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Insect Flight: Aerodynamics, Efficiency, and Evolution Z. Jane Wang Cornell University Copyright Z. Jane Wang, 2007

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Page 1: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Insect Flight:

Aerodynamics, Efficiency, and Evolution

Z. Jane Wang

Cornell University

Copyright Z. Jane Wang, 2007

Page 2: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

E. J. Marey1830-1904

Copyright Z. Jane Wang, 2007

Page 3: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Harper Magazine 1870

Copyright Z. Jane Wang, 2007

Page 4: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

L ~ 1cmf ~ 40HzRe ~ 3000

Copyright Z. Jane Wang, 2007

Page 5: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Copyright Z. Jane Wang, 2007

Page 6: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

walkCopyright Z. Jane Wang, 2007

Page 7: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Lanchester (1897) Kutta (1903) Joukowski (1903) Prandtl (1914) Wagner(1925) Karman and Sears (1938) ……

Dragonflies (~350 Millions years ago)

Rayleigh (1842) Kichhoff (1887) …...

Copyright Z. Jane Wang, 2007

Page 8: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

∂u∂t

+u ⋅ ∇u = −∇P /ρ +ν∇2u

m dvb

dt= F fluid + fext

Navier-Stokes Equation Subject to Wing Motions

Why compute and which methods?

1. Single rigid wing in prescribed or free motion

-Co-moving frame, 4th order in time and space (Phys. Rev. Lett. ‘00, ‘04)

2. Multiple rigid wings in prescribed motion

-Cartesian grid + overset grid, 4th order in time, 2nd order in space

( J. Comp. Phys.’03)

3. Multiple flexible wings in prescribed motion or prescribed force in 2D

-Immersed Interface Method 2nd order in space and time

(SIAM Sci. Comp. ‘05, J. Comp. Phys. ‘05)

4. Multiple flexible wings in 3D

-Immersed Interface Method 2nd order in +space and time (Comp. Mech. ‘07)

∇•u = 0

Boundary condition (no-slip):

ub = vb

Incompressible flow:

Moving wing:

Copyright Z. Jane Wang, 2007

Page 9: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

ZJW, PRL 2000Copyright Z. Jane Wang, 2007

Page 10: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Dragonfly = Drag on Fly

Drag supports majority of the weight.

If drag is neglected, the required lift coefficient ~ 4-5, which is ‘anomalously high’.

down up

Copyright Z. Jane Wang, 2007

Page 11: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Why insects flap their wingsthe way they do?

Copyright Z. Jane Wang, 2007

Page 12: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

~180° phase difference during hovering.~90° phase difference in forward flight.~0° phase difference for large forces.

Ruppell (1989)

Fore-hind Wing Interactions

Copyright Z. Jane Wang, 2007

Page 13: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Copyright Z. Jane Wang, 2007

Page 14: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Fore-Hind Wing Interaction Can Reduce Power for Hovering

Drag reduction

ZJW, D. Russell, PRL 07Copyright Z. Jane Wang, 2007

Page 15: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

~

Passive Wing Pitching

A. Bergou, S. Xu, ZJW, J. Fluid Mech., 2007Copyright Z. Jane Wang, 2007

Page 16: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Copyright Z. Jane Wang, 2007

Page 17: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Optimal ?

Are Insect Wing Motions

Why should nature optimize?

Optimize what?

Copyright Z. Jane Wang, 2007

Page 18: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Do Insect Wing Motions

Minimize Energetic Cost?

Why should they?

Which energy?

Copyright Z. Jane Wang, 2007

Page 19: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Do Insect Wing Motions Minimize

Mechanic Power in Hovering?

Why should they?

Do they?

Copyright Z. Jane Wang, 2007

Page 20: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Energy Minimizing Hovering Wing Motion

Given a wing and a weight, find wing motions

that minimize the aerodynamic power to support the weight

Constrained PDE/ODE optimization

Problem:

Copyright Z. Jane Wang, 2007

Page 21: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Fruitfly:

f = 230Hzηm = 70.2o

Cη = 2.724

Φη = −72.3o

K = .727φm = 89.0o

θm = 2.86o

Φθ = −98.0o

θ0 = 1.75o

N = 2η0 = 89.6o

P* = 14.8Wkg

L −1 = 4.1 ⋅10−10

f = 230 Hz

Optimized Wing Kinematics vs. Observation

Fry, Sayaman, and Dickinson, JEB (2005)

f = 225 Hz

Copyright Z. Jane Wang, 2007

Page 22: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Optimization Results for Insects

Berman and ZJW, J. Fluid Mech. (2007)

Fruitfly Bumblebee Hawkmoth

Copyright Z. Jane Wang, 2007

Page 23: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

12 Species of Drosophila (fruit-flies)Wild-type and Mutants

With I. Cohen GroupCopyright Z. Jane Wang, 2007

Page 24: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

‘Birds’ vs. Plane

Copyright Z. Jane Wang, 2007

Page 25: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

In Classical Aerodynamics Limit

vs

(no need for additional propulsion )

Hovering with

same wing, same weight

Copyright Z. Jane Wang, 2007

Page 26: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

In Classical Aerodynamics Limit

Optimal Steady Flight Sets the Bar

CL/CD ~ 60 CL/CD ~4.7

Independent of Re

Independent of Wing Shape

Copyright Z. Jane Wang, 2007

Page 27: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Including Unsteady Aerodynamics

Copyright Z. Jane Wang, 2007

Page 28: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

U. Pesavento and ZJW, Preprint

Flapping Flight can be Less Costly

Copyright Z. Jane Wang, 2007

Page 29: Insect Flight: Aerodynamics, Efficiency, and Evolution · Title: Microsoft PowerPoint - 2007-APSDFD-JaneWang-Archive.ppt Author: Ken Created Date: 3/18/2008 2:17:57 PM

Fly Group:

Fore-hind wing interactions in dragonfly flight : David Russell

Passive wing pitch reversal: Attila Bergou

Optimization: Gordon Berman

Flapping vs. fixed wing flight: Umberto Pesavento

Fruitfly Exp: Itai Cohen and Leif Reistroph

Falling Paper: Anders Andersen, UP

Immersed Interface Methods: Sheng Xu

http://dragonfly.tam.cornell.eduCopyright Z. Jane Wang, 2007