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. 92 lJ RESISTENCIA A FUNGICIDAS. Jos~ A. Salas F'h.D'~ Contenido :1:) j::;:E:'s:.i!::;1:.E:!riC.i.,::\ (::If::~ :l(::)!.:;:. ~)c:\-!:'C)(",:.I£:~rlc~1!:::, 0:\r1tE~ :i.(::)s (:::C:)fTlr)L.lE?~::,.t:o!::; (:~1.1J..mj.c:(::)s. [] \ '-! ) "-'. t 'n"" J ;:' "-""""" 1. ' ':::i-"-::' I "r" l '. -, f\.II -' C ' 1. ' C ' 1. " j "'",:. '::: 1. .:::i-'!:.' li1.( '- C ,":: ..r r :\""..::>.-,...,...I ~.", .I.;..I=._.~..~_.~_I T ]I i '-:"::""':::' 1 "-:,,, ('-1' 1 '\1::: r ! '::' 1( ' ) ,:::, r' r " c -",' l C""" I"""': cj ':" r "'::"':::' l ":::+'I=:" " iC ' 1. '- - 't " l fJ -1 ( "I l ' C.1' d ;,"":: - , r.",_",.!!.",_..,,1 -,"" ,,--,..! .!1 ,."ld,~ ",' "',-",_.,,-, ,d d "",::1' ,"'~. ' [ '_] ) 1 ".""" (-"-:' 1 -' 1. ' "::"" c,,::: d "'" """""'-' 1. ',:::' t (,::> r ,j- 1. "" ;:::" - i ! "' or - q( -"':: \J ' I\ r,r j ;-' (je '- C ' C - l'.L' J '" C'(:::, ,. 1",._",1",..1,_,_, "., ."--,,,_._,,__,d ,- ,I""J'~ll.,__.,J - (::(,,\_1,1._, -f I_I r-ll:J i, r~ :.1. (j 0:'5 . ' j) - "' l 'r' )( '"". (j .-:. r"'-::,c' 1. ""'i"c- r-,""j'" Y "" E, ] c" C - C - J ' ( "' r" , -',.' .,::, .:, .::, ,,:, .",_":,c!.L,,...,d """,..::!_"",J,. ' jl i 1::'c-c" 1. ,_.,' t""" - i( - 1. "" ('- r-ll '-"- (' j '" " r""",::>,=,=,-",:,,-,.._':\.,"'" "' II i "'. O'1 'l""r1 r ":::. c ' Y"':"""" "-'-::"':: J "":' t "':o r' c -1. "" '" fll" ' )C "' l' C' J ' Cj :::,,:: V",II,.L"_""-,.'!"-'\...""",...,_,,,,,,I ,,-,= ';:1"",1_.. I,jl:[ 1) 1'1~'t(::)d(~)!:~ E?stab1e(:.1.dl'Js par-..":\. E~\...alL.lar" t-e!'5j_s.ten(:,1.,::,\. IX \ 11 "'r' ) """" (- -, .::: OC" J ' O E :' C( ' )r " o """"' j ' c'r" (j 'o, J -, ,,-""c" 1. ""' t c:o r ' iC - 1. '-' '" . fLlrlc "' 1. ' c - 1. ' cl -',::: ".Ilrc:'L"\,,,,1 "."",_"1"""_,1.:"",-""",:"",,,,:,, "-""", """",:,-". ~ X ) Or-~:Jo:'n :,L ,:,:'\(: ,1. t:1r'ies 1 'I't(=f'-ria(~~ i(:J'I,,-" 1 e!:,; r-e 1 a(: ,1.0f1adas COil r"(::;'3.i- 5 'tE~'IC ,1.a. Xl i E :-'j '"o, "' il "' lc "'" cj '" r""""'- 1. """"'co, nc "' l '", - \ 'f Ll" 1qJ ' C " l' C " "'-' "" "" ( ~ c ""'-' t -, R -' J ' c ':::. ",.,-"-:,.,.!,,.::,.:: ..::!":',_"L"::,,,,,,'::\'::' -,""""',~":_I,",J_",":' ,,"". ,1, . S ,i (J ,:,:\ 't 0 v:, -"'-" I-I (= (~J r- a e I-I I:J a r1 a '-I t') . :,';:' . {:~n t:,F' i::'\c rlo~;i <;::,; en pa ~)aya . ::::;. l-,i ,~(~jr1 tar-d ,1,0 f?n papa. "t. E~o't:,J~yt,1.S f'":'rl fF'es,-,,'s. XII) 1_,:.1.'t:er-c:~'t:,I_IF'a ct~)r1!.:;;ul'tada. I) RESISTENCIA DE LOS PATOGENOS ANTE LOS COMPUESTOS QUIMICOS. DE:: mar1f?r-"-,, an..:f\lt')~;~t3 a (:OITIO ap;';:lr-e(:,i(':?Y-OII er1 j:.:J C) (: C) t,iempo :lc:ls !:Ja't('j(;Jet-IOS C 'I I=" ( "" c ') 'n"" r-(::: """:C:: l ..::: t '= r """'-t:-' '" Jc "" "' r " tl ' bl ' (".tl ' c " C """" 'I C)"" J ' r """ e " c --I' C ') "" Y ';' c ":::. r"I' ) o" "::,c!.'f"I,.!"-'",-:!-:',._.,c',L..::!_::,o,.!,,,,,::,1""",J""".!_"y"":"'"I.",:,--"...;, O:',-Q_.:::' r -pt- 1. '"." t ' e -, """"'::..::: ~":::. l n ll" " C)"" 1. " ' 1 "" e( "" t1."'- 1. ' (j ::' s \/ -r""..-, ! ' C'i (4-c- ,'-,,'" ha, '" ' f( "' r -In "' c " o ---"'_",_f\_t_,_, ==I:.1-"'~' ""__.",L,,, '-' 1(::(",.",.,-",."..",1_", -""'" ,f -' ,,-,j t " r r l -' l' E " r) 'I-"" J """" C '::.r"" \s ' f : J ' t ' c r,""f- (" )c ' , =na "" C ' Ll C' C' C), '" r """""' l """""""' r)t ' e -"" -, -' lqlll -' ,-,-, ,,--,-,_._, :L F -:" l "' c) r,-"""" J --c 'J "" r::"'_"'- J c"' fe I'ic-r""' r" t -' n ' er 'i ' t' c' cje F " l . tc ' p "" tc ' )Jo " c"'" ( ~I- )l ' C ' l 'l ' ta ' [r ' i ' t ""' r " ( ,-'":,~:,:~~'~:I,~'L.I,L.!,:,~,.!:,, r~,"':" "':.~i""~ .~~:~~:I"):C:'~~,-":.!~,,,.." ,-,~'~':",_',':~:!""'~;..~~~,- .,,-, , , ""c!- \ ) 11<::lt,l,C_11,:\.l. f~c"tL.I..,\.I,mEr, -,E, f1_"E,,_,LF ~(",t{{:11_, ((.t',l.C1LI1.t"lI CcJ_,,!..a f,J,c_",,). --,1.,-

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Page 1: lJ 92 RESISTENCIA A FUNGICIDAS. - mag.go.cr · 11- '1,::;:_,,11 '" --"\0 -";',,--,c 1 liP r' ~"

.

92lJ RESISTENCIA A FUNGICIDAS.

Jos~ A. Salas F'h.D'~

Contenido

:1:) j::;:E:'s:.i!::;1:.E:!riC.i.,::\ (::If::~ :l(::)!.:;:. ~)c:\-!:'C)(",:.I£:~rlc~1!:::, 0:\r1tE~ :i.(::)s (:::C:)fTlr)L.lE?~::,.t:o!::; (:~1.1J..mj.c:(::)s.

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IX \ 11"'r' ) """" (- -, .:::OC" J' O E:'C( ')r " o""""' j' c'r" (j 'o, J -, ,,-""c" 1.""' t c:or'iC - 1.'-' '" .fLlrlc "' 1.' c- 1.' cl -',:::".Ilrc:'L"\,,,,1 "."",_"1"""_,1.:"",-""",:"",,,,:,, "-""", """",:,-".

~ X ) Or-~:Jo:'n :,L ,:,:'\(: ,1. t:1r'i es 1 'I't(=f'-ria(~~ i(:J'I,,-" 1 e!:,; r-e 1 a(: ,1.0f1adas COil r"(::;'3.i- 5 'tE~'IC ,1.a.

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XII) 1_,:.1.'t:er-c:~'t:,I_IF'a ct~)r1!.:;;ul'tada.

I) RESISTENCIA DE LOS PATOGENOS ANTE LOS COMPUESTOS QUIMICOS.

DE:: mar1f?r-"-,, an..:f\lt')~;~t3 a (:OITIO ap;';:lr-e(:,i(':?Y-OII er1 j:.:J C) (: C) t,iempo :lc:ls !:Ja't('j(;Jet-IOS

C'I I=" ( "" c') 'n"" r-(::: """:C::l..::: t '= r"""'-t:-' '" Jc "" "' r"

tl ' bl ' (".tl ' c" C"""" 'I C)"" J' r""" e" c--I' C') "" Y ';' c":::. r"I' ) o""::,c!.'f"I,.!"-'",-:!-:',._.,c',L..::!_::,o,.!,,,,,::,1""",J""".!_"y"":"'"I.",:,--"...;, O:',-Q_.:::'

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Page 2: lJ 92 RESISTENCIA A FUNGICIDAS. - mag.go.cr · 11- '1,::;:_,,11 '" --"\0 -";',,--,c 1 liP r' ~"

fLlngl.cidas y qLlt:!~ poY"' 10 tanto~ no se yen .::'\fecti:"!dos pOl'"" e:Llos. :JDLlr"::lntli::"! mLlc:l-iOS anos~ CLlalli.:j(:) ="L')10 se LI"l:..iliz~'\r..of1 pl'""otet:.tor-E?S como f.?l

th,J ""'-' m i"1 -' r-'..' I, C"! C ... p. t .' r '. r "' o ="" ()t ..= er ."J-""' on ""=:'."'- r..""""~c r . e."" J.'.- t ".' r . t "".. .IC:\~ ,C:,IO:::'._!1 ,::I.C:lI!1'.=;O"- J=;o..":\1 ,:""""..::._,:..,.!,:.,,:,", .._,..:",=.I""S!1

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-' p ' r "'. ( ... J.' c" Ilr ...' ( .."-r i -' C.I "'. V "" I" it . Llr .. 1...' .i r ...~":.iprlll""'-.= C. LI "' ]1 '= ( I -' C .- Ll Co.-' ..I e 1 .. .' ) ~. j ." 1,::1 ",I,::! _. J .. 1 c:, .. o::."! t. ,,:, . "'! -_!:~~_..:..:.E1. ",,' "::\ "" ".."",~:_:..:._~. \ r_"\ c\ .-, co"\ CJ ..::\ r r. r\ aCE.

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compuE~=..t:c::1 (:~LI:i..rfl.1.c:C:) ~;,f,~ tCil'""I16 inef.1.c:.i.erite ~).::Ir.a cor1tr.olar. a ese horigo

f:?r"l LIlIa ampl.i.::'\ l'""e<;.~i6rl" Desde hacl~ var-i(:)s arIas se sal:)e r.:jLle las

c:e~:ir:"!s:, (:JE" E~r..:.t~..~:.rl.ifi!. 9l!l.'iJ.S:.~~}.r.~~ 1 a bac: ter. i cl cjf!-:.1 t:. i zt')n de i' Uf:.~go, sari

I'""esi,,:;.terites al antibit~tico sistt~mico estl'""eptomicif1a".

r':'r.~\c:tl.c:amf3!nt:f3! t:.odc)s 10s fLlrlgicidcls Llsadc:!s pat-a el coritr.'o]. cje

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1 imi.tados c.-:r. Llf1a al:-.ci6n (je PI"'0.tecc.i6Ii el-1 la SLlpel'""ficie"

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II) HISTORIA RESISTENCIA A FUNGICIDAS SISTEMICOS

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III) PERSPECTIVA DE LOS PROBLEMAS DE RESISTENCIA A FUNGICIDAS.

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~ .1. MINlt'11 ZAR LA F'RESION DE SELECCIOI\! Ei"'J EL F'ATOGENO F'OR EL

V

a) '-..Iso ft_tr\i;}ic::id.:-'ls c~or\ diferentes mc:.lclc_IS de acc:,i6n, b) F'Ltrl~;licidaa(:t,)iTlpanan'te en 1,~ mezcl('~, c) Est,able(:er- r,~o. aplicac,iones/;ano,d) t'1 c:, nit, C) r" f:'~ c:, (j Ii:? 1 1=) a 't, 6 9 to:! r\ 0 .

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atr-ibLt,irjas a Ltr1(:) 0 mcfl!-3 de l(:)s s,igLtientes mf?l.:al'"i,ismos:

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a r- ,}'r'i j'"'-' """"",.,-,,-JI.

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proc:ILIC:"t:Os qLle "t:.1.E'rlen Llrl !:;:,olo sj.tici de ac:c:icln (SIS1'.EI"1ICO~3) y erl

(,;?st(Js c:asos c:Llalquiey-a de los 6 ar-,tey-.ioy-I""?s meCallo1.SmOs, pLleder-,

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[:::N El... ~3 I (:7jlJ J: E/\JTE (:~lJI:)IDF:O SE~ RESIJME El... 1"10DO Df:~ ACC: I 01\1 DE L.OS..::" '._.0' ' ' ' '-I.)IF -EI::;,r::-hl..rE.-r... F" J"'I~. [( .~ Il .-).'.7'c~ c-°I ".. '..E.;-MIC .'OC:..: ": l.~::_!:~ ::::::2 _~..l~- ~..:..-=- !~~::!._~?_..:.::.;:!_. --=- _--o:!--==- .

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V) TIPOS DE RESISTENCIA V SELECCION

Hay j.rll"'c.-Ir.rrlac:i.cir-, CILlf:."'! .1.r1cijoC:c\ C~LI~::: e:o:i.~"""t:er1, a un frILlY bajc.\ porc:entaj~:::~

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I.) I) RESISTENCIA CRUZADA

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eft:"?c t,1.'Y1,cla(j cle ot~c!s fL\ng1,cirjc:'Is ria sidc-1 afectada.

E:t 't~r-liiino ~esis'tencia cF'LIZa(ja se 1_lsa solo en (:,;.iSOS!I cI.\ando el

pat,t~q£,::rlci es trlenos susce~1.t1,t:)lf::~ a ],C)S dos C~LI.1.m1,COS que la F'aza

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,1 . HI I::JF:OCAF:BC:Jr-JOS ARC)t'1A T I (:[15 (CL_ORONEB - B I FEN I LO - D I (:I I:~JRAN - j--" -",-",-, ,_. ,,-,-, ,-,- ,---,._-,_._, ,---

HEXA(::I_OF:OE!Er-JCEr-JO). H'"::IY e_jemplos (jf!!! t-f:?s,istenc:i..:::! Cr-LI;;~,:\(:ja en'tr-~:: Vlc) C'. C-( ')m r c)r""" Y" t ' E'1::: C..,., ""c., t "" Clr .l lr )c " r' cle'r.'.Lc. "" e h "" "..e ri c.,..-'t "" do-" 1,,":,,'._-" ...::: c_,:,,-,'! ::,".r!' "'"III"'!_" -,,- co. I.,...JI,""

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":ii,

VII) MONITORED PARA RESISTENCIA A FUNBICIDAS

LEI prop6~s.i to (je es.te es e 1 de eva lLlar en e 1 CatT1pO 1 a sensibi 1 idad

tiE']. (::)r.~lar-1 i::,mr~) f:"ri e::, tLld ic:) c:l UriC) 0 var..j.os fLlrigic j.d~'t'::; paF.a dec idi r

10s prlJgF.amr:'lS irl.t:'f:-:!(jr-ad(:)s (:1f:~ c~ontro1"

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cje F.if:?!5~4(3" Ilrlp l/::'?tTlen tar- estra-t.eg ia.

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Page 10: lJ 92 RESISTENCIA A FUNGICIDAS. - mag.go.cr · 11- '1,::;:_,,11 '" --"\0 -";',,--,c 1 liP r' ~"

-, .,

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;,VSi se detecta r1c:)r\itoy"eo par-a I.ieter'- Revis:,at-' Y.'E'!C:t:..1.ficar.r-esistenc :i.a 0 (ninaI'"' e>: tens.ion .1 sig- e..:;.tr-ateg .ias.

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r" f= S :i- !3 .t: e r-1 -I.:. f= '3 .

SLt t)se(: Ller-1 .temen te I"1(:)fl .i- -1.:1;3 Y-f=C) (j E' ':'1\.1 ,"::ir! (: E? (::) E:::\I a 1 \_1,3. r I.: c!mpoY- .ta'-Y-e"' jL '-'.'-'.; c'.,r- cj'-:" "" e ,::;,;c.. t' er "- /." e- r""" c'.,": c,, = 1 ".'('..,r-. ty -c'.,]c,L.c I.'-.I ' II-,...IL._J.J"c , 1._.

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(;J :1. ct'::. .VIII) METODOS ESTABLECIDOS PARA EVALUAR RESISTENCIA

La FA() t-a. ".",,::..-:t.) " -':, c- 1.' dc . /j.16 tC " C" ( '.J "" f"-::""'-:.' \1:"' )'-' J. ( "":::, 1..J=,.t.,.i.,rlc::' r-I( -Je::i c' Ll1 "L" J'\.JI-"'~I "-.-,\..,,.\.~..- . c.JJ"""r"""';""'" ~,.L.",; ". 1. FAO. 1982. F~ec:(:)/!\irl~?r-f(:jf={j rn~?-I::t-I(:)(js for- .the (je-l:(;?(::: t:i.(:)r\ arid

measLlr'emE"!!r\t c:,f r-E?s:,is.tar\l.:e oi: ci(;)Y.':i.(:::L\li:ul~c:\l ~:)~:!!~:,.t:. .to F'E:'!::,tic:::i.c:!~:::=.. (4)

dolode Sf:? pr-eselo.ti:3.r-{ /rle.t(:)d(:)s par-a e\I,:i 1 \_\ay" Y..i::?s.is-l::.eloc.ia (je: ~~-~~.!::!.t_!::~..j,..~

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i n f e s .t a 10 '::5 .. F' 1 a E-/TlQP~_r'::.t~. ~..~:.-t;.i.!;';'!:.J_L~ ~ E~:Lr:::.:1_1;;.~::!=-1J:.?:.r_~~:.ii g_.t::.Y...:!;..!:~~. , ,3. v .:'-1 r- .i (:) s--. - J

fLlngj.c .idas:,.

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DUF'ON-r. Sigato~;.cil r-l~':!<;;)y"a y f~irnar.j_11c\. Tecr\il.:as. /"ne~jc.)Y.'acjas ~JcilY."::'I

manejo e ideri1:i'fica,=:i.6rl. 17 p.

IX) IMPACTO SOCIOECONOMICO DE LA R5:SISTENCIB A FUNGICIDAS

QLtierl 'Vi:'. a St"?r- a Fet:tad(:)':'

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(je camlJ.i'3r- a o.tr-o i:::t_\lti\/I::J, (:1eJ. q\_IE"!! !s,31:)e mf"~nos'1 p(jdY.'.i.a ser'. mt-:"?r-I(:)S

r-er\tc:{t:i1~~. c::) -rf:."'~r\et-. C:ILtf::: U!::;c\Y' c.ltr'oS fL\r\~Jj.cidas r)CI~.;jt::,lemli::~rl-l:e mo',ls

car-os.

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Page 11: lJ 92 RESISTENCIA A FUNGICIDAS. - mag.go.cr · 11- '1,::;:_,,11 '" --"\0 -";',,--,c 1 liP r' ~"

di5ponibilidad del pr<3dLlcto (s). c) Menor calidad del prodL,cto

L"(~:;) .

:~;E::.j f::..;:\..DC'J.C.'i"' j,'..r.E:.

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a) F'o~:;,iblE'!s de~mar\dc:ls de lc:)s afectados. b) Mejorar' asistencia

t~I::lj:ic:c~. (:) Df.:::sar-rol1c.ir m~i:odo (5) l'mCII'lit,ol'-eoll Resist£~I'lc,ia. d)

F::'(~r.'c:j 1,ci;::1 c:j(:;! (::or\ f i. ar\ ,:a de l(:)s LlsLlar..i.os. e) Desat-r..o 11 ar EE:, tra teg ias

ant.i-resis.tel-,I: ia

,+ . S,~:-.:.,.,_.f~Xlf:.r'l~~.I.b',~.I_$J:,t:l (Llrl i vt:-:-:o 1". s i, d a ci CI [30 b i. f:~ r,\ 0 )

i:::'!) F::'I:!!!r-(Ji,d,::1. d!'=? col'l'fial'lza df:~l i:~I:JricLllt,or. t)) Desar-rollal'- con

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X) ORGANIZACIONES INTERNACIONALES RELACIONADAS CON RE§ISTENCIA

I)E .b-'i dc ') ::, .j ro" \'- r-.c"J."jJ "" fY."'<::" I .I ! - 1'-".."- J'c" t '= r-" J' -, 1 _"" ( --""-C. 1'-' b "" 1° c-ar ' te c:: se h "' n",'_.".=."!,::,.I"...,..:,,lc:,=,.J'::! ,:,:-::,,_:,,=1,._."-'\ ct~,_c:l-,c:I_,:' "-'\1- I,~ ""

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E "'" ,-" ,_.:..,.::, ,'"::,, " J.F.t',-:F::' «(~r .l\ rJ .") J' r-t""' r ' I"'-' C" J' ( - I-.aJ C""'" F""' odLlcto "" es de,--, to""I'-' .J'.n ~~,,::,""""',!o..., .:) ."L ,1,_,,- .d-o.ll, J"" I 0 I

Agr'I:)(::jU,,_tfl:LI:<:)S) y coor'l:1 :.lric.il'- s~'?mj,I-lar-il:J!5:1 pLlbl i(':c.'l.ciories e

J' r-" E""";:' (. J' C'I-' I .; c"j r.o=""

r"_'.",c:I,._."I,.;:o'.

XI) EJEMPLOS DE RESISTENCIA EN COSTA RICA.

1 )' Resi..-i..er'ri'. (jf'" 1"1v-{");::--j'.'-f:."'rell,- 'fi .1'i'-jj-i<::" ( Si g atol-:a Ne g r- ) a, - - -'" - , 1 ,,-, c:\ - ,_:..L.b.::=-,::.'J_-,,_!.:~-,-,,-=_: :?!:, ..:_- -.,!;.;:!.,-~,:.,_.,~, ' - 0 ~

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t""f~l- 199.1 ,.I!';:lr-rj(J cc.' h-'b'a (je.tec.t;.-(jo r-e<:::i",..ti:::.n-.i.ri a IJ~:'li.Jmil er1 varias ~;~r-_1 , 1.. ._,- ,"_"\.1.,- -- _"\ ..,.-o_.-L.~. -I. _.~:

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aceptC!rl Ilj::ICIC tC::IS cle cclbcl11 erc:)~:; II CjE~ r'ec::(::)merlcjac iC)rlt'!!r::, er) el LIStJ de

BelitJm:i 1; 10s .tf:.~(::n:j_(::os ir11:jic:,:\r-(::JI-1 qLle '5~~ riecl=~s.i'tci Irl'~s que LII'1 acI_IercJo

er . t ."- E' E' J lc ""': 1.,':',""'" n l ""- c:;'::;, "' l """' cj ='- r" f"::"",.t.'-. t . le ' c"""'- r"::. c..c-rr' er-c-I-'c .; or -"."<::: ni le ' "::""""r- 1.1, "-,..,(...1""-1"",-,," ,,,-::=1 .._\,,":lJ :.., "..-JI 1.0."\... 1"""'-1-.~=.;,1

acata(ja~3 obI i(.;lE'Ii::(~1r-:iameri.tl::i!. EJ. ~~rLII:)O (:JE,"!! t~:?cliicos t:orl el apor-'te rjf:-:!

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negra.

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con Beliom.i 1 ind.icar(:)n 16\ preser1cia (je a,i!:-51amier1.tos cje er::,te hongo

resistE:~rlt.E's; '::1 Ben~~imidc:\~~c)lf!!!:;. E:s.t.:\ .irj'for'mE'Icior) 'fLle presel-ltC:'Ida er1

e1 I I COI-)i.";)r-eso Na(::.i(:)r1aJ. de F.itopatologia de la Asoc.iacior1

Co!:.;;tar'r':i.cen~:;e c:le F'i.t.or:lat:.cjlogc.1s del 1.2 a1 14 de 1T1'::'tYCI de 1993, pr.>f-

G. AStl_I,3, L._.F. Ar"al_lz Y C::). Umai,,:\.

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el t:olii:rol rje er1'fermf:?(j'3tjf.?s f.>(:)rJr"{":lI-) r"e'/elar otr-i::Js (::as(:)i::.; (jE?

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se Li.ti.1i~~c\bc\ri mf'ls l;:ic10s tie BE!rlom~il. '..ambit'::!ri erl urla pr-ueba en e1

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f~:J. c:\L.lt(::,j'. ~:)r'c:f~:)c::'rl{,,' lc:\ i'c:lrlllclc::itJn (:IE.' L.lrla (~omisi,6rl riacior\a1 sobr-e

Y".e!:::,i~;{::'E':nl::: La "-"'I i:Llrlgi.(::i(::!as p(:;)~s.ib1emef-1 te .i.n.t:.E'<]r-ac.la por- t-epr-esen.tantes

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~ I I-, g et-i i.f.~r.'o"; 1~(:Jt-()riOm(:;)~:::,; San.idad Ve(Je.ta1 e Inves.tigaciol-"les del MAG;

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Page 14: lJ 92 RESISTENCIA A FUNGICIDAS. - mag.go.cr · 11- '1,::;:_,,11 '" --"\0 -";',,--,c 1 liP r' ~"

Comisj.6n t"i::lc.l.c:)ri,,':\l rje Sj.~.!c:I1::.C,~:.a !\le(;}F",:!; Ir1!:=:.t..jt:LI.cj.(:)nt~!:=:. cje E-=riSenar-I~:a ,0Uni versitat- ia y Caff/8.Y-'::-I r".3C ic~\na:L de InsLlfflos Agr'(.jpecI_IaF"ios; :L a CLI.31

dara F"E~comer\cjaciorIE!s es~:iec:ifj.cas sobr'e E:"-!1 LISO de fLlr\gicidas

propensos a de!-:5c1F"r'oll.:.ir- r"e~;;.ister/c.i,3.~ la cLI.~l tambi~n ~jodt-'a

aseSOF"aY. a a(::jLlel1cfs 1egj.s1.'=Idc.1r"e!:=:. qLle ~Jr"esenter\ pr"oyect-:os de 1E~y

re1at.i'/os a1 LISO de fLlr1gic.ici,;:'\~, como ~jar.te rje 10s a(;;)F"CiqLlim,ic:os como

e1 PF"fcoC'.:er\t 'a do ' P fi r- ] ", !') 'ir' J' L I' !" :' d :::. ":"".-""' r" 0) SEcor", r~F,"f"" (::'1 '-;,'6 CjE' ; LI 'I l' O cje- ""J '" - ,0.'\ ""t ."c.. c", r,=",.., '" ', 1'" ..., ,.- -, ,.,

1991~ ql_Ie I/O c':(.jns,i,(~1ef-~\ el i:.'\sf::Je(::1:r_1 dE'! F"es,is't:f?l/c.ia a plar;)uicidi:1S.

LITERATURA CONSULT ADA

.1. Ar.6r1:.i.mo. Si(].:.i17,op:,a 1-/(':!!~Jr.,~ y am~\F"il1a.. -r<f'cniC:'::-Is ffle_jor'adas par'a

fflcir1ejc) e j.derlt:ific'::lc::.i.c:ln (tamt:)ieri E:!ri ]:r'(;j1t~s). DLlpont

l;grc.1qL\{m.ic:0~5. COf-~~{:1i- Gables F1orid.:.i .17 Ij{;!~l.iri';:'\s.

~~--:I Ac t l \ " G. F l r""'.' LI "" ,.~ lJrr"'~'- 199 -:' r' EO)r""-' l' b 'i ll. ' d '. cj r 'c::.dL 'C " d '"

..:... -:> ..c:l , :I,. -. rl,'::\ .:,., \:J. "Ic:lnc:l. ,,':'. .-:>,. ,~::>, ct ,, =

de- (~ c. ll c=. t ,.' tr ;--I-I' I" i " I] /-':::, ( ",,::r' c- r-': ol. ' d c::.,,:: ;:" l'c:: l "' cj '- (j c::. P-""' j"".'J"" " 1,~-.:!...:,,_.:._'=:..-::~_.:: !:--~:::..!.!..:::,,-,!- ~_..:.:..:::l_"":,,,::l,:-.::;,~-!_,_.!.:_, ,:-::f::,,:;~, ..;\ ,. ,'" "" L! "'" ,-, Jo.-I7 = 0.'\

tiaberic:lazo1e. II Corigrf:'YSO t.Jar.::j,oria1 de Fj,tcjj:icI'to1ogia -

M e m (j F" .i a ,- ~')/ 5 0 (::: ,1, a c: i 6 r1 (::; (.~) s t a F" I~ i C f.'! ri ~::; f":!! cj e F:- i top,~ t 0 log :1. '::'1 . S '::'11-1

Jc """"'" ( ~ c-c:: t "" F" I' C'-. F'J c'l l ' r'- r,, (-).1""~., ,J~,,--- \" I~ c:\.:}, 1(:\ .::.-.

3) Delp. l~.J. 1979. RE!si~::;t.:.irice 'to p1ar1t Disea!.'?e Colltr-ol Agel/d't.

Hol.AJ i::,(j c:or)e Io-Jith j,t.. In't. CcJn<;!y-ess of F'Ic:'r1t j:.1r'otectior1.

l.J a s II :i. r1 (;11: (:j r1 D. (::; .

4) ' F A (1 1 !:f88 F:"""""' c"" me ' ," c!'-"'cj ff,c:;" !" I"' ci c.I<::: fc " r" '!" t""'" C"et c::,c" !" l' C')r" "' r" c". .". ,. . .",.\._-,1111",1",_" ."""I.,I"J J ,,"" ,,1"",..,_..,1 c:lIJ

ff/ea!.'?LlreITler1t o'f 'fLlngic,i(je F:;:f?S,1,stanc:e. F.A.(:.i. ffletho(js :24,-:33_.

F" 1 :::or" t ' r )r ' Q( "" IC' t J' C" r"' t" ll ] ] c" i" ; I_.=1 .. ".I.., .,..J. ,J",.",::., I. ~"\

.. w' , -" .. j

-14-

Page 15: lJ 92 RESISTENCIA A FUNGICIDAS. - mag.go.cr · 11- '1,::;:_,,11 '" --"\0 -";',,--,c 1 liP r' ~"

No. 24 F'rincipios generales~

No. :;:~~:.i M~todc! t-E?!:;:,istE'nc:i.c:t ,~ 1:ur1<;;jic:j.das par'a :~r:l.:!;JJ.r:..!.E:- spp en

mar\zanas y peras.

t-lc:). :;~:6 1"1~t:.C:ICj(:) !=)ar'a y"esister\cia a Berlzimidazoles c.ie ~Q.r1ilj.-E- spp en

'f Y-L\.ti:iS y r-,Lleces.

r,Jo. 2-1 1"1f!-:!.t:ocio !=)c:lr'a r'esister\c::ia a fLlr-I(;,Ijcidas en r1::U..£J..ih\. pOl.YQ~ en

fjfE!p.iliOS.

No. 29 I"1et(:Jcj(::J par'a re!::',ist-::::r\cia C:t fLlr\gicidas er\ [1ilgiL!. .9J:.Qodonoso

(Do~Jny) er1 Livas.

t'Jo. :::;i:) I"1~.t.odo par.::'! Y"E'.!!::',ist.f}!!r\cia er\ tizt!r1 tc.'Ir'd.io F'hytophthor'a) en

papa.

No.3 .1. 1"1 (!! t 0 c:1 0 ~:) <:it r' ,"::\ res j. 'Z'; t e r1 cia er\ E:.Y.r...?:.s-::J::.!J..s£ll Q.!:~z § §

5) F'Llligicide Resistar1(:e j.n crop protection. 1982. Editors J.

De~:.~:.E"!r' and 5.G. Ge(::JrgeopoLllous. Centr'e for AgricLlltural

F'Llblishil-I(J ,and DoCLlmer\.tation - Wagenirtger\ the Netlierlands.

~ 26 ,::. r~).

6) F:-Llr1~Ji.c:Lcje f:-\e~:;i!',;,t:.arlc:~'!! 1.rl North Amer':i.ca 1988. Editor C. J.

Delp. 'T""1f}!! /;/f,el'-.i(::ar1 f:'hytopathological Society. St. F'aLll

Minne!::',ota. 1.33 p.

:7) ~::oJ.J.er!l W. and H. ScheinpflLlg. 1987. FLlngal Resistance to

S.tey-ol B.iosynthesis Inhib.itors: A new challenge. Plant

[ ).. "" "" ;: c:: ,,;;, 71 ( j -:;,) . j (-t66 ' -. 1 (-}7 ~..""""""t_J;:;; \~.. - - . - --.

8) F'est.:i.t:icie F:ei::..is.t,:::\r1ce .- Strategies and .tactics for Management

.1.986. t'Jat.i.c:)j'-,a.J. Ac:."::\deffIY F'res!:::.. Washirlgtor\. D.C. 471 p.

~ ~

-15-

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