18th 19th 20th 21st 22nd 23rd 24th 25th 26th 27th 28th 29th 30thPAPILIONIDAE SWALLOWTAILS

Swallowtails & CattleheartsPolydamas Swallowtail XGreen-celled Cattleheart X X X X XEmerald-patched Cattleheart X X X X X X X X XRuby-spotted Swallowtail X X XAndrogeus Swallowtail X XThoas Swallowtail X X X X X X X XWHITES & SULPHURS

Dismorphilnae Mimic-WhitesJethys Mimic-White X

Coliadinae Sulphurs & YellowsBarred Yellow X XWhite Yellow X X X X X

Guides: Tino Sanchez & Linda Harrison

Day 13 - August 30th; San Francisco Reserve

Day 9 - August 26th; El Salto Road am/Canopy Camp Road pmDay 10 - August 27th; Nuevo Vigia Village

Battus polydamasParides childrenae childrenaeParides sesostris tarquinius Heraclides anchisiadesHeraclides androgeusHeraclides thoas

Enantia jethys

Eurema dairaEurema albula

Papilioninae

PIERIDAE

Day 12 - August 29th; El Salto Trial

Butterflies of Canopy Camp DariénAugust 25th - August 30th, 2019

Panama's Brilliant Butterflies

Day 11 - August 28th; rain am/road to Lajas Blancas

Day 6 - August 23rd; Metropolitan Park am/Old Gamboa road pmDay 7 - August 24th; Pipeline Road & Canopy Tower

Day 8 - August 25th; Bayano Lake & Rio Torti am/Canopy Camp grounds pm

Panama's Brilliant Butterflies:

Butterflies of Canopy Camp Darién :

Day 2 - August 19th; Cerro Gaital am/Finca Macarena pm

Day 4 - August 21st; Altos del Maria

August 18th - August 25th, 2019

TOTAL SPECIES: 336

Lodge & Tower: 245Canopy Camp: 189

Blue=new species added Lodge & Tower: 4Gold=new species added Canopy Camp: 5

Day 1 - August 18th; Arrival, Canopy Lodge gardens/road to Canopy Adventure

Day 3 - August 20th; Valle Chiquito am/Cara Iguana pm

Day 5 - August 22nd; Las Minas & Canopy Adventure am/Canopy Tower pm

list compiled by Linda Harrison

18th 19th 20th 21st 22nd 23rd 24th 25th 26th 27th 28th 29th 30thTailed Orange X X X X

Pyrisitia v. venusta Pale Yellow X X X XMimosa Yellow XCloudless Sulphur X X XApricot Sulphur X X X X X XLarge Orange Sulphur X XStatira Sulphur X X

Aphrissa neleis Pink-spot Sulphur X X XWhitesGreat Southern White X X X

Itaballia demophile centralis Black-banded White X X X XViardi White XCattleheart White X XFrosted Dartwhite XGolden White XHAIRSTREAKS & BLUESHairstreaksWhite-tipped Cycadian XRegal Hairstreak X

Atlides bacis Bacis Hairstreak XImperial Arcas XMexican Arcas XSky-blue Hairstreak X X XPale-clubbed Hairstreak XPearly Hairstreak XTogarna Hairstreak X X X X XThales Hairstreak XEpopea Hairstreak X

Calycopis demonassa Rusty-banded Groundstreak XDusky-blue Groundstreak X X X X X X XTrebula GroundstreakGreat Scrub-Hairstreak XZiba Scrub-Hairstreak X X X X X X XRed-spotted Hairsteak XGray MinistreakMycon Hairstreak XBitias Hairstreak X X

Pyrisitia proterpia

Pierinae

LYCAENIDAE

Pyrisitia nisePhoebis sennae

Evenus regalis

Arcas imperialis Arcas cypria Pseudolycaena damo Theritas hemon Theritas theocritus

Pieriballia viardi noctipennisArchonias brassolis approximata

Phoebis argante Phoebis agaritheAphrissa s. statira

Ascia monuste

Panthiades bitias

Catasticta h. hegemonMelete polyhymnia florinda

Arawacus togranaOcaria thales

Calycopis isobeon Calycopis trebula

Strymon zibaTmolus echion echiolusMinistrymon azia

TheclinaeEumaeus godartii

Thestius epopea

Strymon gabatha

Theclopsis mycon

18th 19th 20th 21st 22nd 23rd 24th 25th 26th 27th 28th 29th 30thPhaleros Hairstreak XNarbal Hairstreak XCelmus Hairstreak XBluesHuntington's Blue X XMETALMARKSMetalmarksViolet-washed Eyed-Metalmark XPurlpe-washed Eyed-Metalmark X X

Leucochimona vestalis leucogaea Vestalis Metalmark XLeucochimona l. lepida Satry Metalmark X

Lagora Metalmark X X XTullius Metalmark X X X XBlue-winged Eurybia X X X X

Eurybia patrona persona Great Eurybia XBlind Eurybia XInca Metalmark X XBarne's Metalmark X X X X X X X X XFulmen Calephelis X

Calephelis schausi Schaus' Calephelis X XIris Calephelis X X X XCalephelis Metalmark XLampeto Metalmark X X XPixie X XElectron Metalmark XRed Metalmark X X

Symmachia p. probetor Red-costa Metalmark XSymmachia accusastrix Accused Metalmark XPhaenochitonia ingipicta Ingipicta Metalmark XAnteros allectus Rounded Anteros XAnteros chrysoprasta Elegant Anteros X

Black-bellied Anteros XAnteros k. kupris Giant Anteros X X

White-patched Emesis XWhite-spotted Emesis XDark Emesis XDotted Metalmark X

Celmia celmus

Emesis lucinda

Emesis ocypore aethaliaArgyrogrammana stilbe

Anteros formosus

Emesis lacrines

perophthalma tullis

Riodinianae

Eurybia e. elvinaAncyluris i. inca

Melanis pixieMelanis electronMesene mygdon

Caria mantinea lampeto

Leucochimona lagora

Eurybia lycisca

Mesosemia telegone Mesosemia lamachus

Detritivora barnesi Calephelis fulmen

Calephelis irisCalephelis sp.

Hemiargus huntingtoni hannoides

Panthiades phaleros

Polyommatinae

RIODINIDAE

Olynthus narbal

18th 19th 20th 21st 22nd 23rd 24th 25th 26th 27th 28th 29th 30thIrenia Metalmark X X X

Thisbe lycorias Banner Metalmark XThisbe ucubis Ucubis Metalmark X

Odites Metalmark XPhasma Metalmark XLucianus Metalmark X

Hypophylla s. sudias White-banded Metalmark XMantus Metalmark XAzanoides Metalmark X XCreamy Metalmark XBlood-spot Metalmark X XAdelina Metalmark XFalcate Theope XStraight-lined TheopeTheope sp. XBRUSHFOOTSSnoutsAmerican Snout XMonarchs & ClearwingsTiger Mimic-Queen X X X XMonarch X X X X X X XCream-spotted Tigerwing X X X X XLysimnia Tigerwing XMenapis Tigerwing X X X X X

Napeogenes tolosa amara Tolosa Tigerwing X XLycasteTigerwing XDiasia Clearwing X X X XHeraldica Clearwing X X

Ithomia iphianassa Iphinassa Clearwing X XPaula's Clearwing X X

Oleria zelica pagasa Zelica Clearwing XCeratinia tutia dorilla Tutia Clearwing X X

Hezia Clearwing XKlug's Clearwing XVariegated Clearwing X X

Heterosais giulia Giulia Clearwing XPseudoscada timnia pusio Timnia Clearwing X

Theope sp.

Calospila lucianus

Nymphidium mantus

Nymphidium ascolia ascolia

Theope phaeo

Nymphidium haematostictum

Nymphidium azaniodes occidentalis

Theope basilea

Juditha odites praeclarum Zelotaea p. phasma

DanainaeLycorea halia atergatis Danaus plexippusTithorea tarricina

Callithomia h. hezia Dircenna klugii

Hypothyris l. lycasteIthomia diasia hippocrenis

Oleria paula

NYMPHALIDAE

Thisbe irenea

LibytheinaeLibytheana carinenta

Mechanitis lysimnia macrinus Mechanitis menapis dariensis

Ithomia heraldica

Godyris zavaleta zygia

Catoyclotis adelina

18th 19th 20th 21st 22nd 23rd 24th 25th 26th 27th 28th 29th 30thLongwingsGulf Fritillary X XJuno Longwing X X XBanded Longwing XJulia X X X X X X XGreen Longwing X X XFine-lined Longwing X X X XZebra Longwing X XCydno Longwing X X XDoris Longwing X XCrimson-patched Longwing X X X X XCrimson-patched Longwing X X X X X XHecale Longwing X X X X XSapho Longwing X X XSara Longwing X X X X X X

Euptoieta hegesia meridiania Mexican Fritillary X X XSistersLeucas Sister XSpot-celled Sister X XCytherea Sister X X X X XErotia Sister X XIphiclus Sister X X XFundania Sister X XZina Sister X XEmperors

Doxocopa l. laure Silver Emperor XPavon Emperor X X XBanners, Crackers & Eighty-eightsEast-Mexican Banner X X XRayed Purplewing X XPale Bluewing XNorthern Ectima X X XRed Cracker X X X X X XGray Cracker X XVariable Cracker X X X X XStarry Cracker X X X XLittle Banner X X X X X X

Hamadryas amphinome mexicana

Hamadryas feroniaHamadryas laodamia Nica flavilla

Hamadryas februa

Heliconius sapho

Adelpha basiloides

Doxocopa pavon

Catonephele mexicana

Myscelia leucocyanaEctima erycinoides

Adelpha malea fundaniaAdelpha zina lacina

Biblidinae

Eunica malvina a;bida

Apaturinae

Heliconiinae

Limenitidnae

Heliconius charithonia Heliconius cydno chioneus Heliconius doris Heliconius erato petiveranaHeliconius erato hydara

Agraulis v. vaniliaeDione juno Dryadula phaetusa Dryas iulia Philaethria dido panamensisEueides aliphera

Heliconius sara

Adelpha barnesia leucas

Adelpha cythereaAdelpha erotiaAdelpha i. iphiclus

Heliconius hecale melicerta

18th 19th 20th 21st 22nd 23rd 24th 25th 26th 27th 28th 29th 30thOrange Banner X X X XDark Sailor X X

Haematera pyrame Blind Eighty-eight XSix-spotted Eighty-eight X XTwo-eyed Eighty-eight X X

Callicore tolima Blue-and-orange Eighty-eight X XDiaethria astala Astala Eighty-eight X

Widespread Eighty-eight X XDaggerwingsMany-banded Daggerwing X X XRuddy Daggerwing X

Marpesia zerynthia Waiter Daggerwing XBeauties & CrescentsOrion Cecropian X X X X XNew Beauty XDirce Beauty X X X X

Smyrna blomfildia Blomfild's Beauty XHypanartia lethe Orange Mapwing X

Red Peacock X XBanded Peacock X X X X X X X X X X X X XWhite Peacock X X X X X X X X XRusty-tipped Page XMalachite X X X X X X X X X X XTropical Buckeye X X X X X XCrimson-patch Checkerspot X

Chlosyne ezra Ezra Checkerspot XBordered Patch X X X X X X XOrange-patched Crescent X X X X XPale-banded Crescent X X XWhite-dotted Crescent X XClio Crescent X X X XVariable Crescent X X XWhitened Crescent X XBlack-bordered Crescent X X X X XLeafwings

Consul fabius Tiger-striped Leafwing XSiderone galanthis Red-striped Leafwing X X

Castilia ofellaEresia clioEresia ithomioides alsinaJanatella leucodesma Tegosa anieta anieta Charaxinae

Chlosyne janais

Chlosyne laciniaAnthanassa d. drusilla Anthanassa tulcis

Anartia jatrophae Siproeta epaphus Siproeta stelenes Junonia evarete

Marpesia petreus

Historis odius dious

Colobura dirce

Anartia a. amatheaAnartia fatima

Marpesia chironCyrestinae

Nymphalinae

Colobura annulata

Temenis laohtoeDynamine paulina

Callicore lycaCallicore pitheas

Diaethria clymena

18th 19th 20th 21st 22nd 23rd 24th 25th 26th 27th 28th 29th 30thIsidora Leafwing XPointed Leafwing X

Fountainea glycerium Angeled Leafwing XMottled Leafwing X X X

Memphis oenomais Edge Leafwing XArchaeoprepona amphimachus White-spotted Prepona X

One-spotted Prepona X X X XTwo-spotted Prepona XMorphos, Owls & Satyrs Cypris Morpho X XCommon Morpho X X X X X X X X XMenelaus Morpho X X X X XStub-tailed Morpho X X XDark Owl-Butterfly XPale Owl-Butterfly X X X XSplit-spotted Owl-Butterfly X X

Opsiphanes cassiae castaneus Cassia's Owl-Butterfly XOpsiphanes quiteria Scalloped Owl-Butterfly X X X X

Narrow-banded Owl-Butterfly XMoon Satyr XPlain Satyr X X X X X X XGold-stained Satyr X XTawny-cornered Satyr XJesia Satyr X XHermes Satyr X X X X X X X X X X X X

Magneuptychia gomezi Gomez' Satry X XBlue-gray Satyr X X XTiessa Satyr XWhite-banded Satyr X XWhite Satyr X X X X X X XYellow-spiked Satyr XKerea Satyr X X X X XThamyra Satyr X X X XRenata Satyr X X X XSKIPPERSLongtails, Flashers, Scarlet-eyesCommon Phanus X X X X

Euptychia hilara

Caligo telamonius menusEryphanis lycomedon

Opsiphanes tamarindiPierella luna luna

Morpho menelaus amathonte Morpho theseus Caligo brasiliensis

Archaeoprepona demophon Archaeoprepona demophoon

Morpho cyprisSatyrinae

Cissia pompilia Cissia pseudoconfusa

Fountainea eurypyle

Memphis arginussa eubaena

Morpho helenor peleides

Zaretis isidora

Hermeuptychia hermes

Magneuptychia libye Magneuptychia tiessaPareuptychia metaleucaPareuptychia ocirrhoe

Eudaminae

Taygetis thamyra Yphthimoides renataHESPERIIDAE

Euptychia jesia

Taygetis kerea

Phanus marshalli

Posttaygetis penelea

18th 19th 20th 21st 22nd 23rd 24th 25th 26th 27th 28th 29th 30thAugiades crinisus Crinisus Skipper X

Dimorphic Skipper X X X X XBroken Silverdrop X XBlurry-striped Longtail X XEight-spotted Longtail XEsmeralda Longtail X X X XDorantes Longtail X X X X X X X X X XTanna Longtail XPlain Longtail XBrown Longtail X X X

Urbanus doryssus White-tailed Longtail X XGreat White Longtail X X X X

Astrapres talus Green Flasher X X XTwo-barred Flasher XFrosted Flasher XGilbert's Flasher X X X X

Astraptes creteus Whitened Flasher XYellow-tipped Flasher X X X

Narcosius nazaraeus Nazareus Flasher XNarcosius parisi helen Narcosius Flasher XCalliades zeutus Zeutes Banded-Skipper X

Broad Banded-Skipper X X XSpike Banded-Skipper X XSharp-banded Skipper X X X X X XFalcate Skipper X XMimosa Skipper X X X

Bungalotis astylos Dark-cheeked Scarlet-eye XDyscophellus p. phraxanor Big-spotted Scarlet-eye X X

Fiery Scarlet-eye XDyscophellus ramon Plain Scarlet-eye X

Spreadwing SkippersMyscelus assricus michaeli Fiery Myscelus XPyrrhopyge zenodorus Red-headed Firetip X

Evan's Firetip XMaculosa Skipper X

Elbella scylla Scylla Skipper XMysoria barcastus venezuelae Barcastus Firetip X

Cogia calchas

Pyrrhopyge evansi

Astraptes anaphus

Autochton neis Autochton longipennis

Spathilepia clonius

Pyrginae

Dyscophellus p. porcius

Hyalothyrus neleus pemphigargyra

Urbanus simplicius Urbanus procne

Urbanus chalco

Astraptes fulgerator Astraptes alardus latia

Urbanus esmeraldus Urbanus dorantes Urbanus tanna

Polythrix octomaculata

Melanopyge maculosa

Epargyreus exadeusChioides catillus albius

Astraptes alector

Autochton zarex

18th 19th 20th 21st 22nd 23rd 24th 25th 26th 27th 28th 29th 30thEligius Flat X XPurplish Tufted-Skipper X XImmaculate Tufted-Skipper X

Pellicia arina Glazed Tufted-Skipper XCopper-headed Sootywing XGolden-snouted Sootywing X X XCentral American Sootywing X XVariegated Skipper X X XBlue-studded Skipper XPale Mylon XYellow-patched Bent-Skipper X X X

Ebrietas evanidus Blurred Bent-Skipper XHelias godmani Godman's Bent-Skipper X

Southern Sicklewing XPale Sicklewing XPanna Skipper X X X X

Zera phila Phila Skipper XCommon Blue-Skipper X X XStriped Blue-Skipper XStigma Skipper XGlorious Blue-Skipper X XRainforest Hoary-Skipper XGlassy-winged Skipper XCommon Spurwing X X X X X X

Timochreon s. satyrus Satyr Skipper XAnisochoria bacchus Northern Snout-Skipper X

Tropical Checkered-Skipper X X X X X X X X X XRuby-eyes & Grass SkippersGreen-backed Ruby-eye X X X XComplanula Skipper X X XSergestus Ruby-eye X X XAlternate Ruby-eye XWhite-edged Ruby-eye X

Carystina aurifer Golden-spotted Ruby-eye XViolet-washed Skipper XImmaculate Ruby-eye X X X XCathaea Ruby-eye X

Pachyneuria licisca

Carystoides cathaea

Ebrietas osyris

Quadrus cerialis

Paramimus stigmaPaches loxusCarrhenes calidius

Eantis thraso

Ouleus pannaAchlyodes pallida

Quadrus c. contubernalis

Talides sergestus

Cobalus virbius

Damas clavusDamas immacula

Antigonus erosus

Pyrgus oileus

Perichares adela

Talides alternata

Mylon pelopidas

Celaenorrhinus e. eligius

Bolla cupreiceps

Staphylus ascalaphusGorgythion begga

Nisoniades rubescens

Staphylus vulgata

Sostrata nordica

Hesperiinae

Xenophanes tryxus

Turesis complanula

18th 19th 20th 21st 22nd 23rd 24th 25th 26th 27th 28th 29th 30thCarystoides l. lebbaeus Lebbaeus Ruby-eye X X

Brazilian Skipper X X X XPanoquina o. ocola Ocola Skipper X X

Purple-washed Skipper XZenis minos Minos Skipper X X

Perching Saliana X X X XMorsa Saliana XViolet-tipped Saliana XSullied Saliana X

Aides dysoni Dyson's Silverpatch X XAroma Skipper XWhitened Faceted-Skipper X

Synapte silius Rain-forest Faceted-Skipper X X XTrailside Skipper XHermione Skipper X

Lento xanthina Xanthina Skipper XCorades Skipper XJuventus Skipper XRadiant Skipper X X X X X X

Callimormus saturnus Saturnus Skipper X XBlack-spot Remella XRita's Remella XPurple-shaded Skipper X X

Methion melas Rusty Skipper XIttona Skipper XTeleata Skipper XJulia's Skipper XAlumna Skipper XFawn-spotted Skipper XVetula Skipper XEufala Skipper X

Papias dictys Bottom-spotted Skipper XLerema liris Liris Skipper X X X

Half-tailed Skipper XNiconiades xanthaphes Stub-tailed Skipper XNiconiades nikko Nikko Skipper XVettius triangularis Tringular Skipper X

Remella remus Remella rita

Cymaenes alumna

Vehilius vetulaLerodea eufala

Nastra julia

Lento hermione

Callimormus radiola

Saliana esperi

Saliana saladin

Synapte lutulenta

Anthoptus epictetus

Calpodes ethlius

Panoquina lucas

Aroma aroma

Callimormus coradesCallimormus juventus

Niconiades incomptus

Saliana salius

Saliana morsa

Eutocus maltidae

Mnestheus ittonaMonca teleata

Cymaenes trebius

18th 19th 20th 21st 22nd 23rd 24th 25th 26th 27th 28th 29th 30thDiversa Skipper X

Justinia justinianus Justinianus Skipper XJustinia norda False Saliana X

Compliant Skipper XImmaculata Skipper X X XOsca Skipper X XFiery Skipper XSouthern Broken-Dash XPompeius Skipper X X X X X

Poanes zabulon Zabulon Skipper XAnthrytone mella Mella Skipper X

Orange-headed Metron XCanna Skipper X X

Cynea cynea Cynea Skipper XNyctelius nyctelius Violet-banded Skipper X

Metron chrysogastra Quinta cannae

Hylephila phyleusWallengrenia othoPompeius pompeius

Vettius diversa

Eutychide complanaEnosis immaculataRhinthon osca

Satyr Skipper Rainforest Hoary-Skipper

Red CrackerSilver Emperor

Scylla Skipper

Orion Cecropian

Blue-winged Eurybia

Red CrackerSilver Emperor

Broken Silverdrop

Isidora Leafwing

Two-eyed Eighty-eight

Ezra Checkerspot