. 1) robust, posterior margin distil®tl;y

rugulose, lateral margin curved and infolded, apically feebly wavy.

Distribution: North East and Western India',. Burma, Java, Sumatra, Born'eo and PhHippines.

Audinetia Ellener

1862. Audinctia Ellener, Nat. Tidsch,.. Nederl. Ind., 24 : 136

2. Audinetia sp~llidens (Fabricius) (Fig. 2, 2A)

1787. Cinzex spil1idens Fabricius, Ment. [nsf., 11 : 285

Body brownish yellow; head w1iltih a median lobe with blackL~ punctures on each slide.; antennal jOlintS black; a smooth line OlD.

pronotal angles; scutellum. at apex clay color; corium at lat~ angl~es clay-color; forewing InenlbralTIleJ smoky; body punctate ventrally; abdomen with blackish fasciae ven.trad; stigmata black.

Length: 14 mnl.

G'enitalia: Clasper (figs.. 2 & 2A) iSltrong1y cuIVled on. inner margin, gradually attenuated apicad, ventral part longitudinally infolded., basal apodrJrne prominent.

Distrihution: North East & South India, Malaya Archipelago, Fizi, Tahiti, East Africa, Abyssinia and Mexico.

DATTA, GHOSH & DHAR : Indian Pentaromoidea 3

Genus Picromerus Amyot & Serville

1843. Picrome,.us Amyot & Serville, Ess. Hem. : 84.

3. Picromerus obtutus Walker (Fig. 3)

1867. Picromerus obtutus Walker, Cat. Het. : 133

Bo<;ly thickly punctate; antennal 3-5 joints black, base yellow: pro notal anglesi black,. re-cury.~, apilces emarginate; costaF area rugu1ose; abdomen with black fasciae; femora speckled.

Length: 12 mm.

Genitalia: Clasper (fig. 3) chelale-shaped, a triangulate .structure project1ed media]]y, apex spatulate ..

Dl:')tribution: North East India & Burma.

F'a mil Y CYDNIDAE

Genus Cydnus Fabricius

1803. Cydinus Fabricius, Syst. Rhyn. : 184

4. Cydnus indicus Westwood (Fig. 4)

1837. Cydnus indicus \Vestwood, Hope. Cat., 1 : 19

Body piceous; h'ead coarsely punctate, laterally refl,exed, pilose; antennal Jast s.egments dilated, Pronotum (male) convex, a transverse impre1ssion, densely punc{at~, pilose laterad; scutellum punctate, apex and basa.f angles impunctate; fore\ving m'embrane smoky.

Length: 7 mm.

Genitalia: Clasper (fig. 4) broader apicad, apical margin ob1iquely dissected, subapicaJ' marglin setose, lateral margin clavalte.

Distribution: Western India, South Africa, S. E. Asia, Australia.

S. Cydnus maurus (Dallus) (Fig. 5)

1851. A:.thus maul'us Dallas, List Hom., 1 : 118.

Body dense],y punctured, shiny; pronotuml wilth a transvers'ely punctured furrow; scuteJlum thi,ckly punctate; corium dark, chestnut brown, veins dotted \vith punctures; abdomen. inlpunctate, J,uteraH) punctate!, postelior part denticulate.

4 REC. ZOOL. SURV. INDIA, OcC. PAPER No. 80

Length: 7 mm.

Genitalia: Clasper (fig. 5) laminate, parallel-sided, short, apex torose.

Distribution: Ind~a, Cey.Jbn.

Genus Macroscytus Fieber

1861. J\1acroscytus Fieber, Eur. Hem. 83 : 262.

6. Macroscytus subaenus (Dallas) (Fig. 6)

1851. A!thus subaenus DaUas, List Hem. 1 : 116.

Head without punct(l1tions, black, oed]j ~1; proootum ,brassy-­black, thicklly pundta1te ; :scutellum sparsely .punctate; forewing m:embrane darker at base; abdomen ventrally brassy-black, nlargin \vrinkled; sternum black, prost'ernum spa.rseJy punc1ate, rostrum and legs black.

Length 9 m1D.

Genitalia Clasper (fig. 6) robusrt, 1a min ate, inner margin rugulose upto lniddle., rest square-shaped with a notch at posterolo. lat~al margin, apical nlargin, convex, subapically with long setae.

Distribution : Bombay, Burma.

Famny DINIOORIDAE

Genus A~pongopus Laporte

7. Aspongopus assamensis Distant (Fig. 7)

1902. Aspollgopus assanlell~is Distant, Faulla Brit. India, 1 285.

Body piceous, antennae and conn,exivium black.

Lel)gth : 16 mm.

Genitalia : Clasper (fig. 7) cordate, 1laterally infolded" bow­shaped, margin finely sietose.

Distribution : Assanl, Khastt" Naga Hills.

Re111arks : Because of struci ural siInlhlarities of the para,mere the species 'should be merged with At,Spongopus nigriventri'S Westwood.

DATTA, GHOSH & DHAR : Indian Pentatom'Oidea 5

8. Aspongopus nigriventris Westwood (Fig. 8)

1837. Aspongopus nigriventris Westwood, Hope. Cat., 1 : 26.

Pronotum and scutellum with fine transver.5e series of punctures; antennaJ 2-3 joints shorter than 4-5 jointsl; legs brassy-black, hind tibiae ampliate mediaUy.

Length : 20 nlnl.

Genitalia : dasper (fig. 8), robust, cordate.

Distribution South India.

Genus Atelides Dallas

1852. Atelides Dallas, A.M.1V.H. X(2) : 360.

9. Atelides centrolineatus Dallas (Fig. 9)

1852. Atelides celltrolineCttus Dallas, A.M.N.H. X (2) : 360.

Body above brass!y-brOlw'Ilj, a centra]. fascia from anterior margin of head to abdomen dorsaUy; connerivium spotted; abdomen v'entromarginaUy fasciate.

l.ength : 18 mm.

Genitalia : Clasper (fig. 9) robust, basal part stalked, rest cordiform.

Distribution : Bangladesh.

Genus Cyclopelta Amyot & Serville

1843. Cyclopeltv, Amyot & Serville, &s. Hem. : 172.

10. Cyclopelta siccifolia (Westwood) (Fig. 10)

1837. Aspongopus sicci/olia Westwood, Hope Cat. 1 : 26.

Body totally bJack; pronotum wrinkl:ed, forewing membrane brownish yeJllow; connexiviium' uniform colo.r without SIPOltSi; scutelIar base with sanguineus .spot; abdonlen with red pigmentations; Inar­ginal spots and stigmata black.

Length : 15 mm.

Genitalia : Clasper (fig. 10) ovall-shaped, distally attenuate, shaft fis'Sate, cliscal margin setose.

6 REC. ZOOL. SURV. INDIA~ Oce. PAPER No. 80

Dilstribution : N. E .. and South India; Burma.

Economic status : The female aduHlts lay ~s on Cajanus cajan, Sesbania aegyptiaca and Erythrina indica., The adultls flock together and through punetures make. brown spots on epidennal tissues of plants.

Genus Megymenum Laporte

1832. MegYlnenum Laporte, Ess. Hem., :. 52.

11. Megymenum parallelum V~)llenhoven (Fig. II, ItA)

1868. Megymenul11 parallelum VollenhoV1("~ ; Foulla Ind. Nearl., In : 481:

Body brassy-black; head with lateral margins tumid; pronotal angles spinous, deeply concavus anteroIaterally, outeI1 margins scalloped; abdominal margiIn:s tubercufute at point of indisures papil­lose at centre of each ~egment.

Length : 16 rnm.

Genjtalic.' : Clasper (fig. 11, I1A) rod-sbapOO, salient lateroapi­C81Hy as figured.

Distribution : N. E. India, Andaman Islands.

Family GRAPHOSOMATIDAE

Genus Adrisa Amyot & Serville

1843. Adris(l Amyot & Serville, Ess. Hem., : 8-3.

12.. A~risa magna (Uhler) (Fig. 12)

1860. AcataleClus maglla Uhler, Proc. A cad. Nat. Soc. Phil.\ : 222.

Shiny, punctured head strongly punctate laterad; pronotum puvc­tured excll'uding smooth area at anterior region; scutellum centrally carinate, rugulate, punctate; corium punctate, forewing membrane smoky.

Length : 11 nlm.

Genitalia : Clasper (fig. 12) notched mediad, apex uncinat~ broader.

Distribution : N. E. India, Burma.

DATIA, GHOSH & DHAR : Indlan Pentatomoidea 7

Genus Podops Laporte

1832. Podops Laporte, Ess. Hem., : 72.

13. Podops lurida (Burmeister) (Fig. 13 )

1834. Telyra lurida Burmeister, Nov. Act. Ac. Leop, XVI : 288.

Body piceous; pronotal angles distinct, punctate, tranS~·l1Se furrow at anterior disc.

Length : 10 mm.

Genitalia : Qasper (fig. 13) broader apicad', feebly bifid, inner margin curved, lateroapically distinctly rugulate.

Distribution : N. E. India, China, Japan.

14. Podops bispinosa (Fabricius) (Fig. 14)

1798. Cimex bespillosa Fabricius, Ent. Syst. Supple : 529.

Body piceous ventratl;y, oensely punctate; antennae apically fuscous; pronotum w.ith a stout spine at anterior ang}le, lateral each a11@le with a slender spine. Scutellum with a piceous foveate CJpot at each basal angJe, basallly trispotted.

Length : 4 mm.

Genitalia : Cla-slper (fig. 14) short, d.ivergent ap'icad slightly, postero~aterad subtriangulate, inner margin serrate.

})islributiol1 : Bombay, Deccan Plateau.

Genus Stibaropus Dallas

1851. Stibaropus Dallas, List Hem., 1 : 125.

15. Stibaropus callidus (Schiodte) (Fig. 15)

1849. Scaptocoris callidus Schiodte, K,.oy. Nat. Tidsker, (2) : 460.

Head, pronotum, and scutellum! castarieous; pfonotum strongly wrinkled, a sha.llow transvers~, furrow nea,r base; scutellum punctate, rugulate; rostrun1 reaching ,length of prosternum; coriunl finely punctate"

Length: 5 mm.

Genitalia: CLasper (fig. 15) unciform, elongate.

Distribution: Eastern India" Burma.

8 REC. ZOOL. SURV. INDIA~ Oce. PAPER No. 80

Family PENTATOMIDAE

Genus Agonoscelis Spinola

1837. Agonoscelis Spinola, Ess. Hem. : 327

16. Agonoscelis nubila Fabricius (Fig. 16)

1775. Agonoscelis nubila Fabricius, 1775, Syst. Ent., : 712.

Body yellowish brown, black punctures along margins, median fascia to h'ead; scutellum antero]alterally with "clay-coloured rug~t-es;" coooexivium with spots at point of incisures; vt1Il~tioo on fOIewing membrane bJack, a double series of Slpdts on stennum and abdomen.

Length: 11 mm.

Genitalia: Clasper (fig. '16) robust~ foot-shaped\, infolded, inn.e.r margin with a long narrow basal plate.

Distribution: N .E. and N. W. India; Ceylon.

Economic ~tatus: The specre.s is common on Wheat and sugar­cane.

Genus Antestia Still

1864. Anteslia StAI, Henl. Afr. i : 200

17. Antestia ancltora (Thunberg) (Fig. 17)

1783. Cinlex Qnchora Thunberg, Nov. Ins. II : 47

Head before each eye with a 1in~r spot; pronotum wilth a series of sub-quadrate spots on posterior margin, an oblique stre'rdk on anterior margin; scutellum hexalIlotat~" basal angle spotted; forewing membrane black, margin transparent; abdomen: ventro~aterally bla"ck, extreme late.ral bands interce.pted.

L,fngth: 11 mm.

G~italia : Clasper (fig. 17) sharply geniculate medially, marginally infolded with an e100gate chrtinised structure extending apicad, margin setose.

Distribution: N.E. India, Burma.

DATTA, GHOSH & DHAR : Indilan Pentatnrnoidea

18. Antestia cruciata (Fabricius) (Fig. 18)

1775. Cimex c1uciafa Fabricius, Syst. Ent. : 714

H'ead with a pair of median 'Sitreaks~ spots near each eye.

Length: 8 mm.

9

Genitalia: Clasp,er (fig. 18) robust. arched wJ'ong inner margin, j·nfolded marginally.

Distribution: Sikkim, Calcutta, Bombay, Nilgiri HiJJs, Ceylon

Econolnic status: The species has: tre.Jmendous econom~c bearing. It occurs 'in great numbers usualUy in November and continues to stay till advent of rainy s:eaSlOn. The incidence of the species is involved in the destruction of II uits and vegetab!es and cereru)1 crops. These are co-nllmon on cofI'ee, fruits of Citrlls, Santalunl album and on Zyzyphus sp.

Genus Eusarcocoris Puton

1866. EusarcGco,.·is Puton, Cat. : 11

19. Eusar.cocoris capitatus Distant (Fig. 19)

1902. Eusarcocoris capitatus Distant, Fauna Brit. India, 1 : 169.

Body brownish ye~~ow, finely punctate ~ head deflected; pronOltuHl with a round SpOIt on anterior area;,: denSlely punctate, a median fascii!, black, extending from meso-sterna to abdomen ~ rostrum at apex ~lack; corium coarsely puncta.te; scutellum at base darker.

Length: 4 mm\.

Genitalia: (Jasper (fig. 19) abruptly narrowed, apex uncina'te.

Distribution: N.E. India.

Genus Stenozygum Fieber

1861. Stenoz.ygunl Fieber, Eur. Hem. : 345

20. Stenozygum speciosum (Dallas) (Fig. 20)

18151 Strachia spedosunl DaHas, List Hem. I : 261.

Head trinot;lte" a central fascia with a spot on each s~dc, two spots on anterior margjn, median fascia extending from pronotum to

2

10 REC. ZOOL. SURV. INDIA, Oce. PAPER No. 80

scutellum; basal angle 'spotted, subapically with a triangular patch; conltlexivium black, spotted yellow; sternum with· double series of spots.

Length: 8 m,m ..

Genitalia: Clasper (fig. 20) i broad~r, rest smoothly curved out forming an uncinate structure, a bunch of stiff bairs alt point of curvature.

Distributioll: N.W. and Southern Itnd~ia.

Genus Strachia Hahn

1831. Strachia Hahn, Wanz. i, : 180

21. Strachia crucigera Hahn (Fig. 21)

1831. Strachia crucigera Hahn, W onz. I : 184

Body blJlack; longitudinal fascia from pronotum to scutellum, later one tralltSversely branded; conium near membrane white, forewing membrane smoky; abdomen ventrallly black, ~n:tercepted by a 1in~.

Length: 9 mm.

Genitalia: Claspec (fig .. 21) aroh'ed along ill1ller margin, sagittaa groove distinct, aJong margin densely setose, clavate, ventro-mesal margin rugose.

Distribution : Kha:si RiBS', Burma, Malaya Peninsula, Indonesim Islands.

Genus Zicrona Amyot & Serville

1843. Zicr.)11c. Anlyot & Serville E~s. Henz., : 86

22. Zicrona caerulea (Linnaeus) (Fig. 22)

1758. Ci,nex cal!l'uiea Linnaeus, Syst. Nat. ed. 10 : 445

Body sky blue; antennal segments and forewing membrane black; surface fine~~ punctate; scutellum hump,.lshaped.

Length: 10 mm.

Genitalia: Clasper (fig. 22) geniculate mediad, basally divergent, abruptly narrowed to a long; proce~s, at right angle to )bng axis of body, in.ner margml infolded, spatulate.

Distribution: Burma, N.W, & N.E4 India, Malaya Archipelago~

DATTA, GHOSH & D'HAR : Indian Pentatomoldea 11

Genus Canthecona Anlyot & Serville

1843. Cantheeona Amyot & Serville, Ess. Hem., : 81.

23. Canthecona jUfeel/ala (Wolff) (Fig. 23)

1801. Cimex furcellala Wolff, Ie. V., : 182.

Body clay color, densely punctate; lateral 10beJs of head brassy­black; rostruln reddish at apex, pronotal all1g1ies projected into ~ort black spine's; scutellum thickly punctate, apically white; abdominal margin with a s'(\ries of green SPOltS.

Length : 15 mm.

Genitaliq : Clasper (fig. 23) genicuJa,te basad, nest scimitar-shaped with a pronlinent basal apodem'e,.

Distribution : Naga Hills.

Econolllic status : Predaceous. The insect feeds on cat~rpiUar Sltages of Jepidopteran insects. Adults! paralyse caterpillar by injcctir.g saliva into caterpHlar. 1st nymphal stages are sap-suckers and 2nd stages onward become predatory.

Genus Cappaea Ellener

1862. Cappaea Ellener, Nat. Tijdschr. Nederl. Ind. XXIV : 146,

24. Cappaea taprobanensis (Dallas) (Fig. 24)

1851. Penratoma taprobanellsis Dallas, List Hem., i : 244.

Body black with yellow markings; central fascia extendilllg: fro III tip of head to scutellar apex; pronotum laterally with linear reti­CUlations, ventrolaterally spoltted black; forewing m'embrane smoky, with brown spots.

Length : 10 mm.

Genitalia : Clasper (fig. 24) scimitar-shaped.

Distribution : N. E. India, Indones;an Islands.

GenU$ Dalpada Amyot & Serville

1843. DalplIda Amyot & Serville, Ess. Henl., : 105.

25. DaJpada affinis Dallas (Fig. 25).

1851. Dalpada affiltis Dallas List Heln. : 185.

Paler hue; pronotunl tuberculafte .late,rad; abdom.en with spar~e punctations, disc impunctate, jarger.

12 REC. ZOOL. SURV. INDIA, 'OcC. PAPER No. 80

Length : 18 mm.

Genitalia : Clasper (fig. 25) robust, f.oot-shaped, apex pointed, lateral margin infolded, densely nodulate, mne.r apical margin d/entate.

Distribution : N of E. India, Burma.

26. Dalpada oculata (Fabricius) (Fig. 26)

1775. Cimex ocuiata Fabricius, Syst. Ent., : 703.

Body marked olive green; head with a fas.cia, close to eyes SOllIe

spots; pronotal angles dark green with apical spot; c.onnexivium \vith alternate green and yellow pigmentations; abdomen black ventrally with a serres of spots.

Length : 11 nlm.

Genzitalia : Clasper (fig. 26) t broader, apex clava.te.

D4stribution : N. E. India, Burma.

Genus Fernelius Distant

1900. Fernelius Distant, Trans. ~"Ilt. Soc.,: : 173.

27. Fernelius indicus Distant (Fig. 27).

1900. Ferllelius ;ndi~us Distant, Trans. £12t. Soc., : 173.

Chestnut brown, covered with yellowish spots; antennae wilh bJ,ack spots; corium at apical area ISpotted, scutellar ang1Ies spotteo; abdomen reddish-yel!ow; connexivium spotted black;\ tibiae, tarsi, sterna, stigma black.

Length : 18 mnl.

Genitalia : Clasper (fig. 27) robuslt,. i distinctly broadeiC, laterally fluted in form of a beak.

Distribution : N. E. India, Burma.

Genus Guliemus Distant

1901. Gulienlus Distant, Entonloiogist : 347.

28. Gulielmus lateralis Distant (Fig. 28)

1901. Gulielnll1s laterlliis Distant, Entomologist : 347.

Extremity of head thick,J,y punctate; proootuln bin 0 tat:e, anteriorly, scutellar aprex with a b~ack spot; scutcllunl basally spotted; conneXl­viunl black; ubdonlen wi,th a black fascia laterad~ punctalte.

DATTA, GHOSH & DIHAR Indian ~entatofnoidea 13

Length : 9 mm.

Genitalia : Clasper (fig. 28) robust, inner margin fluted basad, densely rugulate, foot-shaped, apical margin dissected.

Di~tribution : Western India.

Genus Hoplistodera Westwood

1837. Hopli.'ilodera \Vestwood, Hope Cat. i : 18.

29. Hoplistodera viresceu:; Dallas (Fig. 29).

1851. Hopli.5todera virescells Dallas, List Hem., : 217.

Head orange yellow, sparsely punctate; antennal jointsl tawny; prono1um strongly punctate, anter.ior margin blinotate, salient laterau; scutellum tetranotate, punctate apicad; corium punctate.

l..£ngth : 8 mm.

Genitalia : Clasp~.r (fig. 29) J divergent apically unciform.

Distribution : Sikkim, Naga Hills, Burma.

Genus Menida Motschulsky

1861. Menida ~1otschulsky, Et. Ellt., X : 23.

30. Menida formosa (Westwood) (Fig. 30)

1837.- Pentatol11a fonnosa \Vestwood, Hope Cat., H,cm~ I : 34.

Body chestnut brown, shiny; head trifasciate, a snlaH spot before each eye; pronotunl at anterior nlargin annulate' folIow'ed by fasciae; scutel1llum with a catapult-marking, whlti~h, central line with a series of longitudilllal markings; ste.rnal disc castaneous.

Length : 7 mm.

Genitalia : Clasper (fig. 30) robust" ~pica;l third triangulate, apex pointed.

Distribution : Bombay, S. E. As,ian countries.

31. Menida histrio (Fabricius) (Fig. 31)

1787. Cimex histrio Fabricius, Mant. Inst. II : 296.

Body punctate, black; head tetralineate; pronotunl with a patt of annulate markings at anterior margin; sClItel;lIln at basal angle

14 REC. ZOOL. SURV. INDIA, Oce. PAPER No. sO

spotted; forium with a b]~ck spot at apex; connexivium spotted black; a series of spots Q1n abdomen ..

Length : 7 mm.

Genitalia : Clasper (fig. 31) sta!lked basad, abruptly broader apical half, dens.ely wrinkled.

Distribution : N. E. India, Burma.

Remark : The species is common on wheat and sugarcane.

32. Menida va rip ennis (Westwood) (Fig. 32)

1837. Pentalonu~ varipennis Westwood, Hope Cat. Hem. : 43.

Body brassy black; each eye with a spot at posterior margin; pronotum fasciate with linear tparkings; scut<f)um basally spoilted corium with a disca!1 spot, abdominal spineS1 reaching mid-coxa:e;.

Length : 7 mlm.

Genitalia : Clasper (fig. 32:) foot-shaped, rugulate, apex truncate ..

Distriblltion : N. E. India, Sikkinl, Burma,

Genus Nezara Amyot & Serville

1843. Nezara Amyot & Serville, Ess. Hem. : 143.

33. Nezara viridula (Linn.) (Fig. 33)

1758. Cinzex viridula Linn., Syst. Nat. ed. 10 : 444.

Body dark green, denseJy punctate; head binotate ; allltennae green, apically brown; abdomen ventrolatera!lly black; pronol.lIm trinotate; corium with green, spots.

Length : 14 mnl.

Genitalia : Clasper (fig. 33) robust, foot-sbaped, strongly genj· culate, dens,ely rugulate, apex sharply bent.

DAstribution : Bombay, South India.

Remark : The species is caUed green rice bug and alSOl occurs on wheat mitIets, and fibre crops.

DATTA, GHOSH & DHAR : Indl1an Pentat()/11oidea 15

Genus Priassus Walker

18617. Priassus, Stal, ,O/v. Vct.-A~. Forh : 518.

34. Priassus exemptus (Walker) (Fig. 34)

1868. Prionaca exemptus Walker, Cat. Het. iii : 569.

Body with fine pigmentation in form of punctures; h:e{ad and pronotum with reddish pigtnentation; corium coarsely punctate; abdo­men with a long spine reiachling in1termediate coxae.

Length: 18 mm.

Genitalia : Clasper (fig. 34) robust, sagittal leng.th infolded, forked asym~etrically at apex.

Distriblltion : Naga Hi1Jlts" Burma.

Family PLATASPIDAE

Genus Brachyplatys Boisduval

1.8,15. Brachyplatys Boisduval, Voy. As!r. Ellt. ii : 627.

35. Bracbyplatys punctipes l\Ilontand (Fig. 35).

1894. Brachyplatys punctipes MoIntand, Ann. Mus. Civ. Gen. 34 : 122.

Body shiny, olive color tinged with black, finely punctate; head hexanotate; antenna:e, piceous, pronotum wirh an oblique line sub­marginally; scuteLlum with four brown spots; st1ernum pitchy black; abdomen with a series of fasciae, inte.rcepted by a black stripe; felnur \vith distinct brown spots.

Lengt h: 8 rnm.

Genitalia: Clasper (fig. 35) robust, feebly geniculate, cyrnbiform along long axis, apex .sa~ienit.

Distribution : N.W India, Burnla, China.

36. Bracbyplatys radians Vollenhoven (Fig. 36)

18613.. Brachyplatys radians Vol1enhoven, Fauna Ind. Neerl. I : 53

Head with two transverse lin'es; pronotunl binotate, yellowish; scute1!1\um based biinotate; head, strfmum, abdomen w-ith ray-l'ike yellowish fasciae, acute, shorter.

Length: 7 mm.

16 REC. ZOOL. SURV. INDIA, OcC. PAPER No. 80

Genitalia: dasper (fig. 36) strongly geniculate, apical half baculiform" apex subtriangJUlJ.late.

Di~tributio~l: Andaman Islands, Mala,ya Archipelago, Philippines.

37. Brachyplatys subaeneus (Westwood) (Fig. 37)

1837. Platasp;s subaeneus Westwood, Hope Cat. Hem. I : 17

Shiny b]'ack; head wtith tVt'O bands; antennae piceous; pronatal disc with two spots sinlUarly an scuteIlar base; abdomen with brownisb ! ay-Hke conicaJ short fasciae, a series of submarginal spots.

Length: 7 mm.

Genitalia: Clasper (fig. 37) foot-shaped, proximally at point of curvature wiVh a l?unch of S\eta'~, rest of shaft placed at right angle, sagitaHty infolded, apex obliquely dissected.

Distribution: Andaman Islandsl, Malaya Archipelago, CeJebes.

Genus Coptosoma Dallas

1851. Coptosonla Dallas, List lIem. i : 61

38. Coptosoma contectum Montand (Fig. 38)

1893. Coptosoma contectUJll Montand, Allll. Soc. Ent. Belg. 37 : 567.

Head with a spot at base of central ~lobe;1 II, III, IV segments \vith a double series of linear ~~pots, rest with a sin~ ,series of spots, sternum, piceous; abdomen black~ shiny.

Length: 4 mm.

GenitalifI: Clasper (fig. 38) subapica!.Uy scimitar-9haped :Laterally salient s!l,ightly.

Distribution: N.E. India.

39. Coptosoma cribarium (Fabricius) (Fig. 39)

1798. C;nJex cariboriuln Fabricius~ Ent. Syst. Suppl. : 531.

Body greenish yellow, ventra:llly black, thickly punctate; behind 'each eye a.n obl,ique spot; pronoltum with a transverse series of punctures, anterio.r angle wi,th angulated striae; scutellum punctate; last abdominal s'egm,ent with a ray-like fascia intercepted by a dark line.

l~ngth: 5 mm.

DATTA, GHOSH & DHAR : Indian Pentato111oidea 17

. Gen.ita~ia: Cla'sp'e,r (fig. 39) slender, attenuate distad, apicdl 1/3 curved In form of a beak, point of curvature with m:inute setae.

Distribution: N :E. India, South India, Burma.

Renzark: The species is a common pest on Leguminosac and Compositae.

Genus Tarichea S til 1

1865. l'arichea Still, Ann. Sac. E11t. Fl'. V (4) : 163

40. Tarichea nitens (Dallas) (Fig. 40)

1851. l'iataspis nitens Dallas, List Hen1. I : 74

Body black, tinged with ol'ive green, median fascia extending to. abdomen; sternum pic:eous, head V'~ntral1y with a spot;. antenr,al apex black".

I.lenglh: 9 mm,.

Genitalia : Clasper (fig. 40) scinl,Har-shaped, sagittal groove distinct.

Distribution: N.E. India.

Family PBYLIOCEPHALIDAE

Genu,s Gonopsis Amyot & Serville

1843. GOllopsis Amyot & ServilIe, Ess. Hem, : 180

41. Gonopsis coccinea ( Walker) ( Fig. 41 )

186R. .Macrilla coccinea Walker, Cat. Het. III : 497

Body sanguinscens; ant~nnae apically black; head with anterior margin of pronotum punctate" black, posterior margin f'egulos1e, laterally pignllented upward, a transverse ridge between two projections, laterally serrate; sternuml densely punctate; abdom'en with fine long i­tud!nal and marginal series of black punctures; stigmata black.

Length : 16 mnl.

Genitalia : Clasper (fig. 41) geniculate mediad, apex blunt, strongly infolded sagittally.

Distribution : N .E. India, Burma', Malaya.

3

18 REe. ZOOL. SURV. INDIA, Oce. PAPER No. 80

Genus Tetroda Amyot & Serville

1843. Te'trvcia Amyot & Serville, Ess. Hem. : 177

42. Tetroda divaricata Dallas, (Fig. 42)

1851. Tetroda divaricata Dallas, List Hemp. 'I : 357

Thickly punctate; head, apte.nna1e, pronotum, corium laterob&saUy piceous; abdomen, legs punotate.

Length : 13 rom.

Genitalia : Oasper's (fig. 421) apical part foot .. sihaped, posterior margin rugulose.

Distribution Nepal.

Genus Fitha Walker

1867. Fitha ~alker, Cat. Het. i : 45.

43. Fitha ardens Walker (Fig. 43)

1867. Fitha ardens Walker, Cat. 'Het. I : 45.

Body elliptical, coars~ely punctate; -antennal firSit joint ~eddish,

rest black; pronotal disc and lateral margins hexanotate; ~uteUum uniformly colored,. abdomen on each side w~th purple stripes, an ~l1ow patch at base and apex ; legs red.

Length : 8 mm.

Genitalia: Paranlere (fig. 43) at basal half broader, sagittal groove distinct, genicula~e medially;, shaft denlsely wrjnIded at posterior half, ap'e.x gradually attenuated.

D4stributivn : All over India.

Genus Hotea Arnyot & Serville

1843. Hotea Amyot & Serville, Ess. Hem. : 41.

44. Hotea curcuJionoidea (Herrich-Schaffer) (Fig. 44)

1843. Pachycoris curuliolloidea Herrich-Schaffer, Wanz. Ins. III : 106" 313,1.

Body coarsely punctate; head an9 la1 eral pronotal angles piC'\!ous, pr~notum trifasciate; 5lcutellum with central longitudinal line extending to ~pical spots.

Length : 9 rom.

DATTA, GHOSH & IJHAR : Indian Pentatolnoidea 19

Genitalia : Param·ere (fig. 44) robust, with a flat projection mediad, subapically scimitar-shaped, apex feebly saNent, .5lubgeniculatc medially.

Distribution: N. E. India, Burma.

Genus Lamprocoris Stal

1864. LanlprOcol'is Stal, Hem. Afr. i : 34.

45. Lamprocoris spiniger (Dallas) (Fig. 45)

1849. Callida spiniger Dallas, Trans. Ent. Soc. V : 186.

Body dorsorventrally metallic green; inrrer margin of each eye and an:teri1or angle of pronotunl with a spot, with a fa'scia up to anterior Inargin, on each side, 3 obliq lie linear pots;,. scutellum basally penta­

. notate, posterior one crescentic, 3-4 spots unite,d to a pa1tch.

Length : 20 mm.

Genitalia : Paramere (fig. 45) strongly hamate, shaft i strongly geniculate.

Distribution Sikkim, Bhutan, Khasi, Naga Hills" Bumla.

FamiJ.y SCUTELLARIDAE

Genus Peocilocoris Dallas

1848. Poecilocol'is Dallas, Trans. £nt. Soc. V : 100.

46. Poecilocoris hardwickii (Westwood) (Fig. 46)

1837. Poecilocotis hardwickii (\Vestwood) (Tectocoris) in Hope Cat. i : 13.

1839. PlIC!zYCO";s nepalells;s Herrich-Schaffer, Wal1z~ IllS. IV : 1.

Dark reddish yellow'; pronotal bas'e binotate, black; scutellum with 11 spots~ black; antennae, ventral part violet-black.

Length : 20 m·m.

Genitalia : Paramere (fig. 46) hamate, ~ part flattened.

Distribution : N. E. India, BUflna.

47. Poecilocoris latus Dallas (Fig. 47)

1848. Poecilocoris latus Dallas, TrailS. El1t. Soc. V : 101.

Body thick:ly punctate, reddish yellow; anterior angl'e of pronotul1~ with a spot, base with two spots; scutellar base with a patch. basal

20 REC. ZOOL.-- SURV. INDIA, Oce. PAPER No. 80

ang!~ spotted., beyond middle tetra spotted , last abdomlinal segment \vith a spot; rostrum, legs vio.:~-black; coxae, antennae I & .lI seg­nientsl reddish yellow.

Length: 21 mm.

Genitalia: dasp~r (Fig. 47) with a bunch of setae just bellow the ventromedial tine robust, unciform, subapicaUy tapered.

Distribution: N.E. India, Burma.

Remark: It is a cOmmo.n rea seed bug~ The species also attacks flowers.

48. Poecnoco~s purpurascens (Westwood) (Fig. 48)

1837. Tectocoris purpurascel1s Westwood, Hope Cat. I : 14

Body v~olet color, sonie '!pecitlle~s with tinge of black pigmenta­tions; pronotum tetra spotted , apical margin bright red; body; ventrally shiny, brassy green,; abdom,in~lli seglnents 2-5 with red streaks; legs antennae, rostrum black.

Length: 18 mm.

Genitalia : Clasper (fig. 48) with shaft drawn out to a whip-like process, strongly curved upward, apex pointed.

Distribution: N.E. India, Naga1and.

Genus Scutellera Lamarck

1801. Scutellel'a Lamarck, Syst. : 293

49. Scutellera fasciata (Panzer) (Fig. 49)

1798. Cimex fascia/a Panzer, Voel. Col. IV : 108.

Body meta-Hie green,; inner margin of eaClh eye with a short fascia; pronotum trifasciate, central one continued to scutellum, 7 spots on each s:de of fascia along central axis of scutellum and pronotum; body ventrad, legS', rostrum reddish yellow, abdomen laterad fasciat~; femora tibiae, tarsal apex indigo-blue.,

Length: 20 m'm.

Genitalia: C!asper (fig. 49) robus,t, apex clavate, inner margin curved.

Dl~tribulioll N.E. India, Burma, Malaya Ar.chipelago.

DATTA, GHOSH & DIHAR : Indian Pentalol1widea 21

50.. ScuteIIera nobilis (Fabricius) (Fig. 50)

1775. Cinex Ilobi/is Fabricius, Sysl. En I. : 697

Body m·~tal1ic green, without transverse fascia; scutell urn with a ~edian fascia, hexanotate on disc, spots placed in pairs.

Length : 19 mm.

Genitalia : Oasper (fig. 50) cylincirate nl0r~, than half, bifid apex, chellfornl apicad, robust.

Distribution : .Sind, Karachi. D!ehra Dun, Assam, !vlanipur, 'frivandrum, Ceylon, Bombay, Calcutta.

Relnark: The species: is conlmon on Sandal Wood planta~Ion, Santalilln albunl, Phyllanthus elnblicQ, and alsO' on agricultural crops.

Genus Solenostethium Spinola

1837. Solenoslcli1iunl Spinola, 1'::ss. Henl. : 360

51. Solenostethium rubropunctaturn (Guerin) (Fig. 51)

1830. Scutellera rubropunctatum Guerin, Voy. Cog. Zool., 2(11) : 157.

Body chestnut brown; antenna'e, black!, II and I joints red, prono­tum heptanotate., reddish color; scutellum with ten reddish spots.

Length : 16 mnl.

Genitalia : CJasper (fig. 51) abruptly narrowed mediad, feebly serrate at middle, strongly rostriform.

Distribution : N. E. Ind':a, Sou'th East Asian countries.

F'amily TESSAROTOMATIDAE

Genus Empysurus Martin

1904. Enlpysu rus Martin, Bull. Mus. Paris., : 315.

52. Empysurus johni Oshanin (Fig. 52)

1907. EI11pysurus johni Oshanin, AnJl. Mus. Zool. Acad. lnlp. Sci. Soc. ST. Petersll. xii, : 416.

Ant'ennae yellowish brown; scuteJlLul1 at opaque part of base of membrane narrow, olive green; forewing nlenlbrane vitreous. uncolored, connexivium olive green, laterally bordered by two fine, black lines;

22 REe. ZOOL. SURV. INDIA, Oce. PAPER No. 80

vcntra.J1y yellowish-green, meta- and nlesosterna, sides of posterior and intermediate coxae brownish,· stigmata dark brown; femora yellowish­green, tibiae greenish.

Length : 26 mm.

Genitalia : CJasper (figo\ 52) robust, fissate into two arms, longer one twice shorter one, margin infoi!ded, apex rounddd, shorter arm obliquely truncate, nlargin with stiff. setae.

Dislrib'.l1ion : Ceylon.

Genus Eusthenes Laporte

1832. Ellsthenes Laporte, Ess. Hem. : 64.

53. Eusthenes hercules Stal (Fig. 53)

1870. Eusthellfs hercules Sial, Ent. Ifem. I, : 231.

Body piceous; coriuml chestnut brown; cOnD'exivium black; reddish brown spot at base of each segment; scutellar apex chestnut brown; legs black; tarsi dhestnut brown.

Length : 40 mm.

Genitalia : Clasper (fig. 53) robust, strongly divergent, ! densely setose on disc, latcropostteniorly with a cJ!jg\itiform projection.

Distribution : N. E. India.

54. Eusthenes robustrum Le Peletier & Serville (Fig. 54 ) 1825. Tessaratofna ,.obustuln LePeletier & Serville, Enc~ Meth. X : 591.

Body dark brown" thickly punctate; pranbtum late.rally wrinkled; forewing melnbrane bras'sy yellow·; tarsi chestnut brown; l'egs, antennae b~ack.

Length : 36 mm.

Genitalia : Clasper (fig. 54) laterally sharply salient, arched, l~osterior margin dens·ely setosc.

Distribution : N. E. India, Indon'esian. Islands.

DATI A, GHOSH & DHAR : Indian Pentatol1widea 23

55. Eusthenes saevus Stal (Fig. 55)

1863. Eusthenes sacl'US Stal, Trans. Ell I. Soc.) 1(3) : 597.

Body blackish with rusty brown re!Jections,; corium obscured by dark color, a.ntenna! apical joints, scutellum chestnut brown 2.S tarsal segments; hind femora with double series of obsol'e,te spines~

Length : 25 nlm.

Genitalia : Clasper (fig. 55) robust, dens'ely setO'sl~, apex rhe,Jj .. form, shape a'S; figured.

Distribution : N. E~ Indlia;, North China.

56. Eusthenes scutellaris (Herrich-Schaffer) (Fig. 56)

1839. Te.uaratcnla scutellaris Herrich-Schaffer, H' anz. [nst. IV : 81.

Body brown; pronotuml and scutelIunl stria.te corium punctate membrane brassy browny; po~t'erior fenlur wi:th a long spine at base: apical half with a doub!,1e series of :spiflJes O'n ventral abdonlinal 5.egm'ents.

Length 25 mm.

Genitalia : Clasper (fig. 56) at apical margin truncate, ch~;tlte­

shaped, chitiniscd ntesal margin deeply scu1iform.

Distribution : Nepal:, Khasi HiHs, ·Shillong.

Genus Pycanum Amyot & Serville

1843. pycanum Anlyot & Serville, Ess. Henl; : 171.

.57. Pycanum ponderosum Stal (Fig. 57)

1854. Pycanuln ponderosum Stal, Oft. Vet. -Ak. Fo,.l1. : 234.

Dull obscure c]ay-color; antennae, 'extreme lateral margin of head and pronotum, legs. piceousl; scuteMar ap~x, connex:iviul11 pale clay-color, with Jarge blacklish spots; forewing mem br.ane very pale hraSSIY brown;, pronotal shapes resemb!,e Mattiphus oblongus, anterior' angle quadrate.; P. ponderosum is sleparated from M.~ oblonRus as no central plate (e~~vated) present on meta'siternunl of ponderoslIll1.

Length : 32 mm.

24 REC. ZOOL. SURV. INDIA, OcC. PAPER No. 80

Genitalia : Clasper (g. 57) subrectangulate, robust, with circlet of 10ng sltiff setae, densely arr.anged along margins.

. .1.

Distri hution : Assam" Southern India, Burma, TenassarJ'm.

Genus Tessaratoma Le Peletier and Serville

1824. Tessaratolna Le Peletie:! and Serville~ Enc),cl .. Meth ... X, S90.

58. Tessaratoma nigripes Dallas (Fig. 58)

18'51. Tessaratoll1a lli~ripes D,a11as, List Hemp. I : 341.

Body brownish yellow" ventral1y chestnut brown; head margin bordered black; scutellar ap~x black; forewing membrane bronzy; ste,rnum with dahk Dlarkings; abdomen with distinct central ridge; transver:se striae closl~ to stigmatal spots; legs; rostrum', antennae da'rk ch'estnut brown .

. Length: 30 mm.

Qenitalia: Clasper (fig. 58) :i shaft broader, wV'el"gent abruptly drawn out to a robuSlt process arched inwardly, e!ongat1e.

Distribution: Sikkim" Malaber, S.E. Asian countries.

Family UROSTYLIDAE

Genus Urostylus Westwood

] 837. Urosly/us Westwood, Hope Cat., i ~ 45.

59. Urostylus gracilis Dallas (Fig. 5~)

1851. Urostylus grdcilis Dallas, Ust Hemp. I : 3-15.

Head 'vith~ut punctations; antennal II segmentll pronotum, scutellunl, corium green; body ventrad, legs brownish yellow, rostral apex black.

Length: 10 mm.

Genitalia: Clasper (fig~ 59) appearing scimitar-shaped, basalJy tumescent" pointed apex, dlscal part w'ith shor~ set ale.

Distribution: Sikkim, N.E~, India.

60. Urostylls punl;tigera Westwood (Fig. 60)

1837. Urostylis pUllctigera Wc~twood, Hope Cat. I : 4S.

DATTA, GHOSH & DHAR : Indi.an Pen tat 01110 idea 25

Body punctate; pronoturn binotate, sonletimes absent; rostral apex black, corium j1npunctate with ~~ck discal spots.

Length: 11 mm.

Genitalia: da;s.per (fig., 60) elongate, cy1indrate, sa~ittally in­folded, apex hamate.

Distribution: Sikkim, Kbasi H,ills.

R,emarks: Common p'ests on Magnoliaceae in Bengal, Assam. Highest incidence has been recorded from the foot hills of Darjeeling and Kurseong. T'b:e. nymphs suck sap from the terminal foliage and uppermost branches of the plants, which ultimately lead to vvithering of trees. The adults penetrate stylets into the shoots, main s.tems of young plants: and distintegrate the ti!ssuIes and d~eveloping oell~. The affected trees are further attacked by fungus nam'ely Fusarium sp. and Diplodea SP4

Genus Urochela Dallas

1850. Urochela Dallas, Trans Ent. Soc. I : 2.

61. Urochela quadrlpunctata Dallas (Fig. 61)

1850. Urochela quadrip1.lnctato Dallas, Trans. Ent. Soc. I : 3.

Brownish yellow; antennal segments black ,except lISt joint; pronotum with acentral line along with Jateral lines extJending to scutellum; corium each ~sjd~ with 2 spots; connexivium black; abdomen with a spot on each segment.. anotlrer one on mteral margin black; rostral apex black; f~ora with dusty patches.

Length: 9 mm.

Genitalia: Clasper dig. 61) stender, subapically :serrate, segittally infolded., convergent to a bllunt apex.

Distribution: N .E. Indi·a Burma.

SUMMARY'

The paper sets out to provide a comprehensive account on the genitallic a.rmatures and colour notes of si?Cty one ,~ecies belonging to forty four genera of the Superfamily Pentatomoi1dea4 The economic

4

26 REC. ZOOL. SURV. INDIA, Dec. PAPER No. 80

importance. of the species is mentioned wheI1elVer possi~.' Distribution of all the specileS is incorpolated. All the species are illustra:ted. Re~erences to the original literature are also given in the papett".

REFERENCES

AHMAD, I. and KAMALUDDIN, Sol 1982. A Revision of the genus Poecilocoris DaMa6 (Pe:ntatomoidea : ScutellJeridae) from Indopakistan subcontinent with descriptions of three n\eW species. Orient. Insects, 16(3) : 259'-295.

DISTANT, W. L. 1.902. Rhynchota-Heteroptrera. The Fauna of British Indla including C'eylon and Burma, 1 1-330. London (Taylor and Francis)

DISTANT, W L. 1908. Rhychota-Heteroptera, Ibid., 4 : 420-466.

1)ISTANT, W. L. 1918. Rhynchota-HomoptJera: Appendlix, Heterop­tera : Addenda, Ibid., 7 : -110-151.

LESTON, D. 1952. Notes on the Ethiopian Plentatomoideai (Heteroptera) VIII. Scutellerida:e. Leach of Angola, with remarkis QPon (he male genita1ia arul classification of the subfamilly. Publ. Cult. Cemp. dianl Angola, 16 :. 9-26.

LESTON, D. 1954. Notes on thfe Ethiopian 'P.~tatomoidea (Hetroptera) xx. Steganic.erus Mayr (ScutdJleridae) and. the structure of the Aedeagus. Proc. Roy. ent. Soc. Lond. (B), 23(9-10) .: 164-166.

M'cDoNALD, :F. J. D. 1966. Thle genitalia of North Amellican Penta­tomoidea (Hemiptera : Heteroptera). Q~t' Ent .. , 2 : 7-150.

RUCKES, H., 1963. Insects of Micronesia (Hieteroptera : Pentatomoid~) 7 (7) : 307-356.

SCHOUTEDEN, H. 1909. Catalogues raisOnnes de 1a fauna \:Ultomolo­gique du Congo. Be1ge, HJemipteres,. fame Pentatomidae. Ann. Mus. Congo BeIge, Sere 1(1) : 1-88.

Fig.

Fig.

Fig.

DATIA~ GHOS.H & OHAR . lndifAl1 Pcntqlomoiae(4

". -=,-.~ . , :. •• •

1.

2.

3.

, , • ,

ASOP:lS I1lQ!abar;cus

Audilletia spinidens

Fabr.

(Fabr.)

Picromerus obtusus Walker

E E ("} 2

27

CI~sper

t, ,.

Z8

Fig. 4. CydlJus indicus Westw.

Fig. SA. C. maurus (Dall.)

Fig. 58. C. maUrlU (Dalt.) Fig. 6. Macroscytus subaenus (DaIl.)

Clasper

P art of genitalia

Clasper

J,

DATTA, GHOSH & DHAR Indian PentatOmoidea 29

" \ \ ,~'~

l~j ,(", '~':: Iff l \ l I' It' '~:.f I I ~ \ \.\ " " ' , ,':,::, ' , ~\f\~"" It' t" ,'. 'i '1., t

, ~~" \, 1. \~, • It" ::: • '~ \ .. - '\ ) " t " .... ... .. ~ \ "I

"J... ".~-: ' \ ", " do '. '\'.'" ~ .

• :.::. &. ..t' I~, ~~,"T

~ . ",

Fig. 7. As ponga p'us lisslimensls Disi. Clasper

T~ig. 8. A. nigriventris Westw. " Fig. 9. Ate/ides centl'olineatus Dall. .,

3U

.Fig. 10. Fig. 12.

Fig. 15.

REC. ZOOL. SURV. INDIA, OCC. PAPER No. 80

.. \ .... . 'I" , .... \' , ,,~ , .·.ft . "" .. ., ., . "

eye/opella sicci/olia westw. A drisa Inagna (Uhler. )

Stibaropus callidus (Scb.)

Clasper

DATiA, GHOSH & DHAR Indian PentatOlnoidfa 31

~. -.' c).

Fig. 11. Megymenum parallelum Voll.

Pig. ItA. M. paral/eluln

l'lg. 13. Podops lurida (Bunna.)

F'lg. 14. P. b;~'P;nosa (Fabr.)

0 w 3

14 3

,. " " " ~

1\

f'I

Clasper

part of genitalia

Clasper

"

32 REC. ZOOL. SURV. INOlA, OCC. PAPER No. 80

t(ig. 16. Ag()l1oScelis itubi/a t=:abt. clasper Fig. 17. A Illestia al1chol'a (Thurb.)

" Fig. 18. A. cruciala (Fabr.)

tJ

F,g. "24. Cappaea tapro ban ens;s (Dal1.) ..

DATTA, GHOSH & DHAR

.. .... . . .. " .. , . .. : •. ... ' . . .. . .... : ::::.: ,. -. I' .• ... .. ..... :,: .. ....

Indiun PentatOmoidea

.:' " -.... . :" . .... : . ... . . " ..... ,

.,: . .•.. ::::. \' . ' ...

·f , .:: .' , , ::.

:.:.: t • . l' ':':: .' .'

Fig. 20. Stenozygum speciosuln (Dan.) Clasper

Fig. 21. Strachia crucigera liahn. "

Fig. 22. Zycrona caerulea (Linn.) "

Fig. 23. Canthecona furceJ/ata Wolf "

33

34' REC. lOOL. SU'tV, INOlA, OcC. PAPER No. 80

Fig. 19. Eusarco.coris c.apitatus Dist. Clasper

Fig. 25. DaJpada alfinil Dan. " Fig. 26. D. oculata (Fabr.)

" J;ig.27. Fernelius indlcus Djst.

"

DATI'A, GHOSH & DHAR : Indian Pentatomoidea 35

Fig. 28.. G ulielmus lateralis Dist.

Fig. 29. Hop/istodera virescens Dan

Fig. 30. Menida formosa (Westw.)

Fig. 31. M. histri 0 (Fabr.)

Clasper

" "

"

36 REC. ZOOL. SURV. INDIA, Oce. PAPER No. 80

rig. 32.

Fig. 33.

Fig. 34.

M. varipenllis (Westw.)

Nezara virid'lla (Linn.)

Priasslls exel1tptus (Walk.)

Clasper

"

DATTA, GHOSH & DHAR Indian Penl~/v!11oide~

,

Fig. 35. BrachypJatys punct;pes Montand. Claspff

Fig. 36. B. radians Voll. " Fig. 37. B. Sll baelleu.r.; (Westw.) ~ , Fig. 38. Coptosorl'la conteclunl Montd. 'J

3& REC. ZOOL. SURV. INDIA, OcC~ PAPER NQ, 80

0·3 mm ,

1.,2

Fig. 39. C. cri bnariunl (Fabr.) Clasper

Fig. 40. Tarichea nilells (DiaU.) " Fig. 41. GOllopsis coccinea (Walk) " Fig. 42. Tetro-do divaricata Dall. " Fig. 43. Fitha ardens Walk. u

DATTA, GHOSH & DHAR · Indian PentatOmoidea 39

Fig. 44. Hotea CUfculionoides (Herr .-Sch.) Cla~pcr

Fig. 45. Lomprocoris spiniger (DaB.) " Fig. 46. Poecilocoris hardwicki (Westw.) "

Pig,47. P. lallls D'a,l1. "

40

Fig. 48.

Fig. 49.

Fig. 50.

Fig. 51.

REC. ZOOL. SURV. INOlA, OcC. PAPER No. 80

o • w 3 3

P. purpurasceJlS (Westw.)

Scutellera /asciata (Panz.)

S. nobilis (Fabr.)

SO/clloslet/uHll ru bl'OpullctcllUlll (Guer.)

Clesp~r

" ,. ,.

DATTA, GHOSH & DHAA • Indian Pen/atomoidea

52

Pig. 52. Empysufus john; Oshan. Fig. 53. Eusthenes hercules Stal

Fig. 55. E. saevus StAI

53

Clasper

41

42 REC. ZOOL. SURV. INDIA, OcC. PAPER No. 80

56

l O~mm I 58

Pig. 54. E. robustus (Lep. & Serv.) Clf'.Bper Fig. 56. E. sclltelier;a (Herr.-Sch.)

" Fig. 57. Pycanunl pvndel"OSUnl Stat. "

Fig. 58. TesSaralol11Q nigripes DaU. u

DATTA~ GHOSH &, DHAR Indian Pelltatomoidea 43

59

61

l 0.3 mm

Fig. 59. Urostylus gracilis Dan.

Fig. 60. U. puncti.~era Westw.

GO

Fig. 61. Urochela quadripuflctata Dal'

~OXlrrQd . en

CI~tsper

"

"