genomic diversity and evolution of coronavirus (sars-cov-2 ... · 04/09/2020 · ten were in...

Genomic diversity and evolution of coronavirus (SARS-CoV-2) in France from 309 COVID-19-1

infected patients 2

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Anthony LEVASSEUR1,2,3#, Jeremy DELERCE1,2, Aurelia CAPUTO1,2, Ludivine 4

BRECHARD1,2, Philippe COLSON1,2, Jean-Christophe LAGIER1,2, Pierre-Edouard 5

FOURNIER2,4, Didier RAOULT1,2# 6

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1 Aix-Marseille Université (AMU), MEPHI (Microbes, Evolution, Phylogeny and Infections), IRD, 8

APHM, Faculté de Médecine, Marseille, France 9

2 IHU Méditerranée Infection, Marseille, France 10

3 Institut Universitaire de France, Paris, France 11

4 Aix-Marseille Université (AMU), IRD, APHM, SSA, VITROME, Marseille, France 12

# Corresponding authors: Didier Raoult: [email protected]; Anthony Levasseur : 13

[email protected] 14

Data deposition: genomes were deposited at EMBL-EBI under the BioProject: PRJEB37722 15

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(which was not certified by peer review) is the author/funder. All rights reserved. No reuse allowed without permission. The copyright holder for this preprintthis version posted September 4, 2020. ; https://doi.org/10.1101/2020.09.04.282616doi: bioRxiv preprint

https://doi.org/10.1101/2020.09.04.282616

ABSTRACT 23

The novel coronavirus (SARS-CoV-2) causes pandemic of viral pneumonia. The evolution and 24

mutational events of the SARS-CoV-2 genomes are critical for controlling virulence, transmissibility, 25

infectivity, severity of symptoms and mortality associated to this infectious disease. We collected and 26

investigated 309 SARS-CoV-2 genomes from patients infected in France. Detailed genome 27

cartography of all mutational events (SNPs, indels) was reported and correlated to clinical features of 28

patients. A comparative analysis between our 309 SARS-CoV-2 genomes from French patients and 29

the reference Wuhan coronavirus genome revealed 315 substitution mutations and six deletion events: 30

ten were in 5’/3’ UTR, 178 were nonsynonymous, 126 were synonymous and one generated a stop 31

codon. Six different deleted areas were also identified in nine viral variants. In particular, 30 32

substitution mutations (18 nonsynonymous) and one deletion (Δ21765-21770) concerned the spike S 33

glycoprotein. An average of 7.8 mutational events (+/- 1.7 SD) and a median of 8 (range, 7-9) were 34

reported per viral isolate. Comparative analyses and clustering of specific mutational signatures in 309 35

genomes disclose several divisions in groups and subgroups combining their geographical and 36

phylogenetic origin. Clinical outcomes of the 309 COVID-19-infected patients were investigated 37

according to the mutational signatures of viral variants. These findings highlight the genome dynamics 38

of the coronavirus 2019-20 and shed light on the mutational landscape and evolution of this virus. 39

Inclusion of the French cohort enabled us to identify 161 novel mutations never reported in SARS-40

CoV-2 genomes collected worldwide. These results support a global and continuing surveillance of the 41

emerging variants of the coronavirus SARS-CoV-2. 42

Keywords: French cohort, SARS-CoV-2, coronavirus, evolution, bioinformatics, outbreak, 43

phylogeny, COVID-19 infected patients 44

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INTRODUCTION 48

The new coronavirus SARS-CoV-2 has now spread to every country in the world. SARS-CoV-2 is 49

classified among the positive-sense single-stranded (ss)RNA viruses as the seventh coronavirus known 50

to infect human1. Four genera of coronaviruses have been identified in the subfamily 51

Orthocoronavirinae of the Coronaviridae family (order Nidovirales). Described for the first time in 52

December 2019 in Wuhan, China, the new SARS-CoV-2 coronavirus will naturally accumulate and 53

fix mutations in its RNA genome as the epidemic evolves, partly due to the high immunological 54

pressure in humans. A number of characteristics of coronaviruses are still unknown and one of the key 55

questions is how fast is the SARS-CoV-2 virus evolving? Indeed, mutations could drastically result in 56

difference in virulence, transmissibility, and clinical evolution. Therefore, the frequency and pattern of 57

mutations are essential features to be considered in this ongoing global epidemic. 58

The genome of SARS-CoV-2 is a positive ssRNA of approximately 29.9 kbases in length, with a 59

5’cap structure and 3’polyadenine tail2. Two large ORFs are translated into polyproteins arising from 60

ribosomal frameshift that are subsequently post-translationally processed to produce 16 proteins 61

conserved between coronaviruses3. These 16 proteins are involved in the synthesis of viral RNA and 62

immune evasion4. Four structural proteins (spike S, envelope E, membrane M and nucleocapsid N) 63

and at least nine small accessory proteins, including proteins unique to SARS-CoV-2, are produced 64

during viral genome replication3,5. 65

By comparing alpha- and beta-coronaviruses, it appears that SARS-CoV-2 exhibits notable genomic 66

features such as an optimized binding to the human receptor ACE2 and a functional polybasic (furin) 67

site in the spike protein with predicted O-linked glycans6. The receptor binding domain (RBD) 68

exhibits the most variable fraction of the genome of SARS-CoV-2 (only ~40% amino acid identity 69

with other SARS-CoV coronaviruses). The origin of this virus is debated but two main hypotheses 70

emerged based on either the natural selection in an animal host before zoonotic transfer or natural 71

selection in humans following zoonotic transfer7. Nearly identical RBDs found in pangolins 72

coronaviruses and SARS-CoV-2 could indicate an acquisition in SARS-CoV-2 by recombination and 73


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mutations in a parsimonious scenario6. Metagenomic sequencing identified coronaviruses close to 74

SARS-CoV-2 from Malayan pangolin and confirmed the natural reservoir of this virus in wild animals 75

and the risk of zoonotic transmission8. 76

Being a RNA virus, SARS-CoV-2 is prone to mutate because of the lack of proofreading activity of 77

polymerase. Previous work in SARS-CoV reported an exoribonuclease domain (ExoN) in non-78

structural protein 14 that could provide proofreading activity protecting the virus from high rate of 79

mutagenesis9. Mutations have to be carefully studied and related to the severity of symptoms and 80

clinical evolution associated to patients. In addition, the mutation rate correlates with drug resistance 81

and escape from immune surveillance. The emerging respiratory disease outbreaks caused by the 82

SARS-CoV-2 required an urgent need for comprehensive studies that combine genomic data, 83

epidemiological data, and chart records of the clinical symptoms and clinical evolution of patients. In 84

addition to genomic mutations, transcriptomic studies have revealed several viable deletions in 85

essential proteins, including glycoprotein S reinforcing that several regions are prone to mutate10. 86

The 2019–20 coronavirus pandemic was confirmed to have spread to France since January 2020. In 87

this work, 309 genomes of SARS-CoV-2 from French patients monitored in IHU Méditerranée 88

Infection were studied and compared to the isolates from different countries in all continents. 89

Mutational events were evidenced and compared to the reference Wuhan coronavirus isolate. Finally, 90

clinical issues and impacts of these mutations were investigated. 91

RESULTS 92

Mutational landscape of the SARS-CoV-2 genomes from the French cohort 93

The mutational landscape of 309 coronavirus genomes was tracked in terms of SNPs and 94

insertions/deletions (Table 1). As compared to the reference genome SARS-CoV-2 isolated in Wuhan 95

in December 2019, a total of 321 mutational events were reported in the SARS-CoV-2 genomes from 96

309 patients. A mean of 7.8 mutational nucleotide substitutions (+/- 1.7 SD) and a median of 8 (range, 97

7-9) were reported per viral isolate. Ten substitutions were retrieved in the non-coding part of the 98

genome. In the coding region, 305 nucleotide substitutions were detected representing 178 99


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nonsynonymous (~58.4%) and 126 synonymous mutations (~41.3%) and one stop codon mutation 100

(~0.3%). Some mutations were commonly distributed in the majority of our isolates whereas 208 101

substitutions (~68.2%) were unique to one isolate. Regarding missense mutations, we noted drastic 102

changes of aminoacid (AA) in which the physicochemical properties of the AA were severely 103

interfered (Table 2). In particular, the spike gene contained 30 substitutions (18 nonsynonymous, 60%) 104

and included one deletion of six nucleotides (Δ21765-21770). Among mutations retrieved in the spike 105

gene, the missense mutation P330S (22550, CCT>TCT) was embedded at the border of the receptor-106

binding domain (RBD). A stop mutation was also found in ORF7a at the protein position 38, meaning 107

that the protein was prematurely truncated at the first third of its sequence. 108

Interestingly, we identified six putative deleted regions distributed in nine viral variants among the 109

genes nsp1, nsp2, spike S, ORF3a and ORF7a. These deletions removed entire codons for nsp1 (Δ510-110

518 and Δ686-694, three AA), nsp2 (Δ1605-1607, one AA), spike protein (Δ21765-21770, two AA), 111

ORF7a (Δ27434-27436, one AA) and consequently, conserved open reading frames. Moreover, the 112

deletion event retrieved in nsp1 (position Δ686-694) was found in three different patients and the 113

deletion in nsp2 (position Δ1605-1607) in two different patients that were not epidemiologically-114

related, meaning that these two evolutionary events could be a convergent evolution of the virus. In 115

addition, one single nucleotide deletion impacted ORF3a at the position Δ26161, this frameshift 116

mutation lead to a truncated protein missing 19 AA with a stop codon generated at the protein position 117

259. 118

The occurrence of specific mutations in the 309 genomes was studied and classified. In order to 119

organize our dataset into meaningful groups and identify coherent patterns, a double hierarchical 120

clustering of the detected mutations was generated (Figure 1). The hierarchical clustering revealed 121

preponderant mutations shared by a majority of the isolates. It divided our 309 patients into five main 122

clusters corresponding to specific mutational signatures. Regarding the clustering of mutational 123

events, we detected the presence and co-occurrence of specific mutations in different clusters (Figure 124

1). Among the main clusters, we pointed the cluster 1 (44 patients, 14.2%, positions [28881-28882-125

28883]) with two nonsynonymous mutations in protein N (R203K; G204R). Cluster 2 (39 patients, 126


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12.6%, position 15324) contains a synonymous mutation (C15324T). Cluster 3 (126, 100 and 211 127

patients, at positions 2416, 8371, 25563, respectively) includes one synonymous mutation (C2416T), 128

and two nonsynonymous mutations (nsp3: Q1884H; ORF3a: Q57H). Cluster 4 (68 patients, 22%, 129

position 1059) contains one nonsynonymous mutation (nsp2: T85I). Finally, cluster 5 (from 297 to 130

303 patients, 96-98%, positions 241, 3037, 14408, 23403) displays one mutation in 5’UTR (C241T), 131

one synonymous mutation (C3037T) and two nonsynonymous mutations (nsp12b: P314L, S protein 132

D614G). 133

The genomes of all 309 isolates were integrated in a global phylogenetic reconstruction including all 134

other 200 genomes of SARS-CoV-2 available in France (Figure 2 + supplementary figure 1). We 135

observed different clades corresponding to the previous mutational patterns revealed by the 136

hierarchical clustering. These mutations were positioned in phylogenetic trees. Globally, the 137

distribution of the 309 SARS-CoV-2 genomes from our patients was distributed across all lineages. 138

Noteworthy, we observed a high density of isolates from our cohort in the cluster 3a corresponding to 139

the mutation nsp3: G8371T, Q1884H. Phylogenetic reconstruction was also carried out including all 140

complete genomes available worldwide (5519 genomes on 15th April 2020). All viral isolates of our 141

French cohort were mainly distributed in clades including European isolates (Figure 3). Again, the 142

mutation in the cluster 3a (nsp3: G8371T, Q1884H) regrouped almost exclusively the variants from 143

our French cohort (98/105 genomes). Compared to the Chinese reference, we observed a prevalent 144

distribution of specific mutations at positions 241, 3037, 14808 and 23403 that spread in 145

approximately ~3570 (+/-7) genomes representing ~64.6 % of the isolates worldwide. These mutations 146

have been reported in variants from all continents. Although, we detected some clades depending on 147

the common geographical origin of the patients, a global distribution of variants in all continents was 148

observed, therefore meaning and confirming that SARS-CoV-2 is circulating worldwide. 149

Clinical outcomes of the 309 COVID-19-infected patients were investigated and integrated in our 150

evolutionary analyses (Figure 1-3). Among these patients, we classified the outcomes in three 151

categories including poor clinical outcome (PClinO, defined by either death or transfer to intensive 152

care unit or hospitalization for 10 days or more), poor virological outcome (PVirO, defined by viral 153


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shedding persistence at day 10) and good clinical outcome (GO) (Supplementary Table 1). 154

Coronavirus genomes isolates from 38 patients’isolates with PVirO were widely distributed across the 155

groups, including diverse mutational events meaning that no correlation between higher viral loads 156

and the genomic specificity of the virus was evidenced as previously reported (supplementary figure 157

2). For the 10 patients with PClinO, a majority of isolates were also distributed to all groups 158

(supplementary figure 2). An exception concerned two patients (IHUCOVID-0318, IHUCOVID-159

0333) with one PClinO and one death that were clusterized together in a group of seven different 160

isolates. A common nonsynonymous mutation in the membrane protein M (L129M, CTG>ATG, 161

position 26907) was found in this cluster. 162

DISCUSSION 163

Systematic gene level mutational analysis of the genomes allowed us to identify several unique 164

features of the SARS-CoV-2 genome. In this study, we reported 315 substitution mutations including 165

126 synonymous mutations. Although these later mutations conserved AA, it has been reported that 166

the fitness of RNA viruses with genomes harboring synonymous mutations could be affected11. 167

Regarding the 178 nonsynonymous mutations, alteration due to the replacement of specific AA 168

depends on several factors including their physicochemical properties, steric hindrance and the 169

position of the AA in the tridimensional structure of the resulting protein. A key element of 170

coronavirus host range is determined by the binding affinity between the spike S protein and the 171

cellular receptor12. Here, thirty substitution mutations (including 18 nonsynonymous) were reported in 172

the spike protein S. The spike protein mediates viral entry into host cells by recognizing angiotensin-173

converting enzyme 2 (ACE2) as its receptor13. We detected a missense mutation P330S (CCT>TCT) 174

located at the border of the RBD14. Furthermore, a deletion of six nucleotides Δ21765-21770 removed 175

two AA in the resulting protein. Mutations in the spike surface glycoprotein (A930V (24351C>T)) 176

have already been reported in the Indian SARS-CoV-215. All these mutations in protein S could 177

influence host range and transmissibility of the virus. Consequently, the spike S glycoprotein is a 178

potential target among potential therapeutic strategies. Focusing on the mutations identified in our 179

study enables to drive the strategy to target special positions of the protein. In this line, specific 180


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antiviral therapies on S protein should be conducted in vitro (involving RBD-ACE2 blockers, S 181

cleavage inhibitors, neutralizing antibodies, inhibitors and small interfering)16. Additionally, truncated 182

proteins were also detected for ORF7a with a stop codon mutation (position 38 on 122 AA protein 183

length) and for ORF3a with a frameshift mutation leading to an aborted protein missing 19 terminal 184

AA. These mutations represent major changes in the protein architecture that have to be 185

experimentally tested in order to assess the impact on protein function. 186

In evolutionary biology, convergent evolution is a central concept. The importance of a mutation is 187

greatly strengthened if one can show that the fitness has evolved independently multiple times17. Two 188

deletion events (Δ686-694; Δ1605-1607) were detected in three and two variants while the COVID-189

19-infected patients were not epidemiologically-related. These results strongly support a convergent 190

evolution and could suggest convergent adaptation of the virus. 191

Phylogenetic trees were reconstructed by including isolates from France and all isolates worldwide. 192

Isolates were grouped according to continents, but the large circulation of the virus was evidenced. 193

Interestingly, we detected only two isolates belonging to the S-type in our 309 patients’ cohort18. 194

Noteworthy, an almost exclusive group including SARS-CoV-2 genomes from 98 patients monitored 195

in IHU Mediterranée Infection was phylogenetically close and regrouped in a clade of 105 genomes. 196

Classification based on mutation profiles enables the inclusion of patients in specific groups and 197

correlates this clustering with the clinical features in order to favor a better approach to treatments. We 198

reported widespread variants that extent to different groups and pointed five clustering groups that 199

spread to different lineages. Regarding the clinical evolution of patients, PClinO and PVirO were 200

widely distributed in all groups identified in our clustering analysis as well as in the phylogenetic trees 201

as previously observed19. A single exception was reported in a subgroup with a missense mutation in 202

the gene encoding membrane protein M (L129M, CTG>ATG) in which two PClinO (including on 203

death) were observed. This specific mutation in membrane protein M (L129M) has to be carefully 204

investigated in future large cohort as potential determinant of clinical evolution. But at this stage, more 205

data have to be integrated to state any conclusion or consequence about this mutation. Overall, these 206


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results suggested a clinical outcome that is not dependent on viral variants but rather a patient-207

dependent clinical outcome. 208

In conclusion, the mutational signatures and evolution of the coronavirus SARS-CoV-2 from 309 209

patients monitored in IHU Mediterranée Infection were unraveled. Inclusion of the French cohort 210

allowed us to identify 161 novel mutations never reported before in coronavirus isolates worldwide. 211

This mutations list represents a huge library of potential targets to test experimentally. The isolation 212

and the genomic studies of the 309 SARS-CoV-2 genomes are paving the way for new diagnostic, 213

therapeutic and prophylactic approaches. 214

METHODS 215

Patients 216

The study was conducted at Assistance Publique-Hôpitaux de Marseille (AP-HM), Southern France in 217

the Institut Hospitalo-Universitaire (IHU) Méditerranée Infection (https://www.mediterranee-218

infection.com/). All data from patients were anonymized. The study was conducted on patients 219

included from February 29th to April 4th. Individuals with PCR-documented SARS-CoV-2 RNA from 220

a nasopharyngeal sample20 were selected for genome sequencing. Viral loads ranged as follows: 93 221

patients with PCR Ct <16; 69 patients with [16<Ct<17]; 95 patients with [17<Ct<18]; 30 patients with 222

[18<Ct<19]; 17 patients with [19<Ct<20] and 5 patients with Ct >20. 223

Genome sequencing and assembling 224

From nasopharyngeal samples, genomic RNA was extracted using the EZ1 biorobot with the EZ1 225

DNA tissue kit (Qiagen, Hilden, Germany) as previously described 21. In the reverse transcription (RT) 226

step, cDNA was reverse transcribed from total viral RNA samples using TaqMan Reverse 227

Transcription (Life technologies applied biosystems) following the manufacturer’s recommendations. 228

PCR program was set as follows: 25°C, 10 min, 48°C, 30 min, 95°C, 5 min, 4°C. DNA Polymerase I 229

Large Klenow Fragment (BioLabs) was used for generating double stranded cDNA. 20 µL of each 230

sample were added to a mix (10 µL) of: Buffer 2 Neb 10X, DNA polymerase large Klenow Fragment, 231


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dNTPs working solution (10mM) for 1 h at 37°C. cDNA were purified using beads Agencourt 232

AMPure (Beckman Coulter) and then sequenced with the paired end strategy on a MiSeq sequencer 233

(Illumina Inc, San Diego, CA,USA) with the Nextera XT DNA sample prep kit (Illumina). To prepare 234

the paired-end library, the « tagmentation » step fragmented and tagged the DNA. Then, limited cycle 235

PCR amplification (12 cycles) completed the tag adapters and introduced dual-index barcodes. After 236

purification on AMPure XP beads (Beckman Coulter Inc, Fullerton, CA, USA), libraries were 237

normalized on specific beads according to the Nextera XT protocol (Illumina). Normalized libraries 238

were pooled into a single library for sequencing on MiSeq. The pooled single strand library was 239

loaded onto the reagent cartridge and then onto the instrument along with the flow cell. Automated 240

cluster generation and paired-end sequencing with dual index reads were performed in single 39-hours 241

run in 2x250-bp as previously described 21,22. After sequencing, reads were mapped on the reference 242

SARS-CoV-2 isolate Wuhan-Hu-1 (MN908947.3) using CLC Genomics workbench v.7 with the 243

following thresholds: 0.8 for coverage and 0.9 for similarity. For each sample, detailed per-base 244

coverage information was extracted and all mutations with a coverage ≥5 were considered. 245

Hierarchical clustering 246

Data were ordered and grouped using the distance matrix computation and hierarchical clustering 247

analysis calculated by MultiExperimentViewer23. A double clustering was carried out based on 248

Pearson correlation as a distance matrix and complete linkage clustering as a linkage method. 249

Sequence dataset 250

All genomes available on April 15 of SARS-CoV-2 were downloaded from GISAID (Global 251

Initiative; https://www.gisaid.org/) with acknowledgment 24,25. All genomes were deposited at EMBL-252

EBI under the BioProject: PRJEB37722 253

ABBREVIATIONS 254

AA: Amino Acid 255


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GO: good clinical outcome 256

Indel: Insertion and deletion 257

ORF: Open Reading Frame 258

PClinO: poor clinical outcome 259

PVirO: poor virological outcome 260

RBD: Receptor Binding Domain 261

SD: Standard Deviation 262

SNP: Single Nucleotide Polymorphism 263

UTR: Untranslated Transcribed Region 264

ETHICS APPROVAL AND CONSENT TO PARTICIPATE 265

The study was approved by the Ethical Committee of the University Hospital Institute Méditerranée 266

Infection (N°: 2020-016). Informed consent was obtained from all patients. The study was performed 267

in accordance to the good clinical practices recommended by the Declaration of Helsinki. 268

AVAILABILITY OF DATA AND MATERIALS 269

All genomes were deposited at EMBL-EBI under the BioProject: PRJEB37722 270

COMPETING INTERESTS 271

Authors declare no competing interests 272

FUNDING 273

This work was supported by a grant from the French State managed by the National Research Agency 274

under the "Investissements d’avenir (Investments for the Future)" program with the reference ANR-275

10-IAHU-03 (Méditerranée Infection) and by the Région Provence-Alpes-Côte-d’Azur and the 276

European funding FEDER PRIMI. This research was supported by a grant from the Institut 277

Universitaire de France (IUF, Paris, France) allocated to Prof. Anthony Levasseur. 278

AUTHOR’S CONTRIBUTIONS 279


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AL and DR supervised and conceived the project. AL designed the evolutionary studies and 280

interpreted the data. AL and JD performed the experiments and analyzed the data. JCL, PEF, PC, AC, 281

LB helped with data acquisition. AL wrote the manuscript. The authors read and approved the final 282

manuscript. 283

ACKNOWLEDGMENTS 284

Respectful thanks to all patients and their families enrolled in this study. The authors would like 285

to thank all the engineers, technicians, and clinicians who contributed to this investigation. The 286

authors gratefully acknowledge Olivia Ardizzoni, Vincent Bossi, Madeleine Carrara and Sofiane 287

Regoui for help in sequencing the genomes. The authors acknowledge Yolande Obadia and Audrey 288

Giraud-Gatineau for help in data acquisition. We gratefully thank the authors, the originating and 289

submitting laboratories for their sequence and metadata shared through GISAID (Global Initiative; 290

https://www.gisaid.org/). We acknowledge all the members of CoV-GLUE, Nextstrain 291

(https://nextstrain.org/), and virological.org for sharing their analysis in real-time. 292

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microscope imaging, genome sequencing, and culture isolation of SARS-CoV-2. Eur J Clin Microbiol 355

Infect Dis. 2020. 1‐3. doi:10.1007/s10096-020-03869-w 356

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data management and analysis. Biotechniques. 34: 374-378. 363

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reality. EuroSurveillance. 22(13). 367

FIGURES, TABLES AND ADDITIONAL FILES 368

Table 1. Mutational events detected in the 309 SARS-CoV-2 genomes as compared to the 369

reference Wuhan coronavirus. 370

Table 2. Detailed mutations detected in the 309 SARS-CoV-2 genomes from French patients. 371

Figure 1. Double hierarchical clustering of mutational events in SARS-CoV-2 genomes collected 372

in 309 COVID-19 infected French patients. Top tree: Viral isolates from 309 patients. Left tree: 373

Position of identified mutations according to the reference Wuhan genome. Clusters (1 to 5) were 374

labeled with numbers ranging from 1 to 5. Missing positions were labeled in grey. The mutation type 375

is represented by a colour scale from green (position identical to the reference), red (nonsynonymous 376

mutation), orange (synonymous mutation) and blue (deleted position). 377

Figure 2. Phylogenetic tree of coronavirus in France. The phylogenetic tree was reconstructed 378

from 509 viral genomes collected in France (309 from this study). Number represents the cluster 379


https://doi.org/10.1101/2020.09.04.282616

identified in the double hierarchical clustering. The reference Wuhan coronavirus isolate was 380

underlined in purple. Red star corresponds to PClinO, blue star corresponds to PVirO, grey star 381

corresponds to the loss of follow-up and green star corresponds to deceased patient. Main mutations 382

were labeled along branches. 383

Figure 3. Phylogenetic tree of coronavirus worldwide. The phylogenetic tree was reconstructed 384

from 5519 viral genomes collected around the world. Numbers (1 to 5) represent the cluster identified 385

in the double hierarchical clustering. Red star corresponds to PClinO, blue star corresponds to PVirO, 386

grey star corresponds to the loss of follow-up and green star corresponds to deceased patient. 387

Supplementary Table 1. Baseline characteristics of the 309 COVID-19 infected patients. 388

Supplementary Figure 1. Emergence and phylogenetic tree of coronavirus in France. 389

Supplementary Figure 2. Phylogenetic tree of coronavirus from the cohort of 309 COVID-19 390

infected patients. The phylogenetic tree was reconstructed from 309 viral genomes in this study. 391

Numbers (1 to 5) represent the cluster identified in the double hierarchical clustering. The reference 392

Wuhan coronavirus isolate was underlined in purple. Red star corresponds to PClinO, blue star 393

corresponds to PVirO, grey star corresponds to the loss of follow-up and green star corresponds to 394

deceased patient. Mutations were labeled along branches. 395

AUTHOR INFORMATION 396

Correspondence to Didier Raoult [email protected] or Anthony Levasseur 397

[email protected] 398


https://doi.org/10.1101/2020.09.04.282616

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CO

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-01

62

IHU

CO

VID

-03

75

IHU

CO

VID

-01

72


https://doi.org/10.1101/2020.09.04.282616

France/Valence 425/2020

IHU

CO

VID

-0177

France/HF1795/2020France/A

RA

12253/2020

IHU

CO

VID-0188

France/HF3141/2020

IHU

CO

VID

-001

6

IHUCOVID

-0378

Fra

nce/

HF

2155

/202

0

IHUCOVID-0379

IHU

CO

VID

-010

0

IHU

CO

VID-0060

IHU

CO

VID

-050

2

IHU

CO

VID-0141

France/IDF3345/2020

IHU

CO

VID

-043

8

France/ARA1322/2020

IHU

CO

VID

-021

9

IHUCOVID-0071

France/HF3290/2020

Fran

ce/A

RA

1316

0/20

20

IHUCOVID-0095

France/ARA10170/2020

IHUCOVID-0398

IHUCOVID-0349

IHUCOVID

-0162

IHU

CO

VID

-036

6

Fra

nce/

BF

C27

09/2

020

France/H

F2150/2020

IHUCOVID-0201

IHUCOVID-0447

France/IDF3029/2020

IHU

CO

VID

-0241

Fra

nce/

HF

1995

/202

0

IHUCOVID-0271

IHUCOVID-0160

France

/AR

A11995/20

20

Fran

ce/L

yon

0653

1/20

20

IHU

CO

VID

-010

9

IHUCOVID-0429

IHU

CO

VID

-042

8

Fra

nce

/AR

A12

485/

2020

IHUCOVID-0233


IHUCOVID

-012

0

IHU

CO

VID

-0078

France/B2337/2020

France/C

VL3365/2020

France/IDF3218/2020

Fran

ce/A

RA

1225

0/20

20

IHU

CO

VID

-0300

France/Lyon 0668/2020

Fra

nce/

AR

A12

915/

2020

IHU

CO

VID

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1

IHUCOVID-0229

IHU

CO

VID

-0190

IHU

CO

VID

-0069

IHUCOVID-0365

IHUCO

VID-0

327


France

/AR

A12

238/2020

France/HF1813/2020

IHUCOVID

-033

2

IHUCOVID-0044

IHUCOVID-0176

IHUCOVID-0149

IHUCOVID-0056


France/IDF0386-islP1/2020

Fra

nce/

GE

2843

/202

0

IHUCOVID-0479

IHUCOVID-0106

IHUCOVID-0456

IHUCOVID

-0064

IHU

CO

VID

-0409

IHUCOVID

-0136

IHU

CO

VID

-0222

France/GE1583/2020

IHUCOVID-0239

IHUCOVID

-0375

Fra

nce/

BF

C21

47/2

020

IHUCOVID-0282

France/B

2346/2020

IHU

CO

VID

-0024

IHUCOVID-0049

IHU

CO

VID

-0434

France/ID

F2792/2020

France/HF2797/2020


IHUCOVID-0386

IHU

CO

VID

-007

9

Fra

nce/

HF

1870

/202

0

IHUCOVID-0303

IHU

CO

VID-0389

IHU

CO

VID

-0200

IHUCOVID

-018

9

IHUCOVID-0218

IHUCOVID

-016

7

IHUCOVID-0333

IHU

CO

VID

-050

0

IHU

CO

VID

-028

0France/ARA12499/2020

IHUCOVID-0397

France/IDF2410/2020

France/H

F2586/2020

IHUCOVID

-019

1

IHU

CO

VID

-0057

IHU

CO

VID

-0038

IHU

CO

VID

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5

Fra

nce/

Lyon

066

25/2

020

IHUCOVID-0248

IHUCOVID-0323

Fra

nce/

AR

A10

172/

2020

IHUCOVID-0074

IHU

CO

VID

-0216

IHUCOVID-0286

France/G

E2722/2020

IHU

CO

VID

-0316

IHUCOVID-0226

France/RA739/2020

France/HF3036/2020

France/H

F2151/2020

IHU

CO

VID-0495

Fra

nce/

GE

1977

/202

0

Fra

nce/

AR

A11

980/

2020

IHUCOVID-0151


IHU

CO

VID

-023

1

IHUCOVID

-0494

IHU

CO

VID

-0197

IHUCOVID-0081


France/A

RA

12626/2020

IHU

CO

VID

-0281

IHU

CO

VID

-0128

IHUCOVID

-026

3

IHUCOVID-0293

Fran

ce/H

F274

8/20

20

IHU

CO

VID

-0135


France/IDF3163/2020

IHU

CO

VID

-048

1

IHU

CO

VID-0

110

IHU

CO

VID

-015

8

IHUCOVID-0206


France/H

F1645/2020


France/IDF3040/2020

Franc

e/HF24

96/2

020

IHUCO

VID-0

259

IHUCOVID

-039

1

France/A

RA

12217/2020

IHUCOVID-0203

IHU

CO

VID

-002

0

IHUCOVID-0087

IHU

CO

VID

-0261

IHU

CO

VID-0040

IHU

CO

VID

-0345

IHUCOVID-0099

IHU

CO

VID

-0168

IHUCOVID-0224

France/IDF3274/2020

IHUCOVID-0153

IHU

CO

VID

-0291

Franc

e/G

E3067

/202

0

Fra

nce/

Val

ence

532

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IHUCOVID-0192

IHU

CO

VID

-038

1

IHU

CO

VID

-0371

IHUCOVID-0031

Fran

ce/L

yon

0605

6/20

20

IHUCOVID-0021

France/IDF0373/2020


IHUCOVID-0054

IHU

CO

VID-0238


France/IDF0626/2020

IHUCO

VID-0

208

IHU

CO

VID

-010

7


IHU

CO

VID

-0488

IHUCOVID-0039

Fra

nce/

HF

2948

/202

0

IHU

CO

VID-0

319

IHUCOVID-0211

IHU

CO

VID

-045

5

IHUCOVID-0299

France/HF1871/2020

France/IDF2548/2020

Fra

nce/

HF

3293

/202

0

France/IDF2284/2020

IHU

CO

VID

-0075


IHUCOVID-0308

IHU

CO

VID

-002

8

France/HF1989/2020

IHUCO

VID-0

309

IHUCOVID

-0196

IHU

CO

VID

-0296

IHU

CO

VID

-0337

IHUCOVID-0355

IHUCOVID-0257

IHUCOVID

-0243

IHU

CO

VID

-0485

France/N2223/2020

Fran

ce/A

RA

1297

3/20

20

Fran

ce/H

F259

7/20

20

IHUCOVID-0256

IHU

CO

VID

-0132


Fra

nce/

AR

A10

163/

2020

IHUCOVID-0284

IHUCOVID-0416

France/HF1986/2020

IHUCOVID-0283

IHU

CO

VID

-0329


Fran

ce/L

yon

0648

7/20

20

Fra

nce/

IDF

3230

/202

0

Fra

nce/

Lyon

683

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0

France/HF2405/2020

France/ID

F2534/2020

France/B2330/2020

IHUCOVID-0072

Franc

e/ARA09

686/

2020

IHU

CO

VID

-021

7

Fran

ce/ID

F284

8/20

20

IHU

CO

VID

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9

IHUCOVID-0407

IHUCOVID-0235

IHUCOVID-0012

IHUCOVID-0450


IHUCOVID-0017

IHU

CO

VID-0155

France/H

F2381/2020

IHU

CO

VID

-0240

IHUCOVID-0439

IHU

CO

VID

-0276

France/HF3138/2020

IHU

CO

VID

-022

7

IHU

CO

VID-0360

IHUCOVID-0035

France/B2340/2020

IHUCOVID-0234

France/ID

F3212/2020

Fran

ce/ID

F198

0/20

20

France/HF1988/2020

Fra

nce/

BF

C20

94/2

020

Fra

nce/

HF

1993

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0

IHU

CO

VID

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IHUCOVID-0287

Fran

ce/H

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nce/

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France/B2351/2020

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CO

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-006

3

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Fran

ce/L

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487/

2020

Fra

nce/

Lyon

060

42/2

020

IHUCOVID

-024

7

IHUCOVID-0298

IHU

CO

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-026

9

France/ID

F2989/2020

IHU

CO

VID

-048

3

IHUCOVID-0139

Fra

nce/

CV

L200

0/20

20

Fran

ce/A

RA

1194

3/20

20

France/ID

F2849/2020

IHU

CO

VID

-0496

IHU

CO

VID

-0194


IHUCOVID

-0368

France/IDF2533/2020

IHUCO

VID-0

092

IHU

CO

VID

-0369


IHUCOVID

-0250

IHUCOVID-0344

France/IDF2414/2020

IHUCOVID-0061

Fra

nce/

AR

A09

434/

2020

Fra

nce/

Cle

rmon

t-F

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51/2

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068

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IHU

CO

VID

-0131

France/Macon 06756/2020

France/HF2595/2020

Fran

ce/H

F239

3/20

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France/HF2946/2020

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nce/

HF

2239

/202

0

France/B2335/2020

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CO

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-044

1

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CO

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-0433

IHUCOVID-0125

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CO

VID

-040

5

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CO

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-013

3

IHUCOVID-0214

IHU

CO

VID

-0146

France/IDF2684/2020


IHU

CO

VID

-0421

IHUCOVID-0334

IHUCOVID-0295

IHU

CO

VID

-018

0

France/IDF2256/2020

France/IDF2075/2020

France/H

F2196/2020


IHU

CO

VID-0480

IHUCOVID-0242

France/B2334/2020

IHU

CO

VID-0491

IHUCOVID-0318

IHUCO

VID-0

395

France/B1623/2020

IHU

CO

VID-0094

IHU

CO

VID

-008

4

IHUCOVID-0221

IHUCOVID-0435

France/IDF2532/2020

IHUCOVID-0029

IHU

CO

VID

-003

0

France/HF2237/2020

IHUCOVID-0489

France/ARA1307/2020

IHUCOVID-0116

Fran

ce/ID

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20

IHUCOVID-0032

IHUCOVID

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CO

VID

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France/IDF3276/2020

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CO

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France/HF2039/2020

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ce/H

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CO

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IHUCOVID-0492

France/IDF2420/2020

Wuhan-Hu-1/2019

IHU

CO

VID

-0076

IHUCO

VID-0

249

France/IDF2936/2020


IHU

CO

VID

-0251

IHUCOVID-0449

IHU

CO

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1

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CO

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IHUCOVID

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6


IHUCOVID-0262

France/Bourg-en-Bresse 06813/2020

IHU

CO

VID

-032

8

France/PL1643/2020

IHUCOVID-0122

IHUCO

VID-0

336

France

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CO

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-049

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IHU

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-0082


France/AR

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IHUCOVID-0228

IHUCOVID-0451

IHUCOVID-0209

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CO

VID

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France/B2349/2020

France/AR

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IHUCOVID-0051

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CO

VID

-017

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France/HF2306/2020


France/IDF3324/2020

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CO

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1

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CO

VID

-0304

IHUCOVID-0053

IHUCOVID-0444

France/N1620/2020

France/A

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10251/2020

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-034

2

IHUCOVID-0050

IHUCO

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320

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IHUCOVID-0258

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nce/

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2020

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451/

2020

France/ID

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IHUCOVID-0417

IHU

CO

VID

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Fra

nce/

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3170

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Fra

nce/

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2278

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France

/IDF

2536/2020

France/HF2234/2020

IHUCO

VID-0

432

IHU

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-0006

IHUCOVID-0431

IHUCOVID-0487

France/B2336/2020

Fra

nce/

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2174

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IHUCOVID-0361

IHUCOVID-0204

Fran

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1238

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IHUCOVID-0326France/HF2601/2020

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225

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France/IDF0515/2020

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France/B2348/2020

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CO

VID

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IHUCOVID

-0213

IHU

CO

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France/IDF2279/2020

IHUCOVID-0268

France/A

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11997/2020

Fran

ce/P

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1733

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France/IDF2359/2020

IHUCOVID-0411

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France/B2344/2020

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IHUCOVID

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France/HF3295/2020

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IHUCOVID-0147

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IHU

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France/A

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12630/2020

IHUCOVID-0205

Fran

ce/A

RA

1018

9/20

20

France/G

E1973/2020

IHUCOVID-0232

IHU

CO

VID

-0347

France/IDF0372/2020

IHUCOVID-0045

IHUCOVID-0237

France/HF1465/2020

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CO

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France/B

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CO

VID

-0195

IHUCOVID-0066

IHU

CO

VID

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IHUCOVID-0115

IHUCOVID-0306

IHUCOVID-0490

IHUCOVID-0119

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Tree scale: 0.00001


https://doi.org/10.1101/2020.09.04.282616

Eng

land

/201

3800

41/2

020

Scotland/EDB212/2020

Bel

gium

/ULG

-366

2/20

20

Netherlands/ZuidHolland 74/2020


Eng

land

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4407

2804

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0

Icel

and/

530/

2020

Netherlands/Lim

burg 2/2020

Wales/P

HWC-2

40F7/2020

Australia/VIC154/2020

Taiwan/128/2020

Australia/V

IC40/2020

USA/NY-PV09172/2020

Wal

es/P

HW

C-2

415E

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0

US

A/W

A-U

W214/2020

Wal

es/P

HW

C-2

6FC

B/2

020

England/20129037304/2020

Canada/BC 5522039/2020

USA/N

Y-NYU

MC

150/2020

Iceland/293/2020

France/HF2586/2020

Belgium/LM-030685/2020

Australia

/VIC167/2020

IHUCOVID-0199

Netherlands/Z

uidHolland 20/2020

Australia/NSWID930/2020

Sco

tland

/ED

B08

3/20

20

Icel

and/

480/

2020

England/20142002104/2020

Italy

/SP

L1/2

020

USA/MN-UW264/2020

Hon

gKon

g/H

KP

U45

-050

2/20

20

US

A/W

A-N

H18/2020

Jiujiang/JX22/2020

USA/VA 6171/2020

US

A/N

Y-P

V08

461/

2020

Englan

d/20

1290

4010

2/20

20

Netherlands/N

A 277/2020

Iceland/242/2020

USA/WA-UW-2009/2020

IHUCO

VID-0

092

USA/NY-NYUMC57/2020

USA/WA-UW-2091/2020

Fran

ce/A

RA

1194

9/20

20


Vie

tnam

/CM

295/

2020

USA/ID-UW-4448/2020

England/S

HE

F-C0664/2020

Wal

es/P

HW

C-2

685

7/20

20

Aus

tral

ia/V

IC03

/202

0

USA/NY-PV08453/2020

Iceland/197/2020

HongKong/HKPU57-0702/2020

USA/NY-NYUMC136/2020

Wuhan/IPBCAMS-WH-01/2019

Netherlands/Limburg 10/2020

IHUCOVID-0391

US

A/N

Y-P

V08

429/

2020

Australia/V

IC326/2020

Engl

and/

2009

9072

4/20

20

Aus

tral

ia/V

IC19

0/20

20

Australia/N

SW

27/2020

Luxembourg/LNS4235576/2020

Sco

tland

/CV

R16

9/20

20

Iceland/189/2020

England/NOTT-10E9BB/2020



US

A/C

A4/2020

Wal

es/P

HW

C-2

6927

/202

0

Aus

tralia

/VIC

145/

2020

Australia/V

IC316/2020

Taiw

an/C

GM

H-C

GU

-06/

2020

USA/WI-27/2020

Aust

ralia

/VIC

196/

2020

Icel

and/

246/

2020


Netherlands/N

oordHolland 1/2020

Spain/Valencia79/2020

USA/WA-UW366/2020

Belgium/ULG-10069/2020

Spain/Madrid H12 2107/2020

USA/WA-UW-1998/2020

India/781/2020

IHUCO

VID-0

332

England/CAMB-74711/2020

USA/MN38-M

DH38/2020

USA/W

A-UW

234/2020

Iceland/575/2020

Eng

land

/201

4804

1604

/202

0

Eng

land

/NO

TT-

10E

49C

/202

0

US

A/W

A-N

H5/2020

USA/MN32-M

DH32/2020

Scotla

nd/EDB109/2020

Eng

land

/201

3008

7004

/202

0

Fra

nce/

IDF

0386

-islP

1/20

20

Netherlands/NA 107/2020

Finl

and/

14M

12/2

020

Senegal/306/2020

England/20132001104/2020

USA/NY-NYUMC35/2020

Net

herla

nds/

NA

2/2

020

USA/W

A-UW

297/2020

Wales/PHW

C-24A35/2020

USA/WI-35/2020

Eng

land

/201

3202

2404

/202

0


US

A/W

A-S

93/2020

Por

tuga

l/PT

0069

/202

0

Eng

land

/CA

MB

-748

E1/

2020

Netherlands/NoordBrabant 71/2020

Mala

ysia/

MKAK-C

L-20

20-7

554/

2020

Sw

itzerland/BL0902/2020

Eng

land

/NO

TT-1

0E67

8/20

20

USA/W

A-UW

32/2020



Belgium

/ULG

-6948/2020

Scotland/CVR53/2020

Hangzhou/HZ48/2020

Iceland/565/2020

Wales/PH

WC

-26D94/2020

Por

tuga

l/PT

0083

/202

0

Belgium

/DD

-030593/2020

Belgium/ULG-1

0052/2020

Japan/Hu D

P K

ng 19-031/2020

Belgium/EC-0325128/2020

USA/NY-PV09083/2020

Engl

and/

2013

4064

204/

2020

IHUCOVID-0379

US

A/T

X 2020/2020

US

A/W

A-N

H21/2020

USA

/MA

3699

/202

0

Denm

ark/SS

I-05/2020

US

A/N

Y-N

YU

MC

122/

2020

Iceland/219/2020

USA/CZB-RR057-014/2020

England/20134019402/2020

IHUCO

VID-0

106

Belgium/LPDC-0323115/2020

USA/NY-N

YUMC88/2020

IHUCOVID-0028

England/20102000506/2020

USA/NY-PV09144/2020

Italy

/TE

5780

/202

0

Belgium/BA-02291/2020

US

A/W

A-N

H2/2020

Norway/2087/2020

IHUCOVID-0033

USA/WI-90/2020

Portugal/PT0027/2020

USA/NY-PV09183/2020

Switzerland/GE3121/2020

Iceland/112/2020

Germ

any/N

RW-4

3/20

20

US

A/W

A-U

W-1986/2020

Eng

land

/201

3602

15/2

020

Gua

ngzh

ou/G

ZM

U00

30/2

020

IHU

CO

VID

-012

8

US

A/N

V 0

052/

2020

IHUCOVID-0100

Belgium

/ULG

-10046/2020

Belgium

/LD-030597/2020

Shanghai/S

H01/2020

Belgium/CCM-0324167/2020

US

A/M

A 36

53/2

020

Engl

and/

2013

1102

904/

2020

USA/NY-W

adsworth

-10707-0

1/2020

Belgium

/UM

F-03025/2020

England/20140071404/2020

Engl

and/

CAM

B-74

492/

2020

Portug

al/PT

0079/2020


US

A/W

A7-U

W4/2020

US

A/N

Y-P

V08

401/

2020

US

A/W

A-U

W86/2020


IHU

CO

VID

-0480

USA/WI-66/2020

Por

tuga

l/PT

0075

/202

0

Scotland/CVR20/2020

Engl

and/

2013

6000

1/20

20



Scotland/E

DB

383/2020

England/20130072404/2020

USA/W

A-UW

50/2020

Can

ada/

BC

612

9127

/202

0

Bel

gium

/ULG

-100

82/2

020

US

A/W

A-S

21/2020

IHUCOVID

-0029

Iceland/213/2020

Chongq

ing/IVD

C-C

Q-001

/2020

USA/W

A-UW

218/2020

US

A/W

A-U

W-2203/2020

USA/WA-UW-1732/2020

Iceland/304/2020

Can

ada/

BC

600

4567

/202

0

Belgium/EBN-0323116/2020

Iceland/54/2020

Hungary/49/2020

Netherlands/Lim

burg 5/2020

US

A/W

A-U

W55/2020

England/CAMB-76E56/2020

Italy/UnivPM1/2020

USA/NY-P

V09143/2020


Iceland/253/2020

US

A/W

A-U

W215/2020

USA/PA 1802/2020

Can

ada/

ON

PH

L391

7/20

20

USA/UT-00033/2020

Belgium/DLDJ-030569/2020

Han

gzho

u/Z

JU-0

11/2

020

Aus

tral

ia/N

SW

13/2

020

USA/WA-UW161/2020

USA/W

I-36/2020

Vietnam/CM99/2020

IHUCO

VID-0

432

Por

tuga

l/PT

0033

/202

0

US

A/W

A-U

W-1760/2020

US

A/N

Y-P

V09

067/

2020

Bel

gium

/UG

ent-9

/202

0

Beijing/Wuhan IME-BJ02/2020


Iceland/593/2020

US

A/W

A-S

58/2020

England/20132002004/2020

Wuhan/HBCDC-HB-04/2020


Iceland/27/2020

US

A/W

A-N

H22/2020

England/20140037504/2020

Hangzhou/HZCDC0090/2020

Germany/BAV-V2010840/2020


IHU

CO

VID

-026

9

US

A/W

A-S

50/2020

Wales/PHWC-26620/2020

US

A/W

A-U

W18

5/20

20

US

A/G

A 1621/2020

USA/WI-103/2020

Nether

lands

/NA 16

6/202

0

Wales/PHW

C-24CB7/2020

Wales/P

HWC-2

665D/2020


IHU

CO

VID-0

481

Eng

land

/201

2402

1402

/202

0

Eng

land

/201

3407

8204

/202

0

Iceland/11/2020

USA/WI-70/2020

England/20100004806/2020

Net

herla

nds/

Noo

rdB

raba

nt 2

3/20

20

Aus

tral

ia/N

SW

35/2

020

Finland/14M13/2020

Icel

and/

604/

2020

USA/W

A-UW

302/2020

USA/NY-Wadsworth-10958-01/2020

Engl

and/

2013

6031

8/20

20

Italy

/TE

4953

/202

0

Singapore/31/2020

Canada/M

B 8/2020

USA/NY-PV09190/2020

England/02/2020

Geo

rgia

/Tb-

537/

2020


Wuhan/H

BC

DC

-HB

-01/2019

Netherlands/N

A 286/2020

Belgium/CJE-0325178/2020

Hangzhou/HZ90/2020

Nanchang/JXN3T4/2020

Australia/N

SW

48/2020

Portu

gal/P

T004

3/20

20

USA

/MA

3642

/202

0

Scotland/CVR67/2020

Wal

es/P

HW

C-2

51B

7/20

20

Iceland/90/2020

USA/M

N136-M

DH

136/2020

Germany/NRW-28/2020

US

A/W

A-U

W13

8/20

20

Netherlands/Gelderland 12/2020

Australia/V

IC325/2020U

SA

/WA

-UW

-1294/2020

US

A/M

N42

-MD

H42

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0

Franc

e/M

acon

067

56/2

020

Net

herla

nds/

NA

27/

2020

US

A/W

A-U

W361/2020

Eng

land

/201

4207

9804

/202

0

IHUCOVID-0032

US

A/W

A-S

118/2020

France/HF1795/2020

USA/IA 6391/2020

Kuwait/KU006/2020

USA/UPHL-05/2020

USA/NY-NYUMC48/2020

Iceland/400/2020

USA/WI-47/2020

USA/IA 6401/2020

Nanchang/JX177/2020

England/20139063504/2020


Sin

gapo

re/9

/202

0

US

A/V

A 6382/2020

Wale

s/PHW

35/2

020

Wales/P

HWC-2

4BF6/2020

Wal

es/P

HW

C-2

67B

4/20

20

IHU

CO

VID

-042

8

Taiw

an/C

GM

H-C

GU

-20/

2020

Netherlands/Utrecht 3/2020

Finl

and/

14M

16/2

020

Eng

land

/200

9605

15/2

020

Shang

hai/SH

0126/2020

Wales/P

HW

C-26D

C1/2020

England/20130033104/2020

Japan/Hu Kng 19-437/2020

Netherlands/N

A 171/2020

USA/VA-DCLS-0045/2020

Icel

and/

469/

2020

Taiwan/NTU16/2020

USA

/WA-

UW

-164

5/20

20


Australia/V

IC296

/2020

SouthKorea/KCDC24/2020


US

A/W

A-S

99/2020

Icel

and/

151/

2020

Ger

man

y/N

RW

-35/

2020

Australia/N

T05/20

20

Eng

land

/SH

EF-

BFE

06/2

020

Eng

land

/201

3402

5204

/202

0

IHUCOVID-0329


US

A/W

A-U

W284/2020

Eng

land

/201

3600

37/2

020

Iceland/33/2020

Wales/PHW27/2020

Iceland/548/2020

Wal

es/P

HW

C-2

6E91

/202

0

USA/WA-UW-1422/2020

Iceland/137/2020

US

A/P

A 43

95/2

020

US

A/W

A-U

W320/2020

USA/WA-UW-1425/2020

USA/NY-W

adsworth

-10528-01/2020


US

A/W

A-U

W106/2020

England/20129035304/2020

US

A/N

Y-P

V09

116/

2020


Australia/V

IC37/20

20Bel

gium

/ULG

-973

5/20

20

Engla

nd/C

AMB-7

6D3B

/202

0

USA/NY-PV08492/2020


IHUCOVID-0337

Netherlands/N

A 272/2020


Iceland/97/2020

IHUCO

VID-0

361


USA/N

Y-PV08495/2020

Japa

n/P

4-4/

2020

Net

herla

nds/

NA

47/

2020

Netherlands/N

A 92/2020

Portugal/PT0006a/2020

Japa

n/P

2-1/

2020

England/20134078904/2020

USA/NY-NYUMC65/2020

Net

herla

nds/

NA

212/

2020

USA/NY-NYUMC13/2020

France/IDF2684/2020

Eng

land

/201

3808

2704

/202

0


IHUCO

VID-0

423

Engl

and/

2013

6104

0/20

20

Netherlands/Z

uidHolland 15/2020

US

A/W

A-U

W128/2020

Eng

land

/201

3904

5904

/202

0

IHUCOVID

-039

7

US

A/V

A-D

CLS

-0010/2020

Engl

and/

2013

2027

404/

2020

Australia/WA06/2020


Wales/P

HWC-24BE7/2020

Aus

tralia

/VIC

127/

2020

IHU

CO

VID

-018

6

England/20130052604/2020


England/20142002204/2020

Hon

gKon

g/V

M20

0025

07/2

020

Brazil/S

PB

R-03/2020

France/IDF0626/2020

Austra

lia/V

IC23

4/20

20

Icel

and/

174/

2020

Net

herla

nds/

NA

215

/202

0

Net

herla

nds/

Noo

rdB

raba

nt 2

0/20

20

Iceland/81/2020

Netherlands/N

A 250/2020

Shanghai/SH0009/2020


IHU

CO

VID

-037

6

Canada/ON PHL1095/2020



HongK

ong/

VB2001

7970

/202

0

Aus

tral

ia/V

IC90

/202

0

Canada/BC 6502001/2020


Germany/NRW-42/2020

US

A/M

N69-M

DH

69/2020

Sco

tland

/CV

R23

6/20

20


Iceland/383/2020

Hangzhou/HZCDC0091L/2020

Taiwan/C

GMH-CGU-01/2020

US

A/U

T 8199/2020

Shanghai/S

H0093/2020


Australia/NSW08/2020

Australia/Q

LDID

931/2020

Engl

and/

2012

9036

504/

2020

Aus

tral

ia/V

IC30

/202

0

Aus

tralia

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70/2

020

Wal

es/P

HW

C-2

6C00

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0

Engl

and/

2013

9024

904/

2020

Netherlands/N

A 138/2020

Canada/BC 6489864/2020

Germ

any/N

RW-0

4/20

20


Iceland/179/2020

USA/NY-W

adsworth

-10935-01/2020

USA/WI-GMF-00588/2020

US

A/W

A-S

44/2020

Eng

land

/200

9900

02/2

020

Wales/PHWC-2713D/2020


Icel

and/

449/

2020

England/20139046704/2020

Fran

ce/B

1623

/202

0

Iceland/505/2020

Iceland/476/2020

USA/NY-PV08426/2020

Engla

nd/2

0134

0275

04/2

020

USA/CT-U

W-4

347/2020

Indi

a/11

15/2

020

Han

gzho

u/ZJ

U-0

1/20

20

USA/WA-UW-1954/2020

Belgium/W

AM-030676/2020

US

A/W

A-U

W173/2020


Malaysia/MKAK-CL-2020-5045/2020

IHU

CO

VID

-038

1

Canada/O

N P

HL3877/2020

USA/W

A-UW

-1828/2020

US

A/W

A-N

H24/2020

IHUCOVID

-048

9

IHU

CO

VID

-029

1

USA/W

I-26/2020

Iceland/556/2020


Wales/PHW

C-25E47/2020

USA/NY-PV09180/2020

Wal

es/P

HW

C-2

6918

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0


US

A/W

I-19/

2020

Netherlands/Z

uidHolland 1/2020

Canada/O

N P

HL0142/2020


Scotland/CVR107/2020

Engl

and/

2013

2003

004/

2020

France

/HF3036/2020

USA/WA 5030/2020

Icel

and/

252/

2020

US

A/W

A6-U

W3/2020

England/20130034604/2020


Belgium

/DA

L-030650/2020


Aus

tral

ia/V

IC31

0/20

20


Pingx

iang

/JX5/

2020

Japan/TY-WK-012/2020

Wales/P

HW

C-2617A

/2020

Australia/QLDID923/2020

Net

herla

nds/

NA

73/

2020

Scotland

/ED

B31

8/2020

IHUCOVID

-044

7

Norway/1917/2020



Aus

tral

ia/V

IC23

3/20

20

France/CVL3365/2020

Taiwan/107/2020


Eng

land

/201

3210

0204

/202

0

US

A/W

A-U

W13

3/20

20

USA/NY-NYUMC15/2020

HongKong/HKPU1 2101/2020

Mexico/C

DM

X-InD

RE

01/2020

Englan

d/20

1380

277/

2020

IHU

CO

VID

-043

8

Iceland/507/2020

Portugal/P

T0029/2020


Engl

and/

2013

6023

6/20

20

Icel

and/

275/

2020

Iceland/524/2020


Spain/Galicia201663/2020

Netherlands/N

A 7/2020

US

A/W

A-S

18/2020

Iceland/506/2020

Rus

sia/

StP

eter

sbur

g-35

24/2

020

Poland

/PL

P1/20

20


USA/NY-PV09043/2020

Canada/B

C 3989992/2020


US

A/W

A-S

28/2020

Iceland/432/2020

Netherlands/N

A 150/2020

Eng

land

/201

0909

4206

/202

0

Belgium/KKN-0319184/2020

Luxe

mbo

urg/

LNS

0945

359/

2020

Ger

man

y/B

AV-V

2011

395/

2020

Shenzhen/S

ZT

H-003/2020

Indi

a/16

16/2

020

Sw

itzerland/VD

5615/2020

Scotland/C

VR

32/2020


Wales/P

HW

C-2668A

/2020

France/IDF3235/2020

Aus

tralia

/VIC

218/

2020

Netherlands/N

A 282/2020

Icel

and/

535/

2020

Netherlands/NoordBrabant 9

7/2020

DR

C/K

N-0

060/

2020

Netherlands/Z

uidHolland 83/2020

Belgium/RA-030445/2020

Netherlands/Friesland 4/2020

Icel

and/

115/

2020

USA/WA-UW-2016/2020


England/20129031804/2020

USA/WI-77/2020

Shang

rao/

JX12

15/2

020

USA/WI-04/2020

USA/WA-UW-2131/2020

Spain/V

H198152683/2020

USA/NY-PV09198/2020


USA/MN43-MDH43/2020

USA/CT-UW-4344/2020



USA/WA-UW-1801/2020

Spain/Madrid R2 15/2020



Hon

gKon

g/H

KP

U41

-080

2/20

20


England/20139002204/2020

Scotland/E

DB

057/2020

Iceland/475/2020

US

A/W

A-U

W147/2020

Engl

and/

2010

9056

906/

2020

USA/NY-PV09098/2020

Sco

tland

/CV

R15

5/20

20

Scotland/CVR60/2020

US

A/W

A-S

34/2020


Scotland/C

VR

110/2020

Belgium/MSE-0324131/2020

Sin

gapo

re/3

8/20

20

USA/CT-UW224/2020

Australia/V

IC41/2020

Australia/V

IC251

/2020

Net

herla

nds/

Noo

rdB

raba

nt 5

6/20

20

Iceland/101/2020

Belgium/N

JC-0324126/2020




USA/NY-PV09046/2020

USA/W

A-UW

-1279/2020

England/201380070/2020

Fran

ce/H

F168

4/20

20

IHU

CO

VID

-025

2

Wal

es/P

HW

C-2

43D

0/20

20

Netherlands/N

A 156/2020

Eng

land

/201

3201

8904

/202

0

USA/W

A-UW

112/2020

Spain/PaisVasco201607/2020

Canada/BC 7515663/2020



Shanghai/S

H0050/2020

US

A/N

Y-P

V08

450/

2020

USA/W

A-S90/2020

Nanchang/JX90/2020

Icel

and/

8/20

20

Aus

tral

ia/V

IC13

5/20

20

Italy/TE

5512/2020

Aust

ralia

/VIC

-CBA

2/20

20

Iceland/218/2020

USA/CA-S

CCPHD-UC14/2020

Fujian/8/2020



Wales/PHW

C-2631A/2020

US

A/W

A-S

65/2020

Iceland/552/2020

Iceland/212/2020

USA/NY-NYUMC37/2020

Austria/CeMM0011/2020

Australia/N

SW

50/2020

Beijing/W

uhan IME-B

J05/2020

Netherlands/N

A 56/2020

Australia/V

IC188

/2020

US

A/N

Y-N

YU

MC

126/

2020

Icel

and/

178

/202

0

US

A/W

A-U

W253/2020

Wales/P

HW

C-26D

3A/2020

Taiwan/125/2020

Scotland/CVR66/2020

US

A/W

A-S

54/2020

Eng

land

/201

2803

7406

/202

0

US

A/W

A-U

W-1448/2020

USA/NY-NYUMC46/2020

Scotland/C

VR

152/2020

USA/NY-PV08481/2020

Eng

land

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MB

-737

8B/2

020

Eng

land

/201

4400

2904

/202

0

USA/W

A-UW

172/2020

Netherlands/U

trecht 1/2020

USA/MN40-MDH40/2020

USA/WI-44/2020

USA/NY-PV09138/2020

USA/NY-P

V0900

6/20

20


Eng

land

/201

3006

3604

/202

0

England/20130049704/2020

England/20130065304/2020

Aus

tral

ia/V

IC55

/202

0

Iceland/493/2020

Eng

land

/201

4202

4704

/202

0

Aus

tral

ia/V

IC31

1/20

20


USA/NY-PV08434/2020

Taiwan/NTU21/2020

Australia/V

IC280/2020

Kor

ea/B

A-A

CH

271

8/20

20

Iceland/126/2020

Hangzhou/H

Z162/2020


Iceland/490/2020

US

A/W

A-U

W62/2020

Indi

a/11

04/2

020

USA/NH 0029/2020

Scotland/E

DB

168/2020

USA/W

A-UW

64/2020

Belgium/HP-0319108/2020

US

A/F

L 5091/2020

Net

herla

nds/

NA

198/

2020

IHUCOVID-0409

Aus

tral

ia/V

IC-C

BA

1/20

20


Eng

land

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EF-

C05

85/2

020

USA/MN129-MDH129/2020

Engl

and/

2013

4060

804/

2020

Shanghai/S

H0060/2020

Iceland/286/2020



USA/UT-00349/2020

US

A/W

A-U

W-1327/2020


Wal

es/P

HW

C-2

5B6E

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0

USA/WA-UW288/2020

US

A/W

A-U

W63/2020

Iceland/65/2020

Iceland/447/2020

USA/IN

-Lilly-IPB0170-4bc21/2020

England/20138017804/2020

Italy/FVG-ICG

EB S8/2020

Australia/V

IC240/2020

Engla

nd/2

0134

0211

04/2

020

Sco

tland

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R97

/202

0


Australia/V

IC24/2020

IHUCOVID

-023

6

Australia/V

IC143/2020

Netherlands/Limburg 9/2020



US

A/W

A-S

66/2020


USA/W

A-S96/2020

Icel

and/

470/

2020

USA/UT-0

0297/2020

USA/WI-109/2020


IHUCOVID

-010

8

Iceland/91/2020

USA/NY-PV09020/2020

US

A/W

A-S

63/2020

Eng

land

/201

3005

2004

/202

0

Hon

gKon

g/H

KP

U83

-170

2/20

20

Can

ada/

BC

024

21/2

020

IHUCOVID

-0050


Italy/TE5166/2020


Netherlands/Z

uidHolland 22/2020


US

A/W

A-U

W385/2020

Australia/V

IC118/2020

USA/W

A-UW

229/2020

USA/WI-112/2020

Scotland/E

DB

234/2020


IHU

CO

VID

-030

6


IHUCOVID

-0203

France/IDF2284/2020

Canada/O

N P

HL4464/2020

IHUCOVID-0075

Icel

and/

292/

2020

Eng

land

/201

4407

3804

/202

0

Iceland/438/2020

Canada/BC 4799711/2020

US

A/W

A-S

103/2020

Iceland/24/2020

US

A/T

X 2

817/

2020

Wales/P

HW

C-26787/2020

Wal

es/P

HWC-2

5E65

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0

Engla

nd/2

0148

0061

04/2

020

US

A/W

A-U

W-1700/2020


Sou

thK

orea

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MC

01/2

020

Netherlands/U

trecht 13/2020

Iceland/152/2020

USA/WI-05/2020

USA/IL2/2020


Nor

way

/149

3/20

20

US

A/N

Y-P

V08

489/

2020

Eng

land

/CA

MB

-74E

85/2

020

USA/WA-UW

-2228/2020




Belgium

/UG

ent-6/2020

US

A/N

Y-PV

08124/2020

US

A/W

A-U

W97/2020

England/20108006003/2020

Engla

nd/20129171604

/2020

Germany/NRW-48/2020

England/20132068204/2020

Russia/S

tPetersb

urg-R

II3997/2020

USA/NY-PV08443/2020

Net

herla

nds/

NA

48/

2020

Iceland/244/2020

Ger

man

y/N

RW

-46/

2020

Japa

n/P

3-1/

2020


Slo

veni

a/80

8/20

20

IHU

CO

VID

-0485

England/CAMB-739D0/2020

England/20140009404/2020

USA/NY-NYUMC8/2020



England/20110023706/2020

Indi

a/16

21/2

020

Icel

and/

131/

2020

England/20144003804/2020

USA/NJ-NYUMC84/2020


US

A/M

N25

-MD

H25

/202

0

Taiwan/CGMH-CGU-17/2020

Germany/NRW-32/2020

US

A/W

A2/2020

Eng

land

/201

4201

5104

/202

0

Netherlands/N

A 16/2020

IHUCOVID

-023

4


Engla

nd/2

0136

0538

/202

0

France/Bourg-en-Bresse 06678/2020

Scot

land

/CVR

07/2

020

USA/WA-UW-2238/2020

US

A/W

A-N

H17/2020

USA/RI 0702/2020


Iceland/150/2020

Hon

gKon

g/H

KP

U48

-020

2/20

20

Wales/PHW

C-2680C/2020

USA/NY-PV09039/2020

Net

herla

nds/

NA

238

/202

0

US

A/W

A-U

W-1835/2020

Iceland/264/2020

Sin

gapo

re/5

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0

US

A/W

A-U

W-1729/2020


IHUCOVID

-0411

IHUCO

VID-0

262

Por

tuga

l/PT

0082

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0

Wales/P

HW

C-26D

49/2020

Sco

tland

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B27

0/20

20

Netherlands/N

oordHolland 3/2020

Wal

es/P

HW

C-2

6ED

D/2

020

USA/SC 6370/2020

US

A/W

A-S

106/2020

Scotland/C

VR

49/2020

Bel

gium

/ULG

-645

7/20

20

Iceland/161/2020

Taiw

an/NTU17/2020

Wales/PHW

C-2663F/2020

Eng

land

/201

3610

08/2

020

IHUCO

VID-0

334


Netherlands/ZuidH

olland 28/2020


IHUCOVID-0242

Iceland/221/2020

US

A/W

A3-

UW

1/20

20

USA/NY-PV09088/2020

Iceland/300/2020

Fra

nce/

AR

A12

877/

2020

Canad

a/ON P

HL367

0/20

20

Iceland/20/2020


US

A/W

A-U

W-1

586

/202

0

Fran

ce/A

RA1212

5/20

20

USA/NY-NYUMC14/2020

US

A/N

Y-PV

08122/2020


USA/WA-UW-2094/2020


USA

/WI-2

5/20

20

Scotland/C

VR

33/2020

Belgium/GOF-030578/2020

USA/WI-57/2020

Net

herla

nds/

NA

242/

2020


USA/WA-UW-1570/2020


US

A/W

A-S

70/2020

Iceland/136/2020


Iceland/153/2020


Wales/PHW

C-2682A/2020

France/B

2344/2020

IHUCO

VID-0

122

Wales/PHWC-26F34/2020

Sco

tland

/ED

B04

7/20

20

US

A/W

I-14/

2020

Icel

and/

511/

2020

USA/NY-PV08456/2020

England/20144011504/2020

Taiwan/NTU02/2020

Engl

and/

2013

2021

004/

2020

Wales/P

HWC-2

4455/2020

Engl

and/

CAM

B-73

A19/

2020




Iceland/144/2020

Aust

ralia

/VIC

172/

2020


USA/WA-UW-1706/2020

USA/NY-NYUMC30/2020

DR

C/8

1/20

20

Iceland/267/2020

Eng

land

/201

3808

2004

/202

0

IHU

CO

VID-0054

Fra

nce/

AR

A10

251/

2020


Icel

and/

597/

2020



Shangrao/JX105/2020

Iceland/456/2020

Belgium

/RG

E-0325122/2020


Wales/PHWC-260D7/2020

Fran

ce/A

RA

1226

0/20

20

IHUCOVID

-0153

Canada/ON P

HL4232/2020


Germany/NRW-25/2020

England/20104003002/2020

Singap

ore/

1/20

20

Netherlands/N

A 19/2020

Wales/P

HWC-26A42/2020

Aust

ralia

/VIC

202/

2020

Eng

land

/NO

TT-

10D

F23

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0

USA/CT-UW260/2020



Australia/V

IC59/20

20


Scotland/CVR15/2020

Engl

and/

2013

0053

004/

2020

Scotland/E

DB

561/2020


England/20144004704/2020

USA/WI-73/2020

USA/WI-68/2020

Belgium

/ULG

-100

28/20

20

USA/WI-07/2020

Wales/P

HW

2/2020

Eng

land

/CA

MB

-746

23/2

020

Wale

s/PHW

C-240

06/2

020

Panama/328677/2020

Iceland/596/2020

England/20139064904/2020

Engl

and/

2013

9031

504/

2020

USA/NY-NYUMC56/2020


Portugal/P

T0030/2020



Canada/BC 6404889/2020

Italy/UniSR1/2020

US

A/N

Y-P

V09

061/

2020

US

A/W

A-U

W-1588/2020


US

A/W

A-U

W156/2020

USA/NY-PV09163/2020


Bel

gium

/ULG

-100

72/2

020

Fra

nce/

IDF

3029

/202

0

Iceland/80/2020

US

A/M

A 1

355/

2020

Engl

and/

2013

8002

4/20

20

USA/WI-G

MF-00921/2020


USA/W

A-UW

139/2020

IHUCO

VID-0

490

Han

gzho

u/Z

JU-0

9/20

20

USA/CZB-RR057-011/2020

USA/ID-UW-1925/2020


Aus

tral

ia/V

IC26

2/20

20

Belgium

/LJ-0325148/2020

IHUCO

VID-0

196

France/IDF2414/2020

USA/WA-U

W-1930/2020

Fran

ce/ID

F334

5/20

20

Spa

in/M

adrid

LP

14 3

/202

0

France/A

RA

10282/2020

Norway/1953/2020


USA/NY-N

YUMC107/2020

Netherlands/Z

uidHolland 26/2020

Net

herla

nds/

NA

223

/202

0

Georgia

/Tb-152

3/2020

IHU

CO

VID

-040

5

IHUCOVID-0093


USA/NC 200173/2020

Engl

and/

2013

2024

604/

2020

Shandong/LY008/2020

Eng

land

/201

3200

0904

/202

0

Iceland/128/2020

US

A/W

I-09/2020

USA/WA-UW291/2020

USA/ID-UW-2159/2020

Engl

and/

2014

0064

504/

2020

Iceland/508/2020

USA/NY-NYUMC97/2020









US

A/W

A-U

W369/2020

Wuh

an/W

IV04

/201

9

Australia/N

T02/2020


Japa

n/P

1/20

20

Net

herla

nds/

NA

79/

2020

IHUCOVID

-036

5



Bel

gium

/ULG

-100

14/2

020

IHUCOVID-0021

Iceland/402/2020


Aus

tria/

CeM

M00

07/2

020

Hangzhou/HZ60/2020

Wales/PHWC-26ECE/2020


Bel

gium

/ULG

-100

12/2

020

Singap

ore/

26/2

020

US

A/W

A-U

W-1682/2020

USA/NY-PV08490/2020


Iceland/93/2020

Wal

es/P

HW

C-2

6C88

/202

0

IHUCOVID-0281

IHUCOVID-0342

US

A/W

A-U

W-1744/2020

Iceland/124/2020

USA/WA-UW-1762/2020

USA/WI-100/2020

IHU

CO

VID

-0351

DR

C/241/2020


Wuhan/H

BC

DC

-HB

-05/2020

Net

herla

nds/

NA

59/

2020

Australia/V

IC130/2020

USA/WA-UW-2208/2020

Bel

gium

/ULG

-100

67/2

020

England/C

AM

B-73A

46/2020

Italy/FVG-ICG

EB S9/2020

Icel

and/

595/

2020

Iceland/79/2020

NanChang/JX216/2020

Taiw

an/N

TU20/2

020

Brazil/B

A-312/2020

Aus

tral

ia/V

IC04

/20

20

England/200990723/2020

Eng

land

/201

3200

5704

/202

0

Iceland/390/2020

Eng

land

/201

3905

5204

/202

0

Net

herla

nds/

NA

34/

2020

USA/NY-NYUMC86/2020

Scotland/C

VR

213/2020

Iceland/235/2020

Engla

nd/2

0138

0071

04/2

020

Por

tuga

l/PT

0038

/202

0

IHU

CO

VID

-010

4

Korea/KCDC2002/2020

Sco

tland

/ED

B22

6/20

20

Spain/CastillaLaMancha201328/2020


IHUCOVID-0328

Scotland/C

VR

38/2020



Iceland/37/2020

Engla

nd/2

0140

0022

04/2

020

Belgium/CD-030679/2020

USA/WA-UW

120/2020


IHUCO

VID-0

246

France/IDF2278/2020

France/IDF3276/2020

Icel

and/

414/

2020

USA/WI-50/2020

Iceland/359/2020

USA/ID-UW-4396/2020

Sw

itzerland/GE

9586/2020

Aust

ralia

/VIC

186/

2020

Eng

land

/201

3902

1104

/202

0



USA/NY-PV09077/2020

US

A/W

A-S

36/2020

France/IDF3324/2020


USA/WA-UW210/2020

Nanchan

g/JX176/20

20

USA/WI-79/2020

USA/NY-P

V08403/2020

IHUCO

VID-0

499

IHUCO

VID-0

333

Canada/BC 7553799/2020

Iceland/514/2020

Portugal/C

V62/2020

Shanghai/S

H0055/2020

USA/MN130-MDH130/2020

Canada/O

N P

HL7590/2020

Iceland/158/2020


IHUCOVID

-043

1

Wales/P

HW

C-26F

07/2020

France/H

F3138/2020

Wale

s/PHW

C-268

75/2

020

IHUCOVID

-001

7

Netherlands/N

A 317/2020

Aus

tralia

/VIC

105/

2020

US

A/N

Y-N

YU

MC

47/2

020


USA/NY-P

V09135/2020

Kuw

ait/K

U09

/202

0


US

A/W

A-U

W211/2020

Eng

land

/CA

MB

-76F

62/2

020

Iceland/96/2020

Han

gzho

u/H

ZC

DC

0025

/202

0


USA/NY-PV09021/2020

Mexico/CDMX-INER 04/2020

Belgium

/MC

-030689/2020

Austra

lia/V

IC15

8/2020

Can

ada/

ON

PH

LH64

15/2

020

Wales/PHWC-25F9F/2020

US

A/W

A-S

39/2020

Taiw

an/C

GM

H-C

GU

-08/

2020

Hangzhou/H

Z178/2020

Wales/P

HWC-2

6EFB/2020

USA/N

Y-NYU

MC

4/2020

USA/WA-UW-2212/2020


Colombia/Antioquia79256/2020

US

A/W

A-U

W201/2020

Iceland/431/2020

USA/WA-UW-1921/2020

Guangdong/20SF201/2020


Fran

ce/A

RA

1284

/202

0

IHUCOVID-0400

Iceland/477/2020

USA/WA-UW233/2020

Belgium/R

C-0324168/2020

Belgium/HL-030771/2020

Belgium/U

Gent-24/2020

USA/WA-UW82/2020

USA/WI-78/2020

Iceland/22/2020

Sco

tland

/CV

R46

/202

0

Net

herla

nds/

NA

57/

2020


USA/NY-PV08121/2020

Wales/PHW

C-26F8F/2020


Can

ada/

ON

PH

L097

7/20

20

Iceland/388/2020

USA/OH 0023/2020

Aus

tralia

/VIC

146/

2020

Iceland/165/2020


USA/V

A-DC

LS-0

020/

2020

Iceland/193/2020

US

A/W

A-U

W-1315/2020

Eng

land

/201

4800

4704

/202

0


Netherlands/H

aarlem 1363688/2020

Portugal/P

T0026/2020

Italy/INMI3/2020

Aus

tral

ia/V

IC20

8/20

20


USA/WA-UW-1478/2020

Italy/TE4836/2020

HongKong/HKPU60-0802/2020

Eng

land

/201

3602

06/2

020

USA/WI-93/2020

Iceland/192/2020

Scotland/CVR91/2020


USA

/MA

0020

/202

0

USA/W

A-S73/2020

Eng

land

/201

4403

8204

/202

0

US

A/W

A-U

W32

4/20

20


USA/NY-NYUMC82/2020

IHUCO

VID-0

266

Eng

land

/201

2909

6604

/202

0


USA/W

A-S81/2020

USA/WA-UW-1322/2020

Sco

tland

/ED

B04

8/20

20

USA/NY-PV08491/2020

USA/WA-UW110/2020

Belgium/LV-0324133/2020

Iceland/12/2020

USA/WY-WYPHL006/2020

Wales/PHW

C-24279/2020


USA/AZ1/2020

Belgium/RS-030677/2020


Por

tuga

l/PT

0050

/202

0


US

A/W

A-U

W174/2020

USA/NY-NYUMC74/2020

Australia/N

SW

02/2020

England/20124036306/2020

Italy/FVG-ICGEB S5/2020


Wal

es/P

HW

C-2

649F

/202

0

Scot

land

/CVR

163/

2020

US

A/W

A-S

57/2020

USA/WI-48/2020


England/SHEF-C

042E/2020

US

A/W

A-U

W51/2020

Taiw

an/C

GM

H-C

GU

-16/

2020

Netherlands/ZuidH

olland 98/2020

Canada/O

N P

HL2653/2020

Norway/2

114/2020



IHUCO

VID-0

287

USA/WI-65/2020

IHU

CO

VID

-044

1



Por

tuga

l/PT

0048

/202

0

Wale

s/PHW

C-25A

52/2

020

IHU

CO

VID

-0012

Mexico/Queretaro-InDRE 04/2020

Eng

land

/201

3602

81/2

020

IHUCOVID-0433

Iceland/513/2020


England/20132000304/2020


USA/VA 6389/2020


Engl

and/

SHEF

-C00

B1/2

020

Aus

tral

ia/V

IC12

1/20

20

US

A/W

A-U

W373/2020

USA/ID-UW-2255/2020

Iceland/463/2020



Belgium/D

JMG

-0325156/2020

Sco

tland

/ED

B03

3/20

20


Italy

/TE

6195

/202

0


Scotland/CVR30/2020

Wal

es/P

HW

C-2

6C1F

/202

0

Australia/V

IC115/2020

USA

/MA

4811

/202

0

IHUCOVID-0073

Germ

any/N

RW-0

9/20

20


USA/NJ-NYUMC83/2020

France/HF1645/2020

Iceland/248/2020

Ireland/D

ublin-22428/2020

US

A/W

A-U

W-1271/2020


Fran

ce/H

F239

3/20

20


IHU

CO

VID

-031

6

Norway/2

084/2020

Iceland/418/2020

Portugal/CV63/2020

Australia/WA02/2020

Hon

gKon

g/H

KP

U64

-120

2/20

20

Por

tuga

l/PT

0070

/202

0

USA/WA-UW160/2020

USA/WI-61/2020

Belgium

/VI-03027/2020

USA/W

A-UW

28/2020

IHU

CO

VID

-0188

Germany/NRW-38/2020


Scotland/CVR50/2020

IHUCOVID

-008

1

Ireland/21023/2020

Wales/P

HW

C-2666C

/2020


USA/NY-PV09073/2020


USA/W

A-UW

66/2020

Eng

land

/201

4407

4804

/202

0

Netherlands/N

A 36/2020

Belgium/GMJ-0324132/2020

Iceland/247/2020

Italy

/619

3/20

20

US

A/W

A-U

W154/2020

Iceland/322/2020

USA/NY-NYUMC29/2020

US

A/W

A-S

108/2020


Georgia/Tb-390/2020


USA/GA 1847/2020

US

A/V

A-D

CLS

-0017/2020

USA/NY-PV09122/2020

Por

tug

al/P

T00

22/2

020


HongK

ong/VM

20002849/2020

USA/C

A-CD

PH-U

C1/2020

IHU

CO

VID

-019

8

England/NOTT-10E0B6/2020

England/20142003904/2020

Brazil

/SPBR-11

/2020


Por

tuga

l/PT

0094

/202

0

IHU

CO

VID

-002

4

Engl

and/

2013

2106

004/

2020

Wales/PHWC-26BC7/2020


Belgium

/VW

JM-0324136/2020

DRC/KN-0072/2020


Iceland/181/2020

US

A/W

A-U

W340/2020

Ireland/21145/202

0


Netherlands/Zeewolde 1365080/2020

Germany/NRW-31/2020


Japan/UT-NCGM02/2020

USA/WA-UW-1866/2020




Belgium/GL-030546/2020

Portug

al/PT

0062/2020

USA/NY-NYUMC18/2020

Engla

nd/2

0129

0338

04/2

020

USA/NY-PV09115/2020

US

A/W

I-GM

F-00228/2020

Can

ada/

BC

944

6031

/202

0

Belgium/SJ-030583/2020


England/20130049604/2020

Eng

land

/201

3906

5404

/202

0

England/201340875/2020


France/HF2306/2020


Shanghai/S

H0058/2020

Portugal/P

T0058/2020

USA/NY-N

YUMC108/2020

Austra

lia/V

IC10

1/20

20


IHU

CO

VID

-008

0

Shangrao/JX

1176/2020

Iceland/334/2020

Icel

and/

424/

2020

USA/WA1/2020


USA/NY-PV09121/2020



Sco

tland

/CV

R12

6/20

20

US

A/V

A-D

CLS

-0018/2020

Netherlands/Lim

burg 3/2020

Sco

tland

/ED

B37

4/20

20

Shanghai/S

H0106/2020

France/Pollionay 1733/2020

US

A/W

A-U

W-1863/2020

England/20110003506/2020

Eng

land

/201

3210

6704

/202

0

Shanghai/S

H0020/2020

Belgium/DCS-030796/2020

IHU

CO

VID

-0407


England/20146019704/2020


Can

ada/

ON

PH

L875

1/20

20

Sw

itzerland/GE

1422/2020


Ireland/Limerick-19935/2020


Belgium

/RA

-030664/2020

Iceland/201/2020

USA/WA-UW-1611/2020


US

A/W

I-21/

2020

DRC/253/2020

Engl

and/

2009

4052

7/20

20

Netherlands/N

A 49/2020

US

A/W

A-S

87/2020

Fran

ce/A

RA

1019

2/20

20

USA/W

A-UW

126/2020

England/01/2020

England/20108007002/2020

France/Valence 532/2020

France/B2336/2020


England/S

HE

F-C

0321/2020

Engl

and/

2014

2053

204/

2020

Singapore/4/2020

USA/NY-PV09153/2020

Iceland/464/2020

USA/ID-UW-1934/2020

England/CAMB-738A6/2020

Canada/BC 8897642/2020

Brazil/RJ-314/2020

Wal

es/P

HWC-2

5418

/202

0

Belgium/RDC-030961/2020

US

A/W

A-U

W309/2020

Netherlands/N

oordBrabant 2/2020

Eng

land

/201

0800

6802

/202

0


Eng

land

/201

3906

2804

/202

0

Engl

and/

2012

8036

304/

2020

Belgium/ULG-1

0095/2020

Shang

hai/SH

0022/2020

USA

/MA

3614

/202

0

Netherlands/N

A 91/2020

IHU

CO

VID

-0141

Icel

and/

331/

2020

US

A/N

Y-W

adsw

orth

-110

00-0

1/20

20

Norway/1772/2020

Iceland/472/2020


USA/NY-PV08402/2020

Hon

gKon

g/H

KP

U32

040

2/20

20

USA/AK-PHL077/2020

England/20142053804/2020

Guangdong/2020XN4433-P0040/2020

US

A/W

A-U

W67/2020


Hangzhou/H

ZC

DC

0012/2020

England/20124049402/2020

Iceland/577/2020

Australia/V

IC67/2020

US

A/W

A-U

W165/2020

Germany/NRW-27/2020

Belgium/NML-0325127/2020

Wales/P

HW

C-257F

1/2020



Franc

e/Bou

rg-e

n-Bre

sse

0681

3/20

20

USA/WA-UW203/2020

Senegal/073/2020

Sco

tland

/CV

R12

5/20

20

Portugal/P

T0016/2020

USA/NY-N

YUMC77/2020

US

A/W

Y-WY

PH

L012/2020

Engla

nd/2

0129

0161

04/2

020



Sw

itzerland/GE

1402/2020

Iceland/444/2020

USA/NY-PV08427/2020

US

A/M

N56-M

DH

56/2020

Finl

and/

13M

65/2

020

IHUCOVID-0060

Belgium

/MTR

-03021/2020

USA/NY-PV08477/2020

US

A/N

Y-N

YU

MC

159/

2020

Nonthaburi/74/2020

Scotland/E

DB

038/2020


Sin

gapo

re/7

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0

Engl

and/

2013

2035

604/

2020

Iceland/592/2020



US

A/W

A-U

W279/2020

USA/W

A-UW

-1905/2020


US

A/N

Y-NY

UM

C112/2020

Senegal/620/2020

IHUCOVID

-0035

IHUCO

VID-0

095

IHU

CO

VID

-021

8


Netherlands/NA 11

2/2020

USA/WA-UW-2219/2020

Wales/P

HWC-23D6A/2020

Shang

hai/S

H0109

/202

0

USA/NY-PV09158/2020

Iceland/9/2020


Belg

ium

/DVB

J-03

0468

/202

0

USA/NY-PV09000/2020


Canada/O

N PH

L3458/2020



Netherlands/Z

uidHolland 19/2020

Shang

hai/S

H0011

/202

0

USA/U

T-00020/2020

Icel

and/

116/

2020


Sco

tland

/CV

R45

/202

0

Iceland/398/2020

Wales/P

HWC-2

5524/2020

USA/W

A-UW

-1944/2020

Latvia/0003/2020


Fran

ce/A

RA

1195

0/20

20

Nanchang/JX174/2020

Spain/Madrid201709/2020

Germany/NRW-26/2020

Eng

land

/201

3008

6204

/202

0

US

A/A

Z 8132/2020

Nor

wa

y/13

80/2

020

Germany/N

RW-1

6/2020

Iceland/103/2020

Bel

gium

/ULG

-100

80/2

020


3/2020


DRC/376/2020

Iceland/280/2020

Eng

land

/201

4206

2604

/202

0

Fosha

n/20

SF211/

2020

Australia

/VIC

246/2020

US

A/W

A-U

W88/2020

USA/UT-00139/2020

Kor

ea/B

A-A

CH

271

9/20

20

France/H

F2601/2020

Wuh

an-H

u-1/

2019

Eng

land

/201

4406

1104

/202

0


Net

herla

nds/

NA

190

/202

0

Aus

tralia

/VIC

80/2

020

Shangrao/JX29/2020

Germany/NRW-18/2020


IHU

CO

VID

-021

9

Canad

a/ON P

HL569

4/20

20

Icelan

d/16

6/20

20

USA/SC 3520/2020


Belgium

/GM

H-03022/2020

Taiw

an/C

GM

H-C

GU

-10/

2020



USA/NY-W

adsw

orth

-106

90-0

1/20

20


USA/NY-PV08473/2020

Icel

and/

471/

2020

USA/NY 2929/2020

USA/NJ-PV09072/2020

Wales/P

HW

C-26E

46/2020

US

A/W

A-U

W46/2020

Brazil/SPBR-05/2020

US

A/N

Y-P

V09

307/

2020

US

A/W

A-U

W99/2020

Aus

tral

ia/N

SW

06/2

020

Shanghai/S

H0031/2020

Engla

nd/2

0139

0575

04/2

020

France/IDF3163/2020

Hon

gKon

g/H

KP

U89

-250

2/20

20

IHUCOVID-0056

Wales/P

HW

04/2020

SouthAfrica/R03006-20/2020

Fran

ce/A

RA

1096

8/20

20

Icel

and/

357/

2020


Icel

and/

135/

2020

Sco

tland

/CV

R35

/202

0

England/20134047404/2020

Wuh

an/IV

DC

-HB

-01/

2019


Netherlands/ZuidH

olland 72/2020

Netherlands/Z

uidH

olland 13/2020

Iceland/238/2020

USA/WI-GMF-00466/2020

Thailand/Bangkok-CONI-0020/2020

Italy/INMI10/2020

USA/IL 1375/2020

Geo

rgia

/Tb-

54/2

020

IHU

CO

VID

-0170

Belgium/AKM-030670/2020

Iceland/70/2020

Beijing/233/2020

Sen

egal

/382

/202

0

Netherlands/N

A 130/2020

Norway/1955/2020

Sco

tland

/CV

R10

1/20

20

Net

herla

nds/

Zui

dHol

land

38/

2020

England/20136096504/2020

Scot

land

/CVR

133/

2020


Iceland/177/2020

Engl

and/

2012

4034

702/

2020

USA/NY-PV08493/2020

Wales/P

HWC-244EC/2020

US

A/W

A-U

W39/2020

US

A/N

Y-P

V08

137/

2020


Sco

tland

/CV

R63

/202

0

USA/CA-CDPH-UC26/2020


Iceland/234/2020


USA/NY-PV09171/2020


US

A/W

A-U

W360/2020

US

A/O

R-U

W-1887/2020

Sco

tland

/CV

R10

0/20

20


US

A/N

Y-PV

08123/2020

Icel

and/

120/

2020

US

A/W

I-76/2020

Geo

rgia

/Tb-

1679

/202

0

Portu

gal/P

T003

6/20

20

Australia/V

IC123/2020


US

A/M

N128-M

DH

128/2020

Net

herla

nds/

Noo

rdB

raba

nt 3

9/20

20


IHUCOVID-0259


Netherlands/H

elmond 1363548/2020

Denmark/SSI-03/2020

Iceland/543/2020

Japan/SMU-0311S2/2020

USA/WA-UW-2104/2020

Icel

and/

94/2

020

Irela

nd/2

2901

/202

0

USA/WA-UW-2087/2020

Net

herla

nds/

NA

94/

2020


Belgium

/SE

M-0324120/2020

Wales/P

HWC-2

44FB/2020


Aust

ralia

/VIC

171/

2020

Eng

land

/201

3210

6804

/202

0

IHUCOVID

-005

3


Por

tuga

l/PT

0093

/202

0

Vietnam/19-01S/2020

Eng

land

/201

3006

4104

/202

0

US

A/W

A-U

W-2250/2020

Australia/V

IC231

/2020


Scotland/CVR56/2020

Australia/V

IC128/2020

Singapore/24/2020

USA/NY-N

YUMC45/2020

USA/UT 8906/2020

USA/WA-UW-1446/2020

Australia/V

IC222/2020

Wales/PHWC-254AF/2020

US

A/W

A-U

W-1705/2020

USA/NY-NYUMC72/2020

Eng

land

/201

3005

1004

/202

0

Iceland/57/2020

Georgia/Tb/2020

US

A/N

Y-NY

UM

C75/2020


USA/WA-UW-1436/2020

Iceland/485/2020

USA/NY-PV09177/2020

Wales/PHW1/2020

Net

herla

nds/

NA

152

/202

0

USA/MN124-MDH124/2020


IHUCO

VID-0

099

Iceland/297/2020

US

A/M

N57

-MD

H57

/202

0


Hangzhou/ZJU-07/2020

Eng

land

/201

3005

0904

/202

0

IHUCO

VID-0

378


England/20108034006/2020

US

A/W

A-U

W343/2020

Franc

e/ARA12

249/

2020


US

A/W

A-U

W-1587/2020

Belgium/ZF-0325152/2020

USA/WA-UW-2074/2020


US

A/W

A-U

W57/2020

US

A/W

A-U

W-1338/2020

IHU

CO

VID

-035

4

USA/NY-PV09147/2020



USA/CA7/2020

Germ

any/N

RW-0

2-1/

2020


Finland/14M32/2020

USA/WI-97/2020

Netherlands/N

A 265/2020


IHUCOVID-0069

US

A/W

A-U

W-1805/2020

Net

herla

nds/

Zui

dHol

land

8/2

020

USA/WA-UW-1965/2020

Net

herla

nds/

NA

297

/202

0

Sw

itzerland/GE

8102/2020

Eng

land

/201

3210

0404

/202

0

Iceland/379/2020

CzechRepublic/2741/2020


Gre

ece/

12/2

020

Austria/C

eMM

0019/2020

IHUCO

VID-0

239


US

A/W

A-N

H10/2020

Belgium

/JRH

-030459/2020

US

A/W

A-U

W-1666/2020

DRC/248/2020

Wales/P

HW

C-24349/2020

IHUCO

VID-0

419

Belgium

/GJJ-0325145/2020


US

A/W

A-S

64/2020

England/C

AM

B-73A

28/2020

USA/NY-P

V08463/2020

France/CVL2000/2020

US

A/W

A-S

80/2020


IHUCOVID-0151

Iceland/228/2020

Wal

es/P

HWC-2

51D5/

2020

US

A/W

A-U

W-2

218/

2020

USA/NY-PV09120/2020


England/20144065604/2020

Tianmen/HBCDC-HB-07/2020

Icel

and/

421/

2020

Canada/BC 8159203/2020

USA/NY-NYUMC85/2020

Fra

nce/

IDF

2792

/202

0

Netherlands/ZuidH

olland 41/2020

England/20134023704/2020

Eng

land

/CA

MB

-745

80/2

020


USA

/MA

3878

/202

0


US

A/W

A-U

W24/2020

Franc

e/ID

F2410

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0

Net

herla

nds/

Noo

rdB

raba

nt 7

3/20

20

USA/NY-W

adsw

orth-

1129

1-01

/2020

Wales/P

HWC-25EB0/2020

USA/W

A-UW

379/2020

USA/NY-P

V09170/2020

USA/NY-W

adsw

orth-

1138

0-01

/2020

Belgium/AGR-0324174/2020

Net

herla

nds/

Noo

rdB

raba

nt 2

2/20

20

Japan/Hu D

P K

ng 19-027/2020

India/770/2020

US

A/C

T-U

W21

3/20

20

USA/WA-UW-2169/2020

USA/WA-UW181/2020


Wales/PHWC-260C8/2020


USA

/MA

8933

/202

0

Belgium

/ULG

-10043/2020

Scotland/CVR31/2020

Wales/PHW

C-24F81/2020


Iceland/129/2020

US

A/W

A-U

W277/2020


Guangzhou/GZMU0014/2020

USA/W

A-UW

363/2020

Sco

tland

/CV

R20

1/20

20

Mexico/C

DM

X-IN

ER

05/2020

England/20132082104/2020

USA/NY-PV09102/2020


Eng

land

/201

3903

3404

/202

0

Iceland/186/2020


Fran

ce/H

F206

0/20

20

Shanghai/S

H0053/2020

USA/C

A-CD

PH-U

C9/2020

Franc

e/ARA13

22/2

020


USA/M

N68-M

DH

68/2020

England/20142002904/2020


France/IDF2256/2020

US

A/W

Y-WY

PH

L011/2020

IHUCO

VID-0

257

Eng

land

/201

4800

4904

/202

0


US

A/M

N39

-MD

H39

/202

0

IHUCO

VID-0

284

USA/NY-PV09165/2020


Finland/13M

3/2020

US

A/W

A-U

W-1784/2020

Englan

d/20

1320

2110

4/20

20


USA/AK-PHL006/2020

Finland/F

IN03032020C

/2020

USA/CA-C

DPH-UC4/2020

Nor

way

/137

9/20

20

Iceland/409/2020

Belgium/KA-03042/2020

Eng

land

/201

3906

0504

/202

0

Eng

land

/SH

EF

-C02

CA

/202

0

US

A/W

A-U

W-1351/2020



IHUCO

VID-0

449

Sau

diA

rabi

a/S

CD

C-3

324/

2020

Net

herla

nds/

NA

294

/202

0

Net

herla

nds/

NA

191

/202

0

SouthKorea/KUMC04/2020

Eng

land

/201

4401

8904

/202

0

USA/WA-UW-2240/2020

Brazil/S

PB

R-08/2020

Aus

tral

ia/N

SW

52/2

020

USA/WA-UW-1567/2020

Iceland/111/2020

Por

tug

al/P

T00

08/2

020

Iceland/448/2020

Wales/P

HWC-2

6D2B/2020

USA/WA-UW283/2020

USA/M

N44-M

DH

44/2020

Iceland/180/2020


USA/WI-105/2020

Iceland/243/2020

USA/GA 1320/2020

USA/W

I-110/2020

US

A/W

I-39/2020

Fran

ce/G

E28

43/2

020

Fran

ce/H

F215

5/20

20

Aust

ralia

/VIC

88/2

020

Englan

d/20

1380

276/

2020

Belgium

/ULG

-7242/2020

IHU

CO

VID

-012

9

Australia/WA01/2020

Singa

pore

/11/2

020

Austra

lia/N

SW61

/202

0


Hon

gKon

g/H

KP

U30

290

1/20

20


Australia/V

IC42/20

20

USA/W

A-UW

129/2020

England/20108007302/2020


USA/GA 1445/2020

Hangzhou/HZ576/2020

Wal

es/P

HW

C-2

5BC

8/20

20

Aus

tral

ia/Q

LD09

/202

0

USA/FL 9655/2020

US

A/W

A-S

12/2020

Nethe

rland

s/NA 2

67/2

020

Australia/WA03/2020

Hon

gKon

g/C

UH

K1/

2020

Icel

and/

417/

2020

Fujia

n/13

/202

0

Finland/F

IN-313/2020

Bel

gium

/DV

G-0

3251

46/2

020

USA/WI-67/2020

Aus

tria

/CeM

M00

05/2

020

Icel

and/

38/2

020

Eng

land

/SH

EF

-C05

2B/2

020

Guangdong/2020XN4448-P0002/2020

USA/NY-PV09176/2020

Iceland/314/2020


IHUCOVID

-041

6


Iceland/237/2020

USA/WA-UW-1728/2020

Australia/V

IC226/2020


Norway/1989/2020

Iceland/163/2020

Por

tuga

l/PT

0077

/202

0

US

A/W

A-S

111/2020

USA/WA-UW-1874/2020


Scotland/E

DB

098/2020

Germany/NRW-29/2020

France/BFC2147/2020

US

A/W

A-U

W281/2020

Belgium

/SB

-030990/2020

Iceland/281/2020


England/20128035202/2020

Fra

nce/

AR

A12

217/

2020

USA/LA 0842/2020

US

A/W

A-S

117/2020

Fra

nce/

IDF

0373

/202

0


Engl

and/

2014

4009

104/

2020


Iceland/139/2020

US

A/W

A-S

101/2020

Belgium

/ULG

-668

8/20

20

Netherlands/N

A 119/2020

Australia/V

IC205/2020


USA/W

A-UW

-2141/2020

Italy

/TE

4880

/202

0


Belgium/MJP-030684/2020

Icel

and/

422/

2020

Australia/V

IC111/2020

DRC/521/2020

IHU

CO

VID

-012

1

Hangzhou/HZ638/2020

Iceland/335/2020

IHUCO

VID-0

319

US

A/W

A-S

61/2020

USA/NY-NYUMC28/2020

Canada/O

N-P

HL2445/2020

US

A/U

T-00055/2020

Netherlands/Z

uidHolland 10/2020

Italy/INMI7/2020

Wales/PHWC-253DF/2020

IHUCO

VID-0

326

Germany/NRW-20/2020

Finland/13M27/2020

USA/W

A-UW

101/2020

Aus

tral

ia/N

SW

03/2

020

Iceland/52/2020

Can

ada/

BC

530

6970

/202

0

Wales/PHWC-26EBF/2020

Net

herla

nds/

NA

21/2

020

Netherlands/N

oordHolland 2/2020

Italy/INMI5/2020

USA/WA-UW-4406/2020

USA/TX1/2020

US

A/N

Y-W

adsw

orth

-112

02-0

1/20

20

USA/UT-00035/2020

Belgium

/BS

-030598/2020

Eng

land

/201

3605

08/2

020

USA/ID-UW-2198/2020

Wales/PH

WC

-26A15/2020

England/C

AM

B-77031/2020

Fra

nce/

AR

A13

07/2

020

Aus

tralia

/VIC

203/

2020

Eng

land

/201

4007

0804

/202

0

US

A/P

A 44

05/2

020

England/NOTT-10E8FA/2020


Jian

/JX

169/

2020

France/IDF3212/2020

US

A/W

A-U

W-1702/2020

Italy/CDG1/2020

Net

herla

nds/

NA

312

/202

0


Scotla

nd/E

DB386/

2020


Engl

and/

SHEF

-C02

60/2

020


Hon

gKon

g/V

M20

0025

82/2

020

Netherlands/G

elderland 11

/2020

Iceland/66/2020

USA/NY-PV08412/2020

Taiw

an/N

TU

14/2

020

USA/NY-PV09123/2020

Ger

man

y/B

avP

at2/

2020


Iceland/504/2020

IHUCO

VID-0

299

Por

tuga

l/PT

0076

/202

0

Sco

tland

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R20

3/20

20

Englan

d/20

1240

0390

2/20

20

USA/NY-NYUMC55/2020

France/B2343/2020

US

A/W

A-S

29/2020

Portu

gal/I

GC00

009/

2020

IHUCOVID

-022

4

Denmark/SSI-102/2020

Netherlands/N

A 111/2020


Engl

and/

2013

9060

004/

2020

Netherlands/Z

uidH

olland 24/2020

Wales/PHW

C-2673C/2020

IHUCO

VID-0

243

France/H

F3141/2020

Fran

ce/H

F223

9/20

20

Iceland/585/2020


Chile/Santiago op4d1/2020

France/IDF2420/2020

Taiw

an/C

GM

H-C

GU

-19/

2020

Wales/PHW

C-26D85/2020

US

A/W

A-U

W-1

593/

2020

Australia/V

IC20/2020

USA/NY-PV09181/2020


Scotland/CVR79/2020

Englan

d/SHEF-B

FD45/2

020

US

A/M

N63-M

DH

63/2020

England/20129033904/2020

England/20146060504/2020

IHU

CO

VID-0087

US

A/W

A-U

W124/2020

Iceland/25/2020

Senegal/136/2020

England/SHEF-BFCC0/2020


USA/NY-W

adsworth

-10695-01/2020

Wal

es/P

HW

C-2

6E19

/202

0

Iceland/378/2020

Wales/P

HWC-2

42C4/2020

Eng

land

/201

2003

3002

/202

0

Portugal/P

T0051/2020

Sco

tland

/ED

B30

0/20

20

IHUCOVID-0132

Fran

ce/H

F294

8/20

20

Scotland/C

VR

69/2020

USA/WI-106/2020

Sin

gapo

re/1

7/20

20

IHUCO

VID-0

272


Iceland/172/2020

Italy/FVG-ICGEB S1/2020

USA/WA-UW-1576/2020


USA/W

A-UW

-1785/2020



Aus

tral

ia/N

SW

54/2

020

USA/WA-UW-1646/2020

Hangzhou/ZJU

-04/2020


US

A/W

A-U

W92/2020

US

A/C

T-U

W27

1/20

20


Iceland/199/2020

IHUCO

VID-0

309

USA/WA-UW-1302/2020

Wal

es/P

HW

C-2

7003

/202

0

England/20134021804/2020

USA/WA-UW-2214/2020



IHUCOVID

-022

9

USA/WA-UW123/2020




USA/NY-PV09200/2020

USA/ID-UW-1957/2020

USA/W

A-UW

-1821/2020



Canada/O

N P

HL6884/2020

Engl

and/

2012

4035

802/

2020

England/20109050506/2020

England/20132108104/2020


England/NOTT-10E2B0/2020

Australia/V

IC223/2020

USA/W

A-UW

36/2020

US

A/W

A-U

W235/2020

USA/WI-117/2020

Wuhan/WIV05/2019

Belgium/BJ-030767/2020

USA/W

A-UW

-1441/2020

USA/WA-UW-1739/2020

Iceland/539/2020



USA/NY-NYUMC49/2020

Senegal/087/2020

Can

ada/

ON

PH

L374

1/20

20

Scotland/CVR55/2020



Iceland/375/2020

US

A/A

K-P

HL0

37/2

020

USA/NY-P

V08428/2020

IHUCOVID-0261

DRC/431/2020

USA/WI-64/2020

Japa

n/P

4-7

/202

0

IHU

CO

VID

-047

9

Icel

and/

538/

2020

USA/MN67-MDH67/2020


IHUCOVID

-0061

Chongqing/ZX01/2020

Por

tuga

l/PT

0071

/202

0

Engla

nd/2

0136

0229

/202

0

US

A/N

Y-N

YU

MC

3/20

20

Canada/O

N P

HL6883/2020


Spain/LaRioja201575/2020

USA/W

A-UW

-1850/2020

Iceland/169/2020

Netherlands/N

A 302/2020

Aus

tral

ia/N

SW

24/2

020

Iceland/415/2020

USA/WA-UW-2234/2020

Shanghai/S

H0107/2020


Germany/NRW-21/2020

IHUCOVID-0133


USA/C

A-CD

PH-U

C11/2020


US

A/W

I-99/

2020

Iceland/73/2020

US

A/N

Y-P

V08

136/

2020

IHUCOVID-0213

Japa

n/P

5-1/

2020

Wales/P

HWC-2

4D87/2020

US

A/N

C 6999/2020

USA/NY-NYUMC61/2020

US

A/W

A-S

3/2020

Eng

land

/201

2900

0404

/202

0

IHUCOVID-0250

US

A/W

A-U

W-1321/2020

Iceland/584/2020

USA/NY-PV08462/2020

US

A/IL

008

7/20

20

Taiwan/144/2020

USA/NY-NYUMC70/2020

England/CAM

B-738C4/2020

France/HF2150/2020

US

A/W

A-U

W-1599/2020

Iceland/215/2020

Belgium

/VM

C-030695/2020

Netherlands/N

A 237/2020

USA/WA-UW60/2020

Aus

tralia

/VIC

82/2

020

USA/NY-PV08455/2020

France/B2346/2020

USA/NY-NYUMC53/2020

US

A/W

A-U

W274/2020

Beijing/105/2020

IHUCOVID-0107

USA/W

I-18/2020

Hangz

hou/

HZ-1/2

020

Eng

land

/201

3210

5704

/202

0

US

A/W

A-U

W-2164/2020

Scotland/CVR99/2020

Wales/PHW

C-23FFB/2020

Spain/CastillayLeon201372/2020

Eng

land

/201

4007

0604

/202

0

USA/WA-UW-1651/2020

Por

tuga

l/PT

0066

/202

0P

ortu

gal/P

T00

59/2

020


Engla

nd/2

0134

0155

04/2

020

Engl

and/

2013

9058

304/

2020

Singapore/36/2020

Por

tuga

l/PT

0068

/202

0


Aus

tral

ia/V

IC15

1/20

20

USA/WA-UW153/2020

USA/WI-38/2020


Net

herla

nds/

NA

129

/202

0

Kuwait/KU005/2020

Eng

land

/201

3903

1804

/202

0


Taiw

an/N

TU

11/2

020

Taiwan/NTU01/2020

US

A/W

A-S

49/2020

Hon

gKon

g/V

B20

0249

50/2

020

USA/WA-UW-1733/2020

USA/UT-00290/2020

Belgium

/MTR

-03026/2020

Italy/INMI6/2020

IHUCOVID-0149

Canada/BC 4122951/2020

US

A/W

A-S

68/2020

Australia/V

IC68/2020

Canada/NS 13/2020

England/20104007503/2020


Net

herla

nds/

Ove

rijss

el 2

/202

0

Englan

d/20

1300

3220

4/20

20

Norway/2088/2020

Jiang

su/J

S02/2

020

Australia/QLD04/2020

US

A/N

Y-PV

08146/2020

England/20130061304/2020

Belgium/GE-0317105/2020

Wales/PHWC-266A8/2020

Iceland/39/2020


Iceland/195/2020

DRC/191/2020

USA/NY-PV09045/2020

Ireland/Dublin-22361/2020

USA/NY-PV08474/2020

USA/NY-NYUMC64/2020

Hangzhou/HZ79/2020

Shang

hai/S

H0110

/202

0

Netherlands/N

A 10/2020

Bel

gium

/ULG

-861

7/20

20

USA/W

A-UW

341/2020

Australia/V

IC159/2020

France/N1620/2020

Germany/BavPat1/2020

Iceland/185/2020Iceland/245/2020

Canad

a/ON P

HL369

2/2020

Eng

land

/201

3210

4404

/202

0

USA

/WI-1

13/2

020

USA/WA-U

W-1644/2020

Iceland/3/2020

Bel

gium

/ULG

-100

85/2

020

Eng

land

/201

3906

0804

/202

0

Aus

tralia

/VIC

47/2

020

US

A/W

A-U

W-1

574/

2020

US

A/IL-U

W-1265/2020

IHU

CO

VID

-017

7

Spain/Madrid LP12 21/2020


Engla

nd/2

0138

0869

04/2

020

USA/NE 6

605/

2020



Australia/V

IC153/2020

Netherlands/Z

uidHolland 11/2020

Aus

tral

ia/N

SW

53/2

020

Aus

tral

ia/N

SW

11/2

020


USA/WA-UW170/2020

Portugal/P

T0031/2020

USA/W

A-UW

127/2020

USA/W

A-UW

290/2020


Portug

al/PT

0081/2020

Belgium

/VP

MV

-0325153/2020



Franc

e/ID

F2532

/202

0


Engl

and/

2013

4069

004/

2020

USA/NH 0008/2020

Taiw

an/C

GM

H-C

GU

-18/

2020

USA/NY-PV08150/2020


Australia/Q

LDID

919/2020

US

A/N

Y-P

V09

303/

2020

France/HF2797/2020

IHUCO

VID-0

235

US

A/W

A-N

H13/2020

USA/NY-PV09193/2020

Wales/PHW

C-26E82/2020


India/777/2020

IHU

CO

VID

-011

8

USA/NY-PV08469/2020

Iceland/551/2020


IHU

CO

VID

-0070

USA/WA-UW-1753/2020

Scotland/E

DB

275/2020

USA/IA 6394/2020

Belgium/U

LG-10018/2020

Belgium/VD-0318102/2020

Canada/BC 8894200/2020


SouthA

frica/KR

ISP

0006/2020

US

A/W

A-S

82/2020

Net

herla

nds/

NA

144

/202

0

Belgium

/NL-030447/2020


Indi

a/11

00/2

020

Wales/PHWC-2613E/2020

Denm

ark/SSI-01/2020

USA/WA-UW-1712/2020



Net

herla

nds/

NA

199

/202

0

England/S

HE

F-C

00DF

/2020

US

A/N

Y-P

V08

498/

2020

US

A/N

Y-P

V09

154/

2020

US

A/N

Y-P

V08

135/

2020

US

A/W

A-U

W244/2020

USA/NY-PV08416/2020

Belgium/SC-0319113/2020

Engla

nd/20139046504

/2020

Shangrao/JX

1177/2020

US

A/W

A-S

2/2020

Belgium/RT-030460/2020

IHUCOVID-0344

USA/NY-PV09032/2020

Icelan

d/46

8/20

20

Scotland/E

DB

327/2020

Bel

gium

/VR

JM-0

3231

14/2

020



Wal

es/P

HW

C-2

5B5F

/202

0

Scotland/C

VR

64/2020

USA/WA-UW-2112/2020

IHU

CO

VID

-039

8

IHUCOVID

-028

3


US

A/W

A-U

W23/2020

Shangrao/JX1178/2020

USA/MN55-MDH55/2020

Scotland/CVR70/2020

IHU

CO

VID

-0276

Net

herla

nds/

NA

274/

2020

USA

/PA

1881

/202

0

Canada/B

C 0443574/2020

US

A/W

Y-WY

PH

L001/2020

Belgium/RJL-030588/2020

USA/IL-UW352/2020

USA/NY-PV08485/2020

USA/M

N58-M

DH

58/2020

Belgium/DRV-0325157/2020

Iceland/442/2020

Nethe

rland

s/NA 6

/202

0

Shanghai/S

H0007/2020

Fran

ce/H

F329

3/20

20

USA/NY-PV08127/2020


England/20132091704/2020

IHUCOVID-0145

Sw

itzerland/1000477377/2020


Malaysia/M

KAK-CL-2020-6430/2020

Eng

land

/CA

MB

-737

7C/2

020

Denmark/SSI-1

04/2020

USA/WA-UW-2244/2020

IHU

CO

VID-0421

US

A/VA

-DC

LS-0024/2020

Iceland/260/2020


Aus

tral

ia/V

IC32

2/20

20

Belgium/BS-0325183/2020

USA/WI-0

3/2020

IHU

CO

VID

-024

1

USA/WI-11/2020

Scotland/E

DB

019/2020

France/B

2351/2020

Australia/V

IC29/2020

USA/NY-NYUMC50/2020

Belgium

/FR-03029/2020

USA/WA-UW-1689/2020

Canada/B

C 0554880/2020

England/20139069404/2020

Por

tuga

l/PT

0095

/202

0


USA/WA-UW-4407/2020

Scot

land

/CVR

132/

2020

Fran

ce/H

F199

5/20

20


Canada/O

N P

HL8580/2020

USA/W

A-S119/2020

USA/NY-PV08466/2020

IHUCO

VID-0

494

US

A/C

ZB

-RR

057-

005/

2020

USA/WI-69/2020

Eng

land

/201

3202

0404

/202

0

Iceland/47/2020


Shenzhen/HKU-SZ-005/2020

USA/W

A-UW

-1718/2020

Germany/NRW-15/2020

US

A/W

A-S

20/2020

IHUCO

VID-0

308


Wuhan/WH01/2019

US

A/O

SU

P01

09/2

020


Australia/V

IC314

/2020

Engla

nd/20099038206

/2020

Aus

tral

ia/V

IC17

8/2

020

USA/WA-UW-1329/2020

Eng

land

/201

0909

4506

/202

0



Japa

n/P

4-3/

2020

DR

C/1

58/2

020

Iceland/450/2020

IHU

CO

VID

-0360

Chongqing/YC01/2020

Icel

and/

501/

2020

Australia/V

IC297/2020

Australia/N

SW

45/2020


Icel

and/

330/

2020

US

A/T

X 2

967/

2020

Australia

/VIC64/2020

Por

tuga

l/PT

0020

/202

0

Belgium

/BC

-03016/2020

Iceland/291/2020

Belgium

/VB

K-03

061/2020

Eng

land

/201

0909

4106

/202

0


Iceland/487/2020

Belgium

/UG

ent-12/2020

Scotland/CVR78/2020

USA/G

A 2742/2020

Hangzhou/H

Z477/2020

Wales/PHWC-2681B/2020



US

A/W

A-U

W-1

569/

2020

USA/CT-U

W-4346/2020

US

A/W

A-U

W81/2020

US

A/W

A-N

H14/2020


Eng

land

/201

3901

3406

/202

0


US

A/W

A-S

7/2020

Bel

gium

/NM

A-0

3241

69/2

020


Bel

gium

/ULG

-366

5/20

20


9/2020

England/20139065604/2020

IHUCOVID

-049

8

Iceland/14/2020

Wales/PHW

C-26839/2020

Netherlands/N

A 310/2020

USA/NY-PV08449/2020

Belgium/RHA-0324177/2020

Yunnan/IVDC-YN-003/2020


Icel

and/

502/

2020


US

A/N

Y-P

V09

168/

2020

USA/M

N59-M

DH

59/2020

Germany/NRW-37/2020

Spain/VH000001133/2020

Icel

and/

540/

2020

France/GE1583/2020

IHUCOVID

-020

4

Eng

land

/201

3900

9704

/202

0

Iceland/489/2020

Can

ada/

ON

PH

L097

6/20

20

Sco

tland

/CV

R20

7/20

20

Jiangsu

/JS03/2020

New

Zea

land

/01/

2020

USA/WI-34/2020


USA/WA-UW117/2020

US

A/W

A-U

W-1683/2020

Belgium

/UG

ent-8/2020

Belgium/STM-0324170/2020

Icel

and/

377/

2020

IHUCOVID-0251

Belgium/BM-03012/2020

IHUCOVID

-022

1

US

A/W

A-U

W71/2020

USA/NY-P

V08138/2020

US

A/IL-U

W-1311/2020

England/20128036004/2020

Netherlands/N

A 159/2020


Belgium

/GS

A-030454/2020

Canada/O

N P

HL0178/2020

IHU

CO

VID

-032

4

Singapore/33/2020

England/SHEF-C034F/2020


Wales/P

HW

C-25B

7D/2020

Scot

land

/CVR

42/2

020

Iceland/263/2020




England/SHEF-C02BB/2020


USA/WA-UW109/2020

Australia/V

IC05/2020

Fra

nce/

IDF

2561

/202

0

Australia/V

IC165/2020

Por

tuga

l/IG

C00

010/

2020

USA/W

I-108/2020

USA/WA-UW-1433/2020

US

A/F

L 71

69/2

020


Senegal/016/2020


USA/UT-00032/2020


Belgium/VM

-0325171/2020

Wal

es/P

HW

C-2

4534

/202

0

US

A/W

Y-WY

PH

L018/2020

Luxe

mbo

urg/

LNS

3694

003/

2020


USA/NY-PV09064/2020

Fra

nce/

AR

A10

876/

2020

England/201040081/2020

Wales/PHW

C-26A9D/2020

US

A/N

J 32

01/2

020

Engl

and/

2013

9063

704/

2020

Netherlands/N

A 87/2020


US

A/W

A-U

W-1822/2020

US

A/W

A-S

91/2020

US

A/W

I-111

/202

0

USA/NY-NYUMC76/2020

Shenzhen/HKU-SZ-002/2020

USA/CT-UW259/2020

USA/MN46-M

DH46/2020

Sichuan/IVDC-SC-001/2020

Wal

es/P

HW

C-2

6E37

/202

0

Engl

and/

2013

6018

7/20

20

Hangzhou/H

ZC

DC

0013/2020


USA/NY-NYUMC2/2020

IHUCOVID-0082

Hangzhou/HZ481/2020

Belgium

/JK-0305110/2020

IHU

CO

VID

-033

8

Taiwan/NTU15/2020

Por

tuga

l/PT

0052

/202

0

US

A/C

A-C

DP

H-U

C2/2020

US

A/W

I-59/2020

Australia/N

T01/2020

IHU

CO

VID

-022

2

Indi

a/16

17/2

020

USA/W

A-UW

-1670/2020

IHU

CO

VID

-007

1

England/20140003404/2020

US

A/IL 0089/2020

Aust

ralia

/NSW

36/2

020

USA/WA-UW-1880/2020

Fran

ce/H

F259

7/20

20

US

A/W

A-S

45/2020

Sco

tland

/ED

B01

8/20

20

Australia/V

IC-C

BA

3/2020

Icel

and/

484/

2020

IHUCOVID-0263

USA/NY-PV09127/2020



USA/CA-PC101P/2020

USA/WA-UW241/2020

Belgium/NKR-030645/2020

Iceland/100/2020

US

A/N

Y-N

YU

MC

91/2

020

Eng

land

/201

3800

37/2

020

IHUCOVID-0131

England/NOTT-10E61E/2020

Icel

and/

446/

2020

Wales/PHWC-269AF/2020


Bel

gium

/DM

M-0

3251

80/2

020

Iceland/261/2020

Aus

tral

ia/V

IC14

4/2

020

Spa

in/M

adrid

2017

38/2

020

France/Clermont-Ferrand 650/2020


Belgium

/DW

AE

E-030954/2020

USA/WA-UW-2192/2020


Wales/PHW

C-24507/2020


Finl

and/

13M

60/2

020

Iceland/542/2020



US

A/U

PH

L-03/2020

Bel

gium

/ULG

-969

4/20

20

England/20139058104/2020

Irel

and/

2405

2/20

20

England/SHEF-C03E5/2020

IHUCOVID-0030

Engla

nd/20138014402

/2020


Sco

tland

/ED

B08

6/20

20

USA/N

Y-PV09

142/

2020

England/CAMB-76E92/2020

Icel

and/

341/

2020

Netherlands/N

oordBrabant 58/202

0

Germ

any/N

RW-0

1/20

20

Eng

land

/201

3206

7304

/202

0

Georgia/Tb-712/2020


Austria/C

eMM

0018/2020

Canada/BC 8896915/2020

Can

ada/

ON

PH

L331

8/20

20

US

A/W

I-114/2020

Australia/V

IC241/2020

Belgium/BG-030551/2020

Icel

and/

459/

2020

USA/NY-PV09126/2020

Iceland/555/2020

USA/NY-N

YUMC60/2020

England/20144012404/2020

US

A/W

A-S

33/2020

USA/WA-U

W-1661/2020

USA/WI-02/2020

Iceland/308/2020



Thaila

nd/Bang

kok-CO

NI-0

022/2020

Iceland/269/2020

USA

/MA

9889

/202

0

Iceland/83/2020

Fra

nce/

AR

A11

036/

2020

Netherlands/N

A 89/2020


SouthA

frica/KR

ISP

0012/2020

USA/WI-29/2020

Slovakia/SK-BMC1/2020



Japa

n/P

4-8

/202

0

Iceland/106/2020

US

A/O

R 5

430/

2020

Canada/O

NP

HL8539/2020

Iceland/118/2020


IHUCOVID

-022

8

Sco

tland

/ED

B03

0/20

20

USA/WA-UW388/2020

US

A/W

A-U

W118/2020


Belgium/VNP-0320107/2020

USA/MN35-M

DH35/2020

Shang

hai/SH

0070/2020


US

A/W

A-U

W171/2020

Net

herla

nds/

NA

298

/202

0Han

gzho

u/H

ZC

DC

6789

/202

0

Netherlands/N

oordBrabant 45/202

0

USA/WI-62/2020

IHU

CO

VID

-0488

Wales/PHW

C-26FAD/2020

USA/NY-PV09186/2020

Wuh

an/W

IV06

/201

9

Belgium/VLG

-0325162/2020

Bel

gium

/JJC

-032

5165

/202

0

England/20132096004/2020

USA/NY-P

V08143/2020

Sw

itzerland/SZ

1417/2020


Netherlands/N

A 15/2020

Malaysia/189332

/2020

IHU

CO

VID

-016

0

USA/NY-PV09156/2020

Wales/PHWC-25FAE/2020

USA/O

R-U

W-1849/2020


Wales/P

HWC-2511

1/2020

Engla

nd/S

HEF-

BFFD

6/20

20

US

A/W

A-U

W184/2020

Wales/P

HW

C-25551/2020

Belgium

/ULG

-10027/2020

USA/NY-P

V09132/2020

Iceland/157/2020

England/20122032802/2020


Eng

land

/201

3601

37/2

020

Belgium/ECC-030686/2020

Mexico/C

DM

X-InD

RE

06/2020

Guangdong/G

D2020086-P0021/2020

Indi

a/10

93/2

020

USA/WI-58/2020

USA/W

A-UW

225/2020

Scotla

nd/CVR75/2020

Wales/PHWC-270A9/2020

IHU

CO

VID

-017

6

Wales/PHW

C-2469B/2020


Netherlands/N

oordBrabant 36/2020

Hon

gKon

g/H

KP

U40

-280

1/20

20

Netherlands/N

A 259/2020

France/BFC2709/2020

USA/NY-PV08478/2020

IHUCOVID-0026

Aus

tral

ia/V

IC31

3/20

20

US

A/N

Y-N

YU

MC

156/

2020

IHUCOVID-0062

Iceland/423/2020

Wales/PHW

C-2621D/2020

Belgium/NKMA-0324172/2020

IHUCOVID-0139

India/3118/2020

Netherlands/G

elderland 6/2020

USA/CA 2602/2020

India/3239/2020

Wales/P

HW

C-260A

A/2020

England/NOTT-10E010/2020

USA/WA-UW-1977/2020

Wuh

an/W

H03/2

020

Australia

/VIC

291/2020

Por

tuga

l/PT

0091

/202

0

USA/WA-UW-1697/2020


USA/NY-P

V09137/2020

Aus

tralia

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18/2

020

US

A/W

A-U

W149/2020

Icel

and/

547/

2020

Austri

a/CeM

M00

09/2

020

US

A/G

A 27

41/2

020


Chi

le/S

antia

go-2

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0

Israel/ISR

IT0320/2020


Australia/V

IC180/2020

Germany/NRW

-39/2020

Wales/PHWC-2458F/2020

US

A/W

A-U

W190/2020

Belgium/GHB-03021/2020

US

A/W

A-U

W204/2020

Net

herla

nds/

Noo

rdB

raba

nt 6

9/20

20

Eng

land

/CA

MB

-74B

42/2

020


IHUCOVID-0227

IHUCO

VID-0

209


USA/WI-33/2020

Icelan

d/142

/2020

Sco

tland

/CV

R15

3/20

20

Eng

land

/201

2600

9804

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0

Aus

tral

ia/V

IC34

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20

Icel

and/

492/

2020

Taiwan/3/2020


Iceland/145/2020

Canada/O

N P

HL0052/2020

US

A/VA

-DC

LS-0003/2020

Canada/B

C 1318414/2020

US

A/U

T-00014/2020

Netherlands/Z

uidHolland 103/2020

Engla

nd/2

0130

0534

04/2

020

USA/CT-UW272/2020

Eng

land

/201

3209

2004

/202

0

Taiwan/C

GM

H-C

GU

-13/2020

Wales/P

HWC-2

5867/2020

Eng

land

/201

3905

9904

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0

IHUCOVID

-029

8

Singa

pore/40/2020

Wales/P

HWC-2

422E/2020

England/20100022706/2020


Gua

ngdo

ng/2

0SF17

4/20

20

HongK

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HKPU63-1

402/

2020

US

A/W

A-U

W70/2020

Spa

in/V

alen

cia9

0/20

20

Wales/PHW21/2020

Belgium/UG

ent-25/2020

Belgium/VGA-030672/2020Iceland/187/2020

Eng

land

/201

3800

54/2

020

Net

herla

nds/

NA

295

/202

0

DRC/KN-0058/2020

Eng

land

/201

3402

5904

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0


France/H

F3295/2020

Bel

gium

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L-03

2413

0/20

20

Wal

es/P

HW

C-2

4DD

2/20

20

Icel

and/

483/

2020

Iceland/454/2020

Iceland/594/2020

Can

ada/

ON

PH

L391

9/20

20

Iceland/95/2020


Iceland/452/2020


Indi

a/10

63/2

020

Iceland/405/2020

Iceland/148/2020

Canada/O

N P

HL5705/2020

France/B2335/2020

Icel

and/

283/

2020


IHUCOVID-0493

USA/CZB-RR057-006/2020

Wal

es/P

HWC-2

516C

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0

Bel

gium

/ULG

-100

24/2

020

Nepal/61/2020

Germany/NRW-22/2020

Wales/P

HW

C-26C

4C/2020

Taiwan/NTU13/2020

USA/WA-UW69/2020


Wales/PHWC-246E6/2020

Netherlands/N

A 102/2020

Sco

tland

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R13

9/20

20

US

A/W

A-U

W143/2020

England/20109099206/2020

England/NOTT-10E3AE/2020

USA/G

A 1299/2020

USA/MN48-M

DH48/2020


USA/W

A-UW

49/2020

IHUCOVID-0076

USA/WA-UW-1319/2020



Iceland/317/2020

France/HF2496/2020


Wales/P

HW

C-25F

F9/2020


Canada

/ON

PH

L3459/2020


USA/NY-NYUMC39/2020

Iceland/23/2020

Taiwan/4/2020

IHUCOVID-0349

Gua

ngdo

ng/2

0SF02

8/20

20

Bel

gium

/ULG

-973

2/20

20

US

A/W

A-U

W177/2020

US

A/W

A-U

W-1562/2020

USA/WA-UW78/2020


France/IDF2849/2020

Eng

land

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TT-1

0DFE

7/20

20

USA/NY-NYUMC66/2020

Wuhan/WIV02/2019



Eng

land

/200

9900

06/2

020

Austra

lia/N

SW58

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0

IHU

CO

VID

-050

0

England/20130065604/2020

Iceland/159/2020

Can

ada/

ON

PH

L369

5/2

020

US

A/W

A-U

W93/2020


USA/W

A-UW

74/2020

USA/NY-NYUMC34/2020

Taiwan

/NT

U04/202

0


US

A/W

A-U

W-1473/2020

Engl

and/

2013

9057

204/

2020

USA/WA-UW-1417/2020

US

A/V

A-D

CLS

-0011/2020

US

A/W

I-52/

2020

IHUCO

VID-0

072

Luxembourg/Lux1/2020


Netherlands/N

A 11/2020

Wuhan/IVDC-HB-05/2019

USA/CA-MG0987/2020

Iceland/571/2020

Net

herla

nds/

NA

55/

2020


Icel

and/

339/

2020

US

A/N

Y-N

YU

MC

81/2

020

Brazil/AMBR-02/2020

Wales/PHW

C-240E8/2020

Finl

and/

13M

58/2

020

Hangzhou/HZ49/2020

Wales/PHWC-26F9E/2020



Guangdong/2020X

N4373-P

0039/2020

USA/WA-UW-1796/2020

Belgium/QL-0324173/2020


Sco

tland

/CV

R44

/202

0

Iceland/564/2020

Aus

tral

ia/N

SW

28/2

020

USA/W

A-UW

255/2020

Taiwan/NTU19/2020

IHU

CO

VID

-021

7

Netherlands/N

A 160/2020

USA/NY-PV09309/2020

Australia/V

IC213/2020

IHUCO

VID-0

424

USA/NY-PV08448/2020

US

A/W

A-U

W95/2020

NewZealand/20VR0275/2020

US

A/W

A-S

22/2020

England/201360052/2020

US

A/N

J-NY

UM

C111/2020

Austra

lia/V

IC26

0/20

20

US

A/W

A-U

W194/2020

Iceland/435/2020

USA/WA-UW-1955/2020

England/20132024504/2020

Iceland/2/2020

US

A/W

A-N

H9/2020

IHUCO

VID-0

318


USA/WA-UW-2145/2020


Taiwan/225/2020

US

A/W

I-16

/202

0


USA/WA-UW-1342/2020

IHUCOVID

-029

5

US

A/W

A-S

102/2020

Netherlands/Z

uidHolland 2/2020

US

A/U

T-00012/2020

Bel

gium

/NIM

-032

416

1/20

20

USA/SC 3572/2020

Englan

d/SHEF-C

051C

/202

0

US

A/W

A-U

W-1308/2020

Taiw

an/2

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0


USA/NY-PV09178/2020

US

A/W

I-23/2020

Hon

gKon

g/H

KP

U6

2101

/202

0

India/763/2020

Shang

hai/S

H0008

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0


Engla

nd/2

0140

0028

04/2

020

US

A/W

A-U

W89/2020

USA/C

A-CD

PH-U

C6/2020


USA/CT-UW-4366/2020

USA/NY-PV09192/2020

USA/NY-PV09187/2020

Eng

land

/201

3400

5104

/202

0

Finland/13M69/2020

USA/WA-UW25/2020

Eng

land

/CA

MB

-746

AB

/202

0

Engla

nd/C

AMB-7

46D8/

2020


Scotland/C

VR

190/2020

Netherlands/Z

uidHolland 43/2020

Wales/P

HW

C-262A

4/2020

IHU

CO

VID

-045

0

England/C

AM

B-76F

9F/2020

USA/W

A-UW

-1731/2020


USA/NY-PV08120/2020

England/20142018004/2020

Belgium/DPG-0325123/2020

Eng

land

/201

3400

6504

/202

0

France

/HF2234/2020

Singap

ore/

29/2

020

Iceland/323/2020


Belgium/GLA-0324179/2020

Wales/PHW

C-2575B/2020


Wale

s/PHW

C-268

84/2

020

Aus

tralia

/VIC

81/2

020

Can

ada/

BC

527

5718

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0



Net

herla

nds/

NA

291/

2020

Iceland/372/2020

Eng

land

/CA

MB

-747

5D/2

020


Aust

ralia

/VIC

87/2

020

USA/WA-UW-1608/2020

USA/NY-P

V09164/2020

Engla

nd/2

0132

0177

04/2

020

US

A/W

A-U

W131/2020

Iceland/380/2020

Iceland/125/2020

Senegal/003/2020

Wales/PHW

C-26DA3/2020

Aus

tralia

/VIC

166/

2020

Portugal/P

T0001b/2020

Can

ada/

SK

4/2

020


Wal

es/P

HW

C-2

57B

5/20

20


Iceland/306/2020

Franc

e/HF14

65/2

020

IHUCOVID-0495


USA/MN132-MDH132/2020

USA/WA-UW96/2020

Wal

es/P

HW

C-2

705E

/202

0

Australia/V

IC189/2020

Belgium

/RR

-030699/2020

Canada/BC 9345042/2020


Scotland/CVR83/2020

Engl

and/

2013

4068

604/

2020



Can

ada/

ON

PH

L751

3/20

20

Eng

land

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EF

-C01

EB

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0

Eng

land

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3009

2304

/202

0


US

A/W

A-U

W-1695/2020

Sco

tland

/ED

B15

7/20

20

Iceland/445/2020

Netherlands/N

A 109/2020

Iceland/214/2020

Belgium/DB-03023/2020

Bel

gium

/ULG

-971

4/20

20

Foshan/20S

F207/2020

USA/UT-00028/2020

USA/MN133-MDH133/2020

USA/WA-UW-1306/2020

Por

tuga

l/PT

0072

/202

0

USA/W

A-UW

59/2020

USA/UT-00025/2020

Belgium/FAE-030948/2020

USA/W

A-UW

30/2020

Eng

land

/201

3200

5204

/202

0

Iceland/443/2020

Engla

nd/2

0132

0054

04/2

020

Eng

land

/SH

EF-

C06

0A/2

020


USA/WI-96/2020

Eng

land

/201

3110

4804

/202

0

US

A/C

T-U

W32

8/20

20

Netherlands/U

trecht 10/2020


USA/WI-116/2020

Canada/BC 7277381/2020

USA/NY-PV09057/2020

US

A/A

K-P

HL0

15/2

020



USA/NY-PV09023/2020

England/20148006704/2020


USA/NY-PV08413/2020

US

A/N

Y-P

V09

090/

2020

Iceland/401/2020

Finland/F

IN-508/2020

Wal

es/P

HW

C-2

5F62

/202

0


Iceland/98/2020

IHU

CO

VID

-017

8

US

A/W

A-N

H12/2020



England/C

AM

B-737F

4/2020

Algeria/G

0640 2265/2020

USA/WA-UW362/2020

USA/WI-5

5/2020

US

A/W

A-S

32/2020

IHU

CO

VID

-0155

Can

ada/

ON

PH

L074

3/20

20


Guangzhou/20S

F206/2020

IHU

CO

VID

-023

1


Netherlands/N

A 219/2020


US

A/W

A-U

W179/2020

USA/WA-UW104/2020

Sco

tland

/CV

R20

0/20

20

Sco

tland

/CV

R40

/202

0


Belgium/VRAR-030643/2020


Australia/V

IC173/2020


US

A/W

A-S

77/2020

France

/B2330/2020

Denmark/SSI-04/2020

Shand

ong/IVD

C-S

D-001/2020

US

A/V

A 63

77/2

020

USA/WA-UW247/2020


Netherlands/N

A 13/2020

Eng

land

/NO

TT-

10E

39F

/202

0


Belgium/ULG-1

0088/2020


Icel

and/

223/

2020

USA/W

A-S110/2020

IHUCOVID-0020

NewZealand/20VR0276/2020


USA/W

A-UW

113/2020

DRC/82/2020

Fra

nce/

Lyon

069

3/20

20


Eng

land

/CA

MB

-770

13/2

020

US

A/W

A-U

W164/2020

Icel

and/

498/

2020

US

A/W

A-U

W300/2020

Scotland/C

VR

41/2020

Germany/NRW-42.2/2020

Eng

land

/201

3610

41/2

020

Belgium

/HC

M-0324151/2020

Australia/V

IC229/2020

Netherlands/N

A 37/2020

Finland/FIN-266/2020

Geo

rgia

/Tb-

82/2

020

Belgium

/SQ

-03043/2020

Shanghai/S

H0045/2020

New

Zea

land

/20V

R01

89/2

020

England/20129126204/2020

Iceland/433/2020

DRC/397/2020


Guangdong/2020XN4475-P0042/2020

Aus

tral

ia/V

IC23

9/20

20

Engla

nd/S

HEF-C04

F1/2

020

Peru/010/2020

Fran

ce/A

RA

1091

0/20

20

US

A/W

A-S

116/2020

Aus

tralia

/VIC

116/

2020


Belgium/HC-030760/2020

USA/NY-PV09304/2020

Net

herla

nds/

NA

271/

2020

England/20104004902/2020

Iceland/LUH

-01/2020

Iceland/258/2020

Netherlands/N

A 95/2020

Aus

tral

ia/V

IC45

/202

0

Wales/P

HW

C-26B

4F/2020


USA/ID-UW-2134/2020

USA/W

A-UW

37/2020

Aus

tral

ia/V

IC10

/202

0


Italy/TE4925/2020


Aus

tralia

/VIC

21/2

020

USA/MN60-MDH60/2020

Jiangsu/JS01/2020


Canada/B

C 3842755/2020

Eng

land

/201

3903

2404

/202

0

Wuh

an/IP

BC

AM

S-W

H-0

2/20

19

England/NOTT-10E46F/2020

Sha

ngha

i/SH

0040

/202

0

Italy/TE5472/2020

IHUCOVID

-038

2

Iceland/208/2020

England/20134071804/2020

Iceland/285/2020

Iceland/553/2020

England/SHEF-C01FA/2020

USA/NE 7

025/

2020

Aus

tral

ia/N

SW

21/2

020


IHUCO

VID-0

249

USA/WI-102/2020

Belgium/UM-0318103/2020


Engl

and/

CAM

B-73

7B8/

2020


Jiang

xi/IV

DC-JX-0

02/2

020

England/20132082004/2020

Israel/ISR

JP0320/202

0


Denmark/SSI-09/2020

England/SHEF-BFF6D/2020

Iceland/301/2020

USA/WA-UW169/2020

Franc

e/HF24

05/2

020

US

A/W

A-S

56/2020

USA/WA-UW-2078/2020

US

A/W

A-U

W-1846/2020

USA/N

Y-PV08

435/

2020

US

A/W

A-S

72/2020

Englan

d/20

1360

1550

2/20

20


Iceland/386/2020

Singapore/2/2020

Can

ada

/ON

PH

L335

0/2

020

Aus

tral

ia/V

IC36

/202

0

Netherlands/Z

uidH

olland 29/2020

India/772/2020

Netherlands/E

indhoven 1363782/2020

Australia/N

SW

25/2020

Netherlands/U

trecht 5/2020


USA/WA-UW237/2020

Sco

tland

/ED

B29

7/20

20

Iceland/516/2020

Brazil/S

PB

R-07/2020

IHUCOVID-0049


England/20108004702/2020

Fran

ce/A

RA

1193

9/20

20

Iceland/569/2020

Engl

and/

2013

0072

204/

2020

Belgium/UGent-2/2020

Eng

land

/201

0614

5903

/202

0

Belgium/OS-0325137/2020

Sin

gapo

re/8

/202

0



Algeria/G

0860 2262/2020

Engl

and/

2013

9063

804/

2020

US

A/W

A-U

W-1988/2020

France/V

alence 425/2020

Icel

and/

256/

2020

Kuw

ait/K

U00

8/20

20

USA/MN65-MDH65/2020

IHUCOVID-0135

USA/WI-101/2020

USA

/MA

3672

/202

0

USA/WA-UW-2144/2020

Engl

and/

2014

2021

604/

2020

Singapore/37/2020


Germ

any/Baden-W

uerttemberg-1/2020

US

A/V

A-D

CLS

-0019/2020


Belgium/ULG-1

0089/2020

Chile/Talca-2/2020


IHUCOVID-0117

US

A/N

Y-N

YU

MC

31/2

020

Aus

tral

ia/N

SW

15/2

020


DRC/236/2020


USA/NY-NYUMC78/2020

Belgium/SS-031047/2020

Iceland/227/2020

IHU

CO

VID

-023

7


Taiw

an/N

TU03

/202

0


IHU

CO

VID

-0194

Belgium/SN-03031/2020

USA/WI-G

MF-00744/2020


Wales/PHWC-26B12/2020

Iceland/40/2020

Canada/NB 6/2020

USA/WI-4

2/2020

England/200981386/2020

Wuhan/WIV07/2019


Engl

and/

2010

9039

306/

2020


Engl

and/

2014

4008

004/

2020

US

A/M

D 0

026/

2020

US

A/W

A-U

W289/2020

England/20130035204/2020


USA/WI-72/2020

IHUCOVID-0040

USA/WI-83/2020

Icel

and/

262/

2020


US

A/W

A-U

W167/2020

Engl

and/

2012

4002

102/

2020


Russia/StPetersburg-RII3992/2020

Iceland/110/2020


Italy/TE

4959/2020


Hon

gKon

g/H

KP

U42

-030

2/20

20

USA/WA-UW386/2020

US

A/W

A-S

53/2020

Iceland/428/2020

IHUCOVID-0057

Austra

lia/N

SW57

/202

0


US

A/N

Y-N

YU

MC

36/2

020

USA/NY-PV08447/2020

IHUCOVID-0345

IHUCO

VID-0

258

Iceland/45/2020

US

A/M

N29

-MD

H29

/202

0

IHU

CO

VID

-0240

Nor

way

/169

4/20

20

Fran

ce/ID

F276

8/20

20

Hon

gKon

g/H

KP

U34

300

1/20

20

England/NOTT-10E64B/2020

Portu

gal/P

T003

5/20

20

US

A/W

A-U

W75/2020

USA/W

A-UW

-2180/2020

Sco

tland

/ED

B35

6/20

20

France/H

F2237/2020

Hungary/mbl1/2020

IHUCO

VID-0

211

Zhejia

ng/W

Z-01/

2020

Wales/PHWC-26FBC/2020

England/20142082704/2020

USA/NY-NYUMC69/2020

Austri

a/CeM

M00

02/2

020

Italy

/TE

5056

/202

0

USA/CA-S

CCPHD-UC16/2020

Engl

and/

2012

6012

702/

2020

US

A/C

A2/

2020


IHU

CO

VID

-0254


IHUCOVID

-035

5

Brazil/SPBR-06/2020


Iceland/494/2020


US

A/N

Y-P

V09

063/

2020

Engl

and/

2012

4020

402/

2020

US

A/W

A-U

W54/2020

Por

tuga

l/PT

0061

/202

0

USA/IA 6390/2020

Eng

land

/201

4007

2504

/202

0

Australia/Q

LDID

924/2020

England/20140000404/2020

US

A/N

Y-N

YU

MC

98/2

020

USA/WA-UW-1677/2020

IHUCOVID-0475


US

A/U

PH

L-01/2020

Scotla

nd/EDB391/2020

Japan/SMU-0311S3/2020

DRC/73/2020

France/B2337/2020

IHUCOVID-0233

Hangzhou/HZ62/2020

USA/CA9/2020

US

A/W

A-U

W-1

792/

2020

Net

herla

nds/

NA

74/

2020

Scotland/E

DB

099/2020

England/20129160704/2020

IHUCO

VID-0

375

Canada/O

N P

HL5710/2020

Fran

ce/A

RA

1195

2/20

20

Singa

pore/3Clin/2020

USA/W

A-S104/2020

IHUCOVID-0171

USA/NY-PV09028/2020

US

A/W

A-U

W-1709/2020

Wales/PH

WC

-267A5/2020

Belgium/VHRA-030644/2020

Icel

and/

279/

2020

Scot

land

/ED

B108

/202

0


Can

ada

/MB

10/

202

0

Eng

land

/SH

EF-

C05

C1/

2020

Netherlands/N

A 78/2020



Brazil/SPBR-09/2020

Shang

hai/SH

0023/2020

Belgium

/ULG

-10053/2020

Eng

land

/CA

MB

-744

83/2

020

US

A/N

Y-P

V09

195/

2020

USA/WA-UW-1772/2020

Indi

a/10

73/2

020


Icel

and/

413/

2020

Scotland/C

VR

143/2020

England/CAMB-737D6/2020

US

A/N

Y-P

V09

099/

2020

USA/NJ-NYUMC106/2020

US

A/W

A-U

W-2230/2020

Belgium

/DB

A-03032/2020

Belgium/BGM-030444/2020

Engla

nd/2

0139

0475

04/2

020

HongKong/VM20001061/2020

US

A/W

A-U

W91/2020

Eng

land

/SH

EF

-BF

CE

E/2

020

Aus

tral

ia/V

IC84

/202

0

England/20116022902/2020

USA

/MA

9703

/202

0

US

A/W

A-U

W205/2020

Japan/N

A-20-05-1/2020

USA/W

A-UW

44/2020

Eng

land

/SH

EF

-BF

FA9/

2020

Belgium/DCS-0325158/2020

Scotland/C

VR

115/2020

Wales/P

HW

C-26E

64/2020

Sou

thK

orea

/KU

MC

02/2

020

US

A/W

A-S

26/2020


England/20129036004/2020


Wal

es/P

HW

C-2

6F52

/202

0

USA/WA-UW-4404/2020

Eng

land

/201

3402

3004

/202

0

Finlan

d/14

M74

/202

0

Engl

and/

2014

2019

704/

2020


US

A/W

A-U

W-1756/2020

Franc

e/PL1

643/20

20

Aus

tral

ia/V

IC25

0/20

20

Por

tuga

l/PT

0045

/202

0

USA/WI-71/2020

US

A/M

N41-M

DH

41/2020

IHU

CO

VID

-043

5

Bra

zil/S

PB

R-0

2/20

20

US

A/W

A-U

W152/2020

IHUCOVID-0304


Australia/V

IC46/2020

Iceland/410/2020

USA/WA-U

W-1

716/

2020

IHUCOVID-0225

Guangdong/GD2020233-P0027/2020

US

A/W

A-U

W-2247/2020

Netherlands/Blaricum 1364780/2020

Canada/BC

8150537/2020



England/201360198/2020

Iceland/581/2020


US

A/V

A-D

CLS

-000

9/20

20

USA/IN

2001/2020



Australia/N

SW

37/2020

USA/C

A-CD

PH-U

C7/2020

Netherlands/N

A 54/2020

Germany/NRW-40/2020

Wales/PHWC-24CF3/2020

Engla

nd/N

OTT

-10E

098/

2020

England/CAMB-76DFF/2020

Australia

/VIC168/2020


USA/W

I-91/2020

USA/WA-UW-1797/2020

Aus

tralia

/VIC

301/

2020

Netherlands/N

A 251/2020


Fran

ce/A

RA

0942

8/20

20

Iceland/77/2020

IHUCOVID-0491

USA/NY-PV08468/2020

IHU

CO

VID

-007

4

Icel

and/

202/

2020

IHUCO

VID-0

206

Japa

n/P

5-2/

2020

Aus

tral

ia/V

IC61

/202

0

Aus

tral

ia/V

IC02

/202

0

USA/AZ-ASU2936/2020


USA/WA-UW-1854/2020

Australia/N

SW

38/2020

Englan

d/20

1320

7930

4/20

20

USA/WA-U

W-1872/2020

Belgium

/PC

-0325147/2020

IHUCOVID-0396


Englan

d/20

1300

3650

4/20

20



Georgia/Tb-477/2020

Engla

nd/2

0134

0269

04/2

020

England/SHEF-C06BF/2020

USA/W

A-S76/2020

USA/WI-98/2020

Belgium

/MJP

-030

6100/2

020

Iceland/451/2020

US

A/W

A-N

H4/2020

Iceland/50/2020


Iceland/15/2020


Eng

land

/201

4208

3604

/202

0

Wales/PHW

C-26F25/2020

Portugal/P

T0010/2020

Belgium

/UG

ent-4/2020

USA/WI-G

MF-0

0018

/202

0

Fra

nce/

IDF

0372

/202

0

USA/V

A-DC

LS-0

012/

2020


England/20100121007/2020

Eng

land

/201

3906

2504

/202

0

France/IDF3236/2020

England/SHEF-C0707/2020

Finland/13M26/2020

USA/NY-PV09103/2020

Belgium/PM-0324144/2020

Shenzhen/SZTH-002/2020

USA/SC 3575/2020

Sw

itzerland/1000477797/2020

USA/MN-UW257/2020

US

A/W

A-S

112/2020

Wales/PHWC-25E0B/2020


USA/MN-UW243/2020

Scotland/CVR19/2020

USA/CT-U

W254/2020

Fra

nce/

AR

A12

558/

2020

USA/W

A-UW

85/2020

USA/WA-U

W389/2020

USA/NY-W

adsw

orth-

1049

6-01

/2020

Ganzhou/JX81/2020

IHUCOVID-0110

Germany/NRW-24/2020

USA/WA-UW-1824/2020


Iceland/426/2020


USA/M

N3-M

DH

3/2020


USA/WA-UW-1956/2020

IHUCOVID-0066

Iceland/296/2020

Sco

tland

/CV

R10

/202

0

Sco

tland

/ED

B10

2/20

20

Wales/PHWC-2624A/2020

Scotland/CVR34/2020

US

A/W

A-S

43/2020

Iceland/127/2020

USA

/MA

8932

/202

0




US

A/W

A-U

W378/2020


USA/NY-PV09025/2020

US

A/W

A-S

98/2020




US

A/W

A-S

107/2020

Chi

na/W

H-0

9/20

20

Georgia/Tb-673/2020

Aus

tral

ia/V

IC18

3/2

020

Netherlands/U

trecht 7/2020

Shang

hai/SH

0125/2020

Sco

tland

/CV

R13

6/20

20

US

A/A

Z-A

SU

2923

/202

0

Iceland/130/2020

Iceland/176/2020


Iceland/282/2020

IHUCOVID-0445

Engl

and/

SHEF

-C04

E2/2

020

USA/WI-10/2020

US

A/M

N62-M

DH

62/2020



IHUCOVID-0296


IHUCOVID-0371


Canada/BC 4540462/2020

Por

tuga

l/PT

0087

/202

0

Eng

land

/201

3003

4904

/202

0

Iceland/510/2020


Canada/B

C 6981299/2020

England/20132079204/2020

Braz

il/ES

-225

/202

0

Aus

tral

ia/N

SW

56/2

020

IHUCO

VID-0

286


Australia/V

IC265/2020

Canada/O

N P

HL1083/2020

England/20124003702/2020

Finland/FIN03032020A/2020

Netherlands/N

A 216/2020


Net

herla

nds/

NA

35/

2020

Finland/13M57/2020

Netherlands/N

A 25/2020

Can

ada/

ON

PH

L053

9/20

20

Hon

gKon

g/H

KP

U36

-070

2/20

20

USA/RI 0520/2020

Wales/P

HWC-2

43EF/2020

US

A/W

A-U

W207/2020


Taiw

an/C

GM

H-C

GU

-05/

2020

Bel

gium

/ULG

-100

09/2

020

Fran

ce/ID

F317

0/20

20

USA/UT-00019/2020


Netherlands/ZuidH

olland 97/2020


Iceland/217/2020

USA/W

I-115/2020

US

A/W

A-S

115/2020

USA/WI-G

MF-00707/2020

Bel

gium

/ULG

-974

1/20

20

Finland/FIN-455/2020

Hon

gKon

g/V

M20

0028

68/2

020

US

A/W

A-U

W-1334/2020

Italy/INM

I4/2020


US

A/W

A-U

W-1

864/

2020

Portugal/P

T0054/2020

Wales/PHW

C-25472/2020

Eng

land

/201

3802

82/2

020

Scotland/C

VR

144/2020


Icel

and/

550

/202

0


Bel

gium

/ULG

-701

9/20

20

US

A/W

A-U

W175/2020


India/1135/2020

US

A/M

N134-M

DH

134/2020

Eng

land

/201

4007

3904

/202

0

Fosha

n/20

SF210/

2020


Por

tuga

l/PT

0080

/202

0

Belgium

/KN

-030756/2020

Wales/PHW28/2020

Japa

n/P

5-3

/202

0


Net

herla

nds/

NA

104/

2020

Netherlands/N

A 288/2020

Iceland/6/2020

USA/NY-NYUMC7/2020


Italy

/TE

6222

/202

0

Iceland/573/2020

Belgium

/VJP

-030463/2020

Aus

tralia

/VIC

79/2

020

IHUCOVID-0006

US

A/N

Y-W

adsw

orth

-106

83-0

1/20

20

Seneg

al/61

1/20

20

US

A/W

A-U

W293/2020

Wale

s/PHW

C-25F

EA/202

0

Chile/Santiago-1/2020

Belgium/030959/2020

USA/CZB-RR057-015/2020

IHUCO

VID-0

205

USA/NY-NYUMC63/2020


Aus

tral

ia/N

SW

09/2

020

Eng

land

/201

2801

4806

/202

0

Finland/13M83/2020

Engl

and/

2014

4071

504/

2020

Eng

land

/201

3400

3404

/202

0

USA/NY-PV09199/2020

Netherlands/G

elderland 3/2020

Icel

and/

196/

2020

Netherlands/Z

uidH

olland 16/2020

DRC/80/2020



Engl

and/

2014

2026

304/

2020

USA/W

A-S95/2020


Belgium/DA-030691/2020

Eng

land

/201

2903

6404

/202

0

England/20140048704/2020


England/20109099106/2020

USA/RI 0556/2020

Belgium/PA-030680/2020

Austria/CeM

M0003/2020

IHUCOVID-0323

Beijing/231/2020

Iceland/102/2020


Iceland/255/2020

USA/NY-PV08432/2020


USA/NY-PV08494/2020


Shang

hai/SH

0080/2020

Eng

land

/201

3003

6304

/202

0

Iceland/315/2020

Wuh

an/H

BC

DC

-HB

-03/

2019

Engl

and/

2013

6031

7/20

20

Spain/CastillaLaMancha201329/2020

Netherlands/O

verijssel 1/2020

US

A/W

A-U

W103/2020

US

A/W

A-S

10/2020

Hon

gKon

g/H

KP

U39

-300

1/20

20

Icel

and/

536/

2020

IHUCOVID-0120



Iceland/305/2020

USA/W

A-UW

19/2020


Netherlands/N

A 163/2020

England/20110000706/2020

Eng

land

/201

3202

3004

/202

0

Icel

and/

366/

2020


Iceland/200/2020


Belgium/MS-0324134/2020


Wal

es/P

HWC-2

64DB/2

020

Germany/NRW-23/2020


USA/NY-PV09151/2020

Iceland/190/2020

Wales/PHW

C-2423D/2020

Canada/O

N P

HL3802/2020

England/20142024404/2020

England/201360217/2020

IHUCOVID-0487

Scotla

nd/E

DB106/

2020

IHUCOVID

-0097

IHUCOVID-0430

Belgium/DCB-0324163/2020

Belgium/JLJ-0320101/2020

Scotland/CVR57/2020

USA/CT-UW-4372/2020


Sco

tland

/ED

B11

8/20

20

USA/FL 9656/2020

Netherlands/N

A 186/2020

IHU

CO

VID

-044

0

Iceland/439/2020

USA/NY-PV09044/2020

US

A/A

Z-A

SU

2922/2020


Shanghai/S

H0047/2020

IHUCOVID-0385



Franc

e/ID

F2533

/202

0

Austra

lia/Q

LDID

929/

2020

England/201360057/2020

Wales/PHW

C-2672D/2020

Hungary/2/2020

SaudiA

rabia/K

AIM

RC

-Alghoribi/2020

USA/NY-PV08472/2020




IHUCOVID-0195

USA/NY-NYUMC26/2020

Netherlands/N

A 38/2020

US

A/N

Y-P

V08

471/

2020


US

A/W

A-U

W383/2020

Iceland/336/2020

Iceland/210/2020


USA/NY-PV08148/2020

Iceland/303/2020

Icel

and/

326/

2020

Iceland/313/2020

IHUCOVID

-038

8



Shanghai/S

H0033/2020

US

A/W

A-S

24/2020

IHUCOVID

-042

9

Eng

land

/201

3903

3004

/202

0

US

A/W

A4-U

W2/2020

IHUCOVID

-008

6

US

A/W

I-43/

2020


USA/WA-U

W-1620/2020

Icelan

d/30/2

020


Shanghai/S

H0112/2020

Wales/P

HW

33/2020

Franc

e/ID

F3274

/202

0

USA/WA-UW122/2020

Austra

lia/V

IC22

0/20

20

England/20106005403/2020

Scotland/E

DB

042/2020

Eng

land

/SH

EF

-C08

31/2

020

Engl

and/

SHEF

-C00

66/2

020

France/GE1973/2020



Netherlands/N

A 225/2020

Belgium/DB-0323140/2020

Senegal/370/2020

Scot

land

/CVR

114/

2020

Taiw

an/NTU05/2020

Net

herla

nds/

Tilb

urg

1364

286/

2020

Australia/V

IC163/2020

Finl

and/

14M

26/2

020

Icel

and/

576/

2020

USA/WA-UW-2179/2020

IHUCOVID-0417

Engl

and/

2009

9072

5/20

20


Fran

ce/B

2334

/202

0

Belgium

/ULG

-6216/2020


Iceland/156/2020

USA

/MA

2652

/202

0


USA/NY-PV09130/2020

USA/WI-74/2020

Germany/BavPat3/2020

Wales/PHW

32/2020

Aus

tral

ia/V

IC23

8/20

20


Austra

lia/N

SW62

/202

0

Iceland/63/2020


England/20139070204/2020

IHUCO

VID-0

451

USA/SC 3569/2020


Indi

a/11

11/2

020

US

A/W

A-S

59/2020

USA/WA-UW115/2020

England/CAMB-74D6A/2020

Iceland/381/2020

Scotland/E

DB

120/2020

USA/WA-UW178/2020

IHUCO

VID-0

189

Eng

land

/201

3400

4904

/202

0

Icel

and/

572/

2020

Netherlands/N

oordBrabant 59/2020

Kuwait/KU001/2020

Aus

tral

ia/V

IC12

0/20

20

Fuya

ng/F

Y00

2/20

20

Belgium

/VT-0325143/2020

Iceland/382/2020

Wales/P

HW

C-25542/2020

France/IDF2534/2020

IHUCOVID-0297

IHU

CO

VID

-021

5

Australia/V

IC94/2020

Austra

lia/V

IC19

3/20

20

US

A/W

A-S

48/2020

Iceland/318/2020

Australia/V

IC315

/2020




Shangrao/JX1948/2020

Belgium/UGent-13/2020

USA

/MA

3616

/202

0

Por

tuga

l/PT

0073

/202

0

IHUCOVID-0193

USA/UT-00364/2020


Iceland/72/2020

Netherlands/N

A 293/2020

Wales/P

HWC-2

708B/2020

Nor

way

/153

8/20

20

US

A/VA

-DC

LS-0004/2020

USA/WA-UW-1904/2020

IHUCO

VID-0

064


US

A/W

A-U

W13

7/20

20


US

A/M

N26-M

DH

26/2020

Wuhan/IP

BC

AM

S-W

H-03/2019

Netherlands/N

A 62/2020

USA/NY-N

YUMC11

9/20

20


Eng

land

/201

0905

0806

/202

0

Australia

/VIC22/2020


Nethe

rland

s/Lim

burg

4/2

020


US

A/M

N53

-MD

H53

/202

0

Bel

gium

/ULG

-962

0/20

20

USA/NY-NYUMC62/2020

IHUCOVID-0094

Engla

nd/2

0138

0064

/202

0

IHUCOVID-0016

Wales/P

HWC-25463/2020

England/20139008404/2020

Wales/PHWC-26EEC/2020

US

A/W

A-N

H20/2020

Fran

ce/A

RA

1222

2/20

20

Bel

gium

/ULG

-100

29/2

020

Belgium/WWM-030665/2020


Aus

tral

ia/V

IC29

4/20

20

US

A/W

A-N

H19/2020

Icel

and/

601/

2020

IHUCOVID-0046

USA/NY-PV08445/2020

England/20130064804/2020

Iceland/182/2020


US

A/W

A-U

W-1578/2020

Wales/PHWC-25C3E/2020

Canada/BC 83109/2020

Latvi

a/00

04/2

020


Wales/P

HW

C-26C

E2/2020

USA/NY-NYUMC43/2020

Germ

any/BA

V-V

2011376/2020

Belgium/C

F-0324119/2020

USA/WI-GMF-00232/2020

Net

herla

nds/

Zui

dHol

land

81/

2020

Aus

tral

ia/V

IC21

2/20

20

Eng

land

/201

3904

7104

/202

0


Eng

land

/201

4401

8704

/202

0

USA/GA 2343/2020

Iceland/364/2020

Nethe

rland

s/Zuid

Hollan

d 27

/202

0

Sco

tland

/CV

R21

/20

20

Wal

es/P

HW

C-2

6D0D

/202

0


Scotland/C

VR

27/2020

Eng

land

/201

4804

3704

/202

0

Sw

itzerland/GR

2988/2020


Iceland/338/2020

USA/NY-NYUMC44/2020

US

A/W

A-U

W189/2020

England/CAMB-7477B/2020

USA/ID-UW-4375/2020

Tur

key/

ER

AG

EM

-001

/202

0


USA/W

A-UW

176/2020

China/IQ

TC

01/2020

Por

tuga

l/PT0

021/

2020


US

A/W

A-S

62/2020

USA/WI-GMF-00557/2020

Mexico/EdoM

ex-InDRE 03/2020

US

A/W

A-U

W-1782/2020

Netherlands/N

A 231/2020

Iceland/503/2020

Ger

man

y/N

RW

-19/

2020

Belgium/GS-030549/2020



Taiwan/NTU09/2020


USA/WI-46/2020

Belgium/RS-030257/2020

Spa

in/V

alen

cia1

1/20

20

Iceland/36/2020

IHU

CO

VID

-0051


Geo

rgia

/Tb-

468/

2020


Wales/PHWC-264BD/2020

Iceland/287/2020

Belgium

/ULG

-10068/2020

Eng

land

/SH

EF-

C01

72/2

020

US

A/W

I-12/

2020

Netherlands/N

A 8/2020

Englan

d/CAM

B-770

22/2

020

Aust

ralia

/VIC

307/

2020

Iceland/599/2020

France/GE3067/2020

US

A/W

A-U

W130/2020


USA/UT-

0001

0/20

20

Iceland/82/2020

Eng

land

/201

3401

6802

/202

0


USA/W

A-UW

-1765/2020

Belgium/GE-030573/2020

Singapore/32/2020

Sco

tland

/CV

R46

2/20

20

Wales/P

HW37/2020

IHUCOVID-0058

England/20129017104/2020


Engl

and/

2013

2024

904/

2020

England/20144074504/2020

Netherlands/R

otterdam 1364740/2020

Canada/ON P

HL3680/2020

Bel

gium

/DM

E-0

3241

35/2

020

IHU

CO

VID

-011

5


Australia/V

IC08/2020

US

A/W

A-S

109/2020

Bel

gium

/ULG

-963

4/20

20

IHUCO

VID-0

109


USA/WA-UW296/2020

Can

ada/

BC

597

9789

/202

0

Arg

ent

ina/

C12

1/2

020

Guangdong/2020XN4243-P0035/2020

Netherlands/G

elderland 8/2020

Australia/V

IC237

/2020

USA/UT-00301/2020

England/20144037304/2020

Finl

and/

FIN

0303

2020

B/2

020

Eng

land

/201

0903

5906

/202

0

Net

herla

nds/

NA

244/

2020

Belgium/VAG-03013/2020

IHU

CO

VID

-005

5

US

A/W

A-N

H8/2020

Portugal/PT0011/2020Belgium/BCM-0324160/2020

Finland/14M14/2020

US

A/D

C 0004/2020


IHUCO

VID-0

226

Belgium

/DJ-030452/2020

USA/SC 3571/2020

US

A/W

A-U

W33/2020


Belgium/OT-0319109/2020

Italy/INMI8/2020

Eng

land

/CA

MB

-770

7D/2

020

Iceland/578/2020

Australia/N

SW

44/2020

Belgium/CJM-0323175/2020

Eng

land

/NO

TT-1

0DF7

E/2

020

England/20144076304/2020

Eng

land

/201

4201

9004

/202

0


Por

tuga

l/PT

0047

/202

0

US

A/W

A-U

W217/2020


USA/W

A-UW

367/2020


Senegal/020/2020

Iceland/270/2020

USA/CA5/

2020

USA/CA8/2020

Guangdong/GDSZ202015-P0019/2020

Taiw

an/C

GM

H-C

GU

-03/

2020


Taiw

an/C

GM

H-C

GU

-12/

2020

USA

/MA

9704

/202

0

France/IDF2936/2020


IHUCO

VID-0

288


Wuhan/IVDC-HB-04/2020




Eng

land

/CA

MB

-744

74/2

020


Austra

lia/V

IC25

8/20

20

Beijing/235/2020

Nan

chan

g/JX

155/

2020

Iceland/113/2020

US

A/W

A-U

W-2

232/

2020

US

A/W

A-U

W-1295/2020

USA/NY-N

YUMC89/2020

Iceland/473/2020

Arg

entin

a/C

1374

/202

0

USA/NY-PV09134/2020

England/20132090604/2020

Australia/V

IC110/2020

England/20140064004/2020

Por

tuga

l/PT

0086

/202

0

Singapore/6/2020

USA/WA-UW248/2020

Engl

and/

2012

9037

204/

2020

Iceland/140/2020

England/20140000604/2020

USA/W

I-49/2020

Eng

land

/NO

TT-1

0EA

6D/2

020


IHUCOVID-0083

Belgium/BEG-0324149/2020

Iceland/74/2020

Engl

and/

2012

8003

506/

2020

IHUCOVID-0347


Australia/W

A09/2020


DRC/94/2020

USA/ID-UW-2127/2020

Belgium/VL-0318104/2020



USA/MN52-M

DH52/2020


US

A/W

A-U

W-1341/2020

Hon

gKon

g/H

KP

U44

-010

2/20

20

DRC/214/2020

Scotla

nd/CVR29/2020

Norway/1811/2020

Eng

land

/201

4804

4604

/202

0

US

A/W

I-63/

2020


Iceland/325/2020

USA/NY-PV09079/2020


US

A/W

A-S

83/2020



Canada/O

NP

HL4181/2020

Wales/P

HWC-2

6A51/2020

USA/WA-UW-1690/2020


Sw

eden

/01/

2020

Hon

gKon

g/V

M20

0029

07/2

020

US

A/W

A-U

W15/2020

US

A/M

N64-M

DH

64/2020

Eng

land

/201

3007

1504

/202

0

England/20139072204/2020


Xinyu/JX122/2020

IHU

CO

VID

-007

8

Sco

tland

/ED

B31

4/20

20

Engla

nd/2

0128

0144

04/2

020

US

A/W

A-S

75/2020

US

A/W

A-U

W270/2020

US

A/W

A-U

W-1778/2020

France/B

2349/2020



US

A/W

A-S

52/2020


India/1-31/2020

Hon

gKon

g/H

KP

U74

-130

2/20

20

Sco

tland

/ED

B39

6/20

20

Senegal/139/2020

Scotland/CVR28/2020

Net

herla

nds/

NA

14/

2020

IHUCOVID-0216

Wales/PHW

C-24525/2020


Net

herla

nds/

NA

85/

2020

USA/NY-N

YUMC42/2020

US

A/W

A-U

W52/2020


US

A/N

Y1-

PV

0800

1/20

20

IHUCOVID

-029

3

Wal

es/P

HWC-2

6BA9/

2020

Belgium

/FM-030592/2020

US

A/W

A-U

W111/2020

US

A/W

A-U

W222/2020

US

A/T

X 2039/202

0

IHUCOVID-0497

Hangzhou/ZJU

-05/2020

Netherlands/U

trecht 2/2020

Brazil/S

PB

R-13/2020

Canada/BC 3968175/2020

USA/WA-UW193/2020

Eng

land

/201

4202

3704

/202

0

Norway/1951/2020

Netherlands/N

A 64/2020

USA/W

A-UW

26/2020

USA/NY-PV09125/2020

Wales/PHW

C-25779/2020

Spain/Madrid LP15 4/2020


England/C

AM

B-7708C

/2020

Australia/V

IC199/2020

USA/W

A-UW

220/2020


US

A/W

A-U

W-1855/2020

Belgium

/BD

W-030694/2020

Eng

land

/201

0403

5803

/202

0

Netherlands/Z

uidHolland 18/2020

England/NOTT-10E2FC/2020

USA/MN45-MDH45/2020

France/HF1871/2020

IHUCOVID-0047

Eng

land

/201

4803

9404

/202

0


USA

/PA

2317

/202

0

USA/WA-UW134/2020

Wal

es/P

HW

C-2

6981

/202

0

US

A/N

Y-NY

UM

C95/2020

Belgium

/MA

C-0324142/2020



IHUCOVID

-004

5

England/20104008402/2020

Germany/NRW

-34/2020

Ger

man

y/B

AV

-V20

1285

6/20

20

Australia/WA11/202

IHU

CO

VID

-021

0


Eng

land

/201

3800

44/2

020

Wales/PH

WC

-268A2/2020

USA/WA-UW-1994/2020

Scotland/CVR18/2020


USA/NY-PV08459/2020

Aus

tral

ia/V

IC27

2/20

20

Eng

land

/201

3007

1804

/202

0

England/20132004704/2020

Iceland/412/2020

USA/NY-PV08444/2020


US

A/W

A9-U

W6/2020


Fran

ce/H

F274

8/20

20



USA/W

A-UW

38/2020

USA/WI-06/2020

Eng

land

/201

3902

2704

/202

0

Germ

any/BA

V-V

2010492/2020

USA/IA 6395/2020

Eng

land

/SH

EF-

C08

40/2

020

France/IDF2359/2020


Zhejia

ng/W

Z-02/

2020

France/IDF3230/2020

Scotland/CVR77/2020

Italy

/VR

20C

OV

21-

26/2

020

Sco

tland

/CV

R20

6/20

20


Englan

d/20

1320

8040

4/20

20

IHUCO

VID-0

191

Belgium/DBOM-030566/2020

USA/WI-104/2020

USA/WA-UW-1707/2020

USA/WI-22/2020


IHUCOVID-0119

Wales/PHW

C-260F5/2020

Iceland/582/2020

Malaysia/188407

/2020

Sha

ngha

i/SH

0012

/202

0

Wales/P

HW

C-25D

D1/2020


Eng

land

/201

4206

2204

/202

0

Iceland/356/2020

Nanchang/JX39/2020


US

A/W

A-U

W19

7/20

20


Aus

tral

ia/V

IC10

9/20

20

Sw

itzerland/1000477757/2020


Iceland/360/2020

USA/W

I-81/2020

USA/W

A-UW

202/2020

USA/NY-PV09056/2020


Iceland/89/2020

Aus

tral

ia/V

IC25

/202

0

Iceland/290/2020

USA/IN-Lilly-IPB0170-4bc20/2020

Eng

land

/201

3004

9804

/202

0

Icel

and/

203/

2020

Iceland/321/2020


IHU

CO

VID

-040

1

Iceland/368/2020

England/20134016902/2020

US

A/W

A-U

W-1843/2020

Wuh

an/W

HU01/2

020

USA/CT-UW273/2020

Engla

nd/2

0132

0327

04/2

020


USA/W

A-S114/2020

Netherlands/N

A 213/2020

USA/WA-UW365/2020

Belgium/RT-03035/2020


Wales/PHW

C-2573D/2020

Belgium

/GJ-030458/2020

USA/NY-PV09097/2020



USA/W

A-UW

84/2020

US

A/C

A3/2020

Iceland/495/2020

Iceland/370/2020

Eng

land

/201

3401

9904

/202

0

Ireland/Limerick-19934/2020

Aus

tral

ia/V

IC25

2/20

20

USA/NY-NYUMC27/2020


Aus

tralia

/VIC

169/

2020

Eng

land

/201

3210

8404

/202

0


Belgium

/VS

-030542/2020

US

A/W

A-U

W195/2020

IHU

CO

VID

-020

0

England/20132081104/2020


Fran

ce/L

yon

0668

/202

0

US

A/C

T-U

W22

3/20

20

IHU

CO

VID

-019

2

Australia/V

IC134/2020

Icel

and/

457/

2020

Gua

ngdo

ng/G

D20

2008

7-P

0008

/202

0

Aus

tral

ia/N

SW

19/2

020



USA/WA-U

W-2

236/2020


USA/WA-U

W-1680/2020

Iceland/21/2020

Net

herla

nds/

NA

60/

2020

Wales/PHW

C-26DEF/2020


US

A/W

A-S

42/2020

Nonth

abur

i/61/

2020




Germany/NRW-12/2020

IHU

CO

VID

-0434

USA/W

A-S11/2020

Eng

land

/SH

EF

-BF

ED

9/20

20

US

A/N

Y-P

V09

194/

2020

England/20129023104/2020

USA/NY-NYUMC33/2020

Australia/V

IC269/2020


Germany/NRW-44/2020

IHUCO

VID-0

126

Germany/N

RW-1

4/2020

Icel

and/

119/

2020

Hon

gKon

g/H

KP

U29

010

2/20

20

England/CAMB-745BD/2020

Kuw

ait/K

U18

/202

0

USA/WI-54/2020

US

A/N

Y-N

YU

MC

51/2

020

Por

tug

al/P

T00

23/2

020

US

A/W

A-S

17/2020

Belgium/DA-0319106/2020



Canada/BC 4118226/2020

USA/MN31-MDH31/2020

Eng

land

/NO

TT-1

0E06

B/2

020

Singa

pore/41/2020

US

A/W

A-S

105/2020

Iceland/333/2020

Shandong/LY003/2020

Wal

es/P

HW

C-2

46B

9/20

20

Eng

land

/200

9600

41/2

020

Belgium/RS-0324138/2020

France/H

F1988/2020


US

A/W

A-U

W-1297/2020

Scotland/CVR12/2020


USA/WA-UW-1865/2020

USA/WA-UW-1947/2020

Wales/P

HW

C-26E

A0/2020

Iceland/133/2020


US

A/W

A-U

W79/2020

Taiw

an/1

70/2

020

Engla

nd/SH

EF

-BF

EE

8/2020

Por

tug

al/P

T00

34/2

020

Iceland/53/2020

Iceland/257/2020

USA/WA-UW-1888/2020

USA/NY-NYUMC19/2020


USA/WI-94/2020

USA/W

I-08/2020

Icel

and/

579/

2020

Wales/P

HW

C-26796/2020

US

A/W

A-U

W29

5/20

20

Argentina/C3013/2020

IHU

CO

VID

-003

8

England/201380050/2020

Iceland/123/2020

Iceland/367/2020

Engla

nd/2

0134

0609

04/2

020

Icel

and/

570/

2020

Bel

gium

/ULG

-100

66/2

020



USA/WI-82/2020

Vie

tnam

/CM

296/

2020

Japa

n/P

4-2/

2020

Kor

ea/B

A-A

CH

260

4/20

20

England/20142078304/2020

Singa

pore/42/2020

Eng

land

/09c

/202

0


France/IDF2848/2020

Singapore/28/2020

USA/W

A-UW

-1320/2020


England/201360224/2020

France/HF2595/2020

USA/NY-NYUMC80/2020

US

A/W

A-U

W375/2020


US

A/F

L 5125/2020

Senegal/618/2020

India/c32/2020

USA/UT-0

0034/2020

USA/WA-UW-1827/2020

Taiw

an/C

GM

H-C

GU

-11/

2020

USA/WI-75/2020

Eng

land

/201

3110

3404

/202

0

Wal

es/P

HW

C-2

6444

/202

0

Denmark/SSI-101/2020

Engla

nd/2

0132

0674

04/2

020

Aus

tralia

/VIC

23/2

020

Wal

es/P

HW

C-2

5946

/202

0

IHU

CO

VID

-025

6

US

A/A

Z 4811/2020

Fran

ce/A

RA

1055

2/20

20


Canada/BC 8718874/2020

USA/WI-92/2020

US

A/W

A-U

W-1730/2020

SouthAfrica/KRISP 0011/2020

Xinyu/JX

124/2020

Netherlands/N

A 12/2020

Iceland/168/2020

US

A/W

A-U

W-1583/2020

Engl

and/

CAM

B-76

EB0/

2020



USA/AK-PHL002/2020

Iceland/437/2020

France/IDF2989/2020

Australia

/VIC147/2020

Wales/PHW

C-26013/2020

Fran

ce/G

E19

77/2

020

Austra

lia/V

IC26

/202

0

Japa

n/P

2-2/

2020

USA/NY-PV08487/2020

Wales/PHWC-2715B/2020


Shandong/LY005/2020


Sco

tland

/CV

R47

/202

0

Iceland/134/2020

Iceland/233/2020


Singapore/20/2020

Scotland/CVR76/2020


IHUCOVID-0167

Bel

gium

/ULG

-100

26/2

020


Australia/V

IC162/2020


US

A/W

A-U

W-1

826/

2020

Sco

tland

/CV

R13

4/20

20

IHUCO

VID-0

327


US

A/W

A-U

W-2082/2020

Iceland/29/2020

Eng

land

/201

3400

7204

/202

0

IHUCOVID

-004

4

IHU

CO

VID-0

366

Wales/PHW

C-2419A/2020

Iceland/109/2020

Net

herla

nds/

Noo

rdB

raba

nt 2

6/20

20

Austri

a/CeM

M00

01/2

020

Iceland/394/2020

US

A/W

A-U

W32

2/20

20

US

A/W

A-U

W-1

745/

2020

USA/UT-00023/2020

USA/ID-UW-4355/2020

US

A/W

A-U

W209/2020

Netherlands/Z

uidHolland 14/2020


Norw

ay/1867/2020


USA

/VA-

DC

LS-0

016/

2020

England/20144006004/2020

Aus

tral

ia/V

IC15

6/20

20


USA/WA-U

W-1799/2020

England/20140005304/2020

Georgia/Tb-273/2020

Engl

and/

2012

0002

102/

2020

Bel

gium

/ULG

-100

03/2

020


Netherlands/N

A 26/2020

USA/ID-UW-2160/2020

US

A/N

Y-P

V08

425/

2020



Sin

gapo

re/3

9/20

20


Eng

land

/CA

MB

-76D

95/2

020

Iceland/249/2020


Australia/V

IC181

/2020

US

A/W

A-U

W-2

181/

2020

Germany/NRW

-33/2020

USA/WA-UW-2100/2020


USA/NY-PV09060/2020


Eng

land

/CA

MB

-737

C7/

2020

Bel

gium

/FM

C-0

3231

59/2

020

Engl

and/

2013

4023

104/

2020


England/20126007102/2020

USA/WI-40/2020

Bel

gium

/PC

-032

5182

/202

0



Iceland/183/2020

Slov

akia

/SK-

BMC

2/20

20

Engla

nd/2

0136

0143

/202

0

Icel

and/

411/

2020

Taiwan/NTU12/2020

Belgium

/DB

D-03024/2020

Belgium

/RY

R-030649/2020

Canada

/ON

PH

L0141/2020

Australia/V

IC43/20

20

Iceland/175/2020

USA/NY-PV08436/2020

Por

tuga

l/PT

0078

/202

0

England/C

AM

B-73888/2020

IHUCO

VID-0

214

IHU

CO

VID

-028

0

IHUCOVID-0232



Sco

tland

/ED

B56

6/20

20

Wales/P

HW

C-26D

D0/2020

Sha

ngha

i/SH

0056

/202

0

US

A/W

A-N

H23/2020

USA/NJ 3592/2020

Wale

s/PHW

C-23E

2B/2

020


IHUCO

VID-0

492

Mexico/CDMX-INER 01/2020

USA/WI-95/2020

US

A/W

A-U

W180/2020

Wales/PHW

C-244CE/2020

Iceland/171/2020

Belgium/CG-0

30158/2020

US

A/W

A-U

W364/2020

England/20129059204/2020

US

A/W

I-15/2020

Iceland/460/2020

Net

herla

nds/

NA

299

/202

0

Switzerland/TI9486/2020


USA/W

A-UW

-1708/2020

England/20126096602/2020

Bel

gium

/ULG

-667

0/20

20

Australia

/VIC175/2020

IHU

CO

VID-0501

Iceland/416/2020

Japa

n/P

3-2/

2020

IHUCOVID-0290

Iceland/316/2020


US

A/W

A-U

W183/2020

Iceland/568/2020



Hon

gKon

g/V

M20

0024

93/2

020

US

A/IL-U

W350/2020

US

A/W

A-S

40/2020

Iceland/19/2020

US

A/W

A-U

W316/2020

Sw

itzerland/100047779

6/2020

Belgium/ULG-7

626/2020

Iceland/602/2020


Bel

gium

/BE

G-0

3231

54/2

020

Chile/Talca-1/2020

Engla

nd/2

0132

0825

04/2

020

Eng

land

/201

3605

06/2

020

Bra

zil/S

PB

R-1

0/20

20


Scotland/CVR72/2020

Netherlands/N

A 9/2020

USA/CZB-RR057-013/2020

Bel

gium

/DV

-032

4117

/202

0

France/HF2381/2020

Hangzhou/H

ZCDC0162/2020

Australia

/VIC

185/2020

Netherlands/U

trecht 4/2020

Eng

land

/201

3203

16/2

020

Norway/1526/2020

Iceland/590/2020

England/CAMB-76FF9/2020


Belgium

/ULG

-6942/2020



Senegal/600/2020

Eng

land

/201

3806

3804

/202

0

US

A/W

A-U

W-1296/2020

Iceland/216/2020

US

A/W

A-U

W-1968/2020

Luxembourg/LN

S7866283/2020

USA/NY-PV08149/2020

Wales/PHW

C-26CA6/2020

USA/NY-PV09157/2020

Belgium/PJM-0325166/2020

Finl

and/

13M

64/2

020

Iceland/419/2020

England/201380055/2020


Belgium

/MM

J-03034/2020

IHUCOVID-0125



USA/CT-UW-1365/2020

Belgium

/TG-030757/2020


Sau

diA

rabi

a/S

CD

C-3

321/

2020

Eng

land

/201

4602

5304

/202

0

Bel

gium

/ULG

-663

8/20

20

USA/WI-56/2020

USA

/MA

3626

/202

0



Germany/NRW-47/2020


Engla

nd/2

0132

0231

04/2

020

Bel

gium

/ML-

0324

164

/202

0

USA/WA-UW-2132/2020

England/NOTT-10E21A/2020

Eng

land

/CA

MB

-76F

CC

/202

0

USA/CA1/2020

Aus

tral

ia/V

IC58

/202

0

Canada/O

N P

HL0654/2020

Wales/P

HWC-2

5120/2020

Iceland/589/2020

Bel

gium

/ULG

-100

45/2

020

US

A/W

A-S

6/2020

Netherlands/N

oordBrabant 14/2020

Hon

gKon

g/H

KP

U52

-310

1/20

20

Canada/BC 9345715/2020

Bel

gium

/DN

S-0

3241

76/2

020

Bel

gium

/ULG

-100

57/2

020

Net

herla

nds/

NA

269/

2020

Australia

/VIC104/2020

Aus

tral

ia/N

SW

40/2

020


Iceland/294/2020

Fran

ce/H

F217

4/20

20

Net

herla

nds/

NA

201

/202

0


Iceland/226/2020

USA/WI-88/2020

France/BFC2094/2020

Iceland/425/2020

England/CAMB-745AE/2020

Icel

and/

499/

2020

US

A/W

A-U

W-1775/2020

USA/NH 0033/2020

Icel

and/

230/

2020

USA/NY-PV09092/2020

Wales/P

HWC-2

6D1C/2020

US

A/W

A-S

92/2020

USA/NY-N

YUMC11

8/20

20

USA/WA-UW269/2020

Scotland/CVR51/2020


USA/WA-UW-1748/2020

US

A/A

Z 8135/2020

USA/NY-PV09189/2020

US

A/F

L 11

77/2

020

Shandong/LY004/2020

USA/WA-UW268/2020

US

A/W

A-N

H11/2020

Engl

and/

2011

0136

302/

2020

USA/NY-PV09139/2020

US

A/W

A-U

W-1913/2020

IHUCO

VID-0

212

Wales/P

HWC-24A62/2020


USA/WA-UW-1470/2020

Iceland/455/2020


Australia/V

IC52/2020

USA/NY-PV08109/2020

Engla

nd/2

0130

0527

04/2

020

England/20139015304/2020



US

A/W

I-24/2020


Iceland/307/2020

Aus

tral

ia/N

SW

14/2

020

USA/UT-00008/2020

Greece/16/2020

US

A/W

A-U

W-1860/2020


Can

ada/

ON

PH

L353

6/20

20

Icelan

d/479

/2020

Aus

tral

ia/V

IC32

4/20

20


Franc

e/ARA12

388/

2020

USA

/NC

002

4/20

20

USA/NY-PV09169/2020


Iceland/583/2020

Sw

itzerland/BE

6651/2020


Guangdong/G

D2020234-P0023/2020

England/S

HE

F-C

0752/2020

France/GE2722/2020

Canada/B

C 3808524/2020

USA/NY-PV08446/2020

Canada/BC 8486790/2020

USA/NY-PV08125/2020

Eng

land

/SH

EF

-C02

8E/2

020

Taiw

an/C

GM

H-C

GU

-07/

2020

Iceland/241/2020

Australia/V

IC276

/2020

Wal

es/P

HW

C-2

465F

/202

0

France/ID

F0515/2020

Singapore/35/2020

USA/W

A-S94/2020

Australia/V

IC160/2020

USA/WA-UW-1723/2020

IHU

CO

VID

-0065

Australia/W

A04/2020

IHUCOVID-0439

England/20144008204/2020


USA/NY-NYUMC87/2020


USA/W

A-UW

162/2020

US

A/V

A 63

52/2

020

Belgium

/ULG

-10006/2020

Eng

land

/201

3801

2802

/202

0

Sco

tland

/ED

B10

1/20

20

Aus

tral

ia/V

IC21

9/20

20

Belgium/PAN-030681/2020

Australia/V

IC176/2020

Fin

land

/13M

33/2

020

Wales/P

HWC-26C97/2020

Taiw

an/N

TU

06/2

020

Wales/PHW

C-260B9/2020


Iceland/31/2020

IHU

CO

VID

-005

9

Russia/M

oscow P

MV

L-1/2020


US

A/N

Y-P

V09

301/

2020

USA/ID-UW-1935/2020



Finland/1/2020

USA/NY-PV09305/2020

Wuh

an/IP

BC

AM

S-W

H-0

4/20

19




USA/MN30-MDH30/2020

IHUCO

VID-0

162

Engl

and/

2013

9064

404/

2020

Belgium

/ULG

-3843/2020

USA/MN-UW251/2020


Sco

tland

/CV

R68

/20

20

USA/NY-PV08464/2020

USA/IA 6399/2020


Iceland/224/2020

USA/ID-UW-4380/2020

Singapore/34/2020

Engl

and/

2014

0070

004/

2020

Nanchang/JX14/2020

USA/NY-PV08486/2020

Gua

ngdo

ng/2

0SF04

0/20

20

Iceland/117/2020

Belgium/MC-0323129/2020

Spa

in/V

alen

cia4

5/20

20

Netherlands/N

A 275/2020

Finland/13M29/2020

USA/NY-PV09197/2020

Iceland/61/2020

Iceland/467/2020


Iceland/236/2020

France/GE2720/2020

IHU

CO

VID

-0300

CzechRepublic/951/2020

Icel

and/

146

/202

0

Wales/P

HWC-24419/2020

USA/WA-UW227/2020

Icel

and/

204

/202

0

Wale

s/PHW

C-255

7F/2

020

US

A/W

A-S

86/2020

IHU

CO

VID

-016

8

Wales/P

HW

C-266B

7/2020

IHU

CO

VID

-019

7


Eng

land

/CA

MB

-745

44/2

020

Belgium/BM-030687/2020


Iceland/427/2020


Wales/PHWC-26C3D/2020

USA

/KS

4126

/202

0

Net

herla

nds/

NA

29/

2020

USA/WI-45/2020

Georgia/Tb-1352/2020

Bel

gium

/VP

E-0

3065

0/20

20

USA/WA-UW-1879/2020

USA/WI-41/2020


Aus

tral

ia/W

A10

/202

0

USA/NY-PV08500/2020

USA/WA-UW282/2020

IHUCOVID-0084

Belgium/Human/C

S-031052/2020

Aus

tral

ia/V

IC97

/202

0

Eng

land

/SH

EF

-C03

5E/2

020


Eng

land

/201

3202

4104

/202

0

Iceland/436/2020


Por

tuga

l/PT

0049

/202

0

Eng

land

/CA

MB

-739

EF

/202

0

Belgium/NA-0323141/2020

US

A/W

A-U

W182/2020

Guangdong/G

D2020080-P

0010/2020

Irel

and/

2404

2/20

20

Engla

nd/C

AMB-7

3A0A

/202

0

Icel

and/

358/

2020

England/20134078004/2020

US

A/N

Y-PV

09075/2020

Belgium

/BG

-030955/2020

USA/NY-PV08452/2020

Engla

nd/2

0136

0220

/202

0


IHUCO

VID-0

444

USA

/VA-

DC

LS-0

005/

2020

Aus

tral

ia/V

IC26

8/20

20


Wales/P

HWC-26CC4/2020

Hon

gKon

g/H

KP

U19

040

2/20

20

US

A/N

Y-N

YU

MC

90/2020

France/H

F2946/2020

Wales/P

HWC-2

4F72/2020

Taiw

an/N

TU08

/202

0

USA/WA-UW155/2020

Scotland/CVR05/2020

USA/NY-PV09119/2020

Engla

nd/S

HEF-C02

24/2

020


Sw

itzerland/GE

3895/2020

Australia/V

IC305/2020

US

A/W

A-U

W10

8/20

20

US

A/W

A-S

55/2020

Eng

land

/201

4402

0004

/202

0

England/C

AM

B-74D

4C/2020

Icel

and/

170/

2020

US

A/W

A-U

W-1

722/

2020

Icel

and/

206/

2020

IHUCO

VID-0

455


Fran

ce/H

F199

3/20

20

US

A/W

A-S

47/2020

Eng

land

/201

2903

5404

/202

0

Belgium/VHC-030655/2020

Scotla

nd/E

DB22

7/20

20

Engla

nd/20142016504

/2020

Iceland/59/2020

Icel

and/

403/

2020

USA/NY-NYUMC94/2020

Australia/V

IC57/2020

England/CAMB-76EDE/2020

Wales/PHW

C-263FC/2020

Eng

land

/201

3210

4104

/202

0


US

A/F

L 95

90/2

020

Aus

tralia

/NS

W41

/202

0

IHUCOVID-0037

USA/MN2-M

DH2/2020

Aus

tral

ia/N

SW

12/2

020

Iceland/209/2020

Iceland/395/2020


US

A/N

Y-P

V09308/202

0

USA

/PA

2937

/202

0

Iceland/149/2020

Net

herla

nds/

NA

230

/202

0

USA/WA-UW-1269/2020

US

A/W

A-U

W144/2020

Icel

and/

554/

2020

France/IDF2548/2020

Hangzhou/HZ91/2020

IHUCO

VID-0

268

England/20132068304/2020

USA/W

A-UW

146/2020

Fra

nce/

AR

A12

371/

2020

US

A/W

A-S

69/2020

Portugal/P

T0055/2020

Eng

land

/201

3008

7204

/202

0


Belgium/CA-0319111/2020

Switzerland/AG0361/2020

Eng

land

/SH

EF

-C04

00/2

020

Indi

a/16

52/2

020

Scotland/CVR52/2020

IHUCOVID-0395

Guangdong/2020XN4459-P0041/2020

US

A/W

A-U

W286/2020

DRC/108/2020

Netherlands/N

A 285/2020

Japa

n/P

4-1/

2020

Canada/BC 6997898/2020

China

/IQTC

02/2

020

Netherlands/N

A 284/2020

Scot

land

/CVR

37/2

020

US

A/VA

-DC

LS-0001/2020

Iceland/10/2020


Aus

tral

ia/V

IC09

/202

0


Wales/P

HW

C-26C

5B

/2020

Belgium

/MF

-030546/2020

Eng

land

/201

3906

5204

/202

0

USA/W

A-UW

265/2020

Vie

tnam

/396

07/2

020

USA/WI-51/2020


Iceland/132/2020

Australia/V

IC204/2020

Shanghai/S

H0114/2020

US

A/U

T-00016/2020


Icel

and/

544/

2020

Netherlands/U

trecht 21/2020

Aus

tral

ia/N

SW

18/2

020

Ger

man

y/N

RW

-13/

202

0

IHUCO

VID-0

453

Iceland/365/2020

Kuw

ait/K

U17

/202

0

England/200990660/2020

Netherlands/N

A 1/2020

Belgium/GM-0323139/2020

USA/W

A-UW

-1840/2020



US

A/W

A-U

W-1300/2020

Senegal/315/2020

Netherlands/U

trecht 14/2020

England/20132047304/2020

Iceland/254/2020

Iceland/167/2020

Sw

itzerland/BS

0914/2020

Bel

gium

/SR

-031

9112

/202

0

USA/WA-UW287/2020

Engl

and/

2013

4019

502/

2020

Iceland/312/2020

USA/WA-UW-1435/2020

USA/NY-PV09087/2020

USA/W

I-80/2020

Eng

land

/201

0012

2106

/202

0

USA/WA-UW73/2020


IHU

CO

VID

-0238

Belgium

/ULG

-2971/2020

US

A/W

A-N

H6/2020

Australia/QLD

ID926/2020

Net

herla

nds/

NA

276

/202

0

Engl

and/

SHEF

-C00

EE/2

020

Iceland/376/2020

IHU

CO

VID

-014

7

Brazil/S

PB

R-04/2020

Scotla

nd/EDB002/2020

USA/W

A-UW

116/2020

Belgium

/RJ-030552/2020

Eng

land

/CA

MB

-738

E2/

2020

IHUCOVID-0158


Iceland/369/2020

IHUCO

VID-0

303

Australia/V

IC69/2020

USA/M

N-U

W334/2020

Engla

nd/C

AMB-7

4614

/202

0

Bel

gium

/ULG

-300

0/20

20

Franc

e/ARA09

588/

2020

US

A/W

A-S

19/2020


Hangzhou/H

Z185/2020

Icel

and/

500/

2020

Icel

and/

240/

2020

USA/NY 1922/2020

Icel

and/

497/

2020

Germ

any/N

RW-0

6/20

20W

ales/P

HWC-2

6FE9/

2020

US

A/W

A-S

84/2020

USA/WA-UW-1890/2020

US

A/C

T-UW

258/2020


USA/WA-U

W-2

213/

2020

US

A/W

A-U

W221/2020

USA/WI-31/2020


Iceland/259/2020

USA/MN66-MDH66/2020

Iceland/461/2020

IHU

CO

VID

-007

9



Icel

and/

430/

2020

Canada/BC

3972884/2020

Icel

and/

496/

2020

Engl

and/

2013

9008

704/

2020



USA/NY-NYUMC52/2020

Eng

land

/NO

TT-

10E

2CF

/202

0

SouthAfrica/KRISP 0007/2020


Por

tuga

l/PT

0096

/202

0

Net

herla

nds/

Noo

tdor

p 13

6422

2/20

20

Japan/O

S-20-07-1/2020


Wales/PHWC-26BD6/2020


Portugal/P

T0007/2020

USA/WI-GMF-00498/2020

Pakistan/G

ilgit1/2020

USA/IA 6396/2020

USA/SC 3570/2020

US

A/N

Y-N

YU

MC

140/

2020

Shanghai/S

H0051/2020

Bel

gium

/VH

V-0

3241

18/2

020

Iceland/84/2020

IHUCO

VID-0

208

IHUCO

VID-0

446


Eng

land

/201

3209

8804

/202

0

Engl

and/

2013

4067

904/

2020

USA/WI-118/2020

Iceland/361/2020

USA/NY-PV09069/2020

IHUCOVID-0179

Pingxiang/JX151/2020

Singap

ore/

25/2

020

US

A/C

ZB

-RR

057-

007/

2020



USA/WA-UW-1440/2020

USA/W

A-UW

142/2020

Netherlands/N

oordBrabant 55/2020


Eng

land

/201

3407

8804

/202

0

Iceland/462/2020

Iceland/64/2020

Fran

ce/A

RA

1275

9/20

20

Belgium/FFJ-0324124/2020

Scotland/C

VR

215/2020

Scotland/CVR36/2020



Por

tuga

l/PT

0017

/202

0




Sco

tland

/CV

R16

8/20

20


Eng

land

/201

4406

0204

/202

0

Japan/Hu D

P K

ng 19-020/2020

Shang

rao/

JX19

74/2

020

Netherlands/Lim

burg 8/2020

England/20132003304/2020

Fran

ce/B

2340

/202

0

Gre

ece/

10/2

020


England/CAMB-7381F/2020

US

A/N

Y-N

YU

MC

71/2

020

Belgium/VLM-03011/2020


US

A/W

A-S

5/2020


Icel

and/

324/

2020

Eng

land

/201

3400

7104

/202

0

Scotland/CVR23/2020

England/20148006604/2020

Scotland/CVR24/2020

Canada/O

N P

HL8458/2020

IHUCOVID-0019

Sw

itzerland/AG

7120/2020

Belgium/SH-03014/2020

IHU

CO

VID

-019

0

US

A/N

Y-NY

UM

C100/2020

USA/WI-GMF-00441/2020

USA/WA-UW-1758/2020


USA/NY-PV09068/2020


Aus

tral

ia/N

SW

26/2

020


Sco

tland

/ED

B53

0/20

20


IHUCOVID-0320

Eng

land

/201

3007

3304

/202

0

US

A/F

L 6318/2020

Engla

nd/2

0136

0188

/202

0

US

A/M

O 4470/2020

USA/ID-UW-2174/2020

Iceland/184/2020

USA/WA-UW-1907/2020

Scotland/CVR85/2020

England/S

HE

F-C

01CD

/2020

Netherlands/Z

uidH

olland 25/2020

England/S

HE

F-C

0646/2020

Bel

gium

/BJ-

0325

181/

2020


IHUCOVID

-020

1

Eng

land

/201

3006

4304

/202

0

Por

tuga

l/PT

0065

/202

0

US

A/W

A-S

122/2020

Icel

and/

273/

2020

Belgium/VDLH-030548/2020

Icel

and/

441/

2020

Wales/PHW

C-2566D/2020

Engl

and/

CAM

B-74

4A1/

2020

Belgium/MCW-030574/2020


Spa

in/A

ndal

ucia

2012

72/2

020

Bel

gium

/HA

A-0

3095

3/20

20

Wales/PHW

C-255E8/2020


USA/ID-U

W-4

400/

2020

Hon

gKon

g/H

KP

U54

-030

2/20

20


IHU

CO

VID

-027

4

Portugal/P

T0028/2020

Spain/V

alencia81/2020

IHUCOVID

-0412

Senegal/094/2020

Iceland/7/2020

Wales/PHWC-26CB5/2020

Aus

tral

ia/N

SW

07/2

020

USA/WA-UW-1287/2020

Aus

tral

ia/V

IC30

9/20

20

US

A/W

A-U

W319/2020


Brazil/S

PB

R-12/2020

Eng

land

/201

3602

49/2

020

Iceland/62/2020


Iceland/284/2020

Iceland/374/2020

Belgium/CK-0324150/2020

Iceland/164/2020

Iceland/205/2020


England/201360200/2020

Wal

es/P

HW

C-2

65E7

/202

0

Engla

nd/S

HEF-C02

F7/2

020

Belgium

/MS

E-0324121/2020

Englan

d/20

1320

5460

4/20

20

France/N2223/2020

IHUCOVID-0369

Sin

gapo

re/2

7/20

20


Canada/O

N-V

IDO

-01/2020

USA/W

A-UW

-1538/2020

Hangzhou/Z

JU-06/2020

Engla

nd/S

HEF-

C04

97/2

020

USA/WA-UW356/2020

USA/WI-GMF-00986/2020

Wales/P

HW

C-263D

E/2020

Italy/INMI9/2020

US

A/M

A1/2020

Eng

land

/201

3005

2204

/202

0

Mexico/Puebla-InDRE 05/2020

Franc

e/Ly

on 5

08/2

020


Iceland/155/2020

Sin

gapo

re/1

0/20

20

Scotland/E

DB

148/2020

England/20142002704/2020

France/Clermont-Ferrand 651/2020

US

A/W

A-S

16/2020

US

A/V

A-D

CLS

-000

8/20

20

Senegal/026/2020

Iceland/138/2020

US

A/N

Y-PV

09182/2020

Icel

and/

537/

2020

USA/AK-PHL003/2020

Eng

land

/201

3401

7404

/202

0

USA/W

A-UW

228/2020

England/S

HE

F-C0619/2020

USA/W

A-UW

80/2020

Netherlands/N

oordBrabant 3/2020

Japa

n/P

4-5/

2020

US

A/W

A-U

W13

6/20

20


Por

tuga

l/PT0

040/

2020


USA/W

A-UW

72/2020

Bel

gium

/ULG

-100

13/2

020

England/20144071004/2020

Belgium

/ULG

-6950/2020

USA/WA-UW-1741/2020

England/201360298/2020

Australia/V

IC177/2020

DR

C/523/2020

Shanghai/S

H0094/2020

Italy/UniMI03/2020

France

/B2348/2020



USA/WA-UW252/2020

Denmark/SSI-02/2020

US

A/VA

-DC

LS-0002/2020

Netherlands/N

A 51/2020


IHU

CO

VID

-014

6

England/20136094704/2020



USA/NY-PV09037/2020

Wales/PHW

C-25973/2020

US

A/W

A-S

71/2020

Guangdong/2020XN4273-P0036/2020

Wale

s/PHW

C-26D

B2/20

20

Por

tuga

l/PT

0074

/202

0

Kuw

ait/K

U12

/202

0

Aus

tral

ia/V

IC24

9/20

20


USA/NY-PV08415/2020

US

A/W

A-U

W370/2020

Iceland/141/2020

Eng

land

/201

2900

9304

/202

0


USA/CA-C

DPH-UC3/2020

Iceland/488/2020

US

A/W

A-S

100/2020

Latvia/0001/2020

France/IDF3165/2020

Wales/P

HWC-2

4288/2020

Eng

land

/CA

MB

-738

B5/

2020

US

A/W

A-U

W77/2020

Sw

itzerland/VD

0503/2020

Iceland/387/2020

Portugal/P

T0037/2020

Sco

tland

/CV

R89

/202

0


USA/WA-UW-1770/2020





Belgium/UNJP-0324125/2020

USA/NY-PV09019/2020

USA/W

A-UW

261/2020

USA/NY-P

V08496/2020


USA/W

A-UW

148/2020

USA/NY-PV09140/2020


US

A/O

R-U

W-1851/2020

USA/NY-NYUMC11/2020

Engl

and/

2014

4014

504/

2020


Eng

land

/201

1804

4606

/202

0

US

A/W

A-U

W125/2020

IHUCOVID-0496

IHUCOVID-0483

Hangzhou/HZ551/2020

US

A/W

A-U

W-2249/2020

Scotla

nd/E

DB40

9/20

20


Belgium/LT-030956/2020

US

A/W

A-U

W-1603/2020

Guangdong/G

D2020246-P

0028/2020

Norw

ay/2090/2020

Finland/13M77/2020


USA/NY-N

YUMC12/2020

Net

herla

nds/

NA

145/

2020

Switzerland/TI2045/2020

IHUCO

VID-0

248


Por

tug

al/P

T00

53/2

020


Belgium/AC-030982/2020

USA/WA-UW192/2020

Iceland/5/2020

Finl

and/

14M

20/2

020

England/20129158804/2020


IHU

CO

VID

-018

0

Eng

land

/201

1211

2106

/202

0

USA/WA-UW307/2020

Belgium/ULG-1

0032/2020

France/H

F3290/2020

Finland/13M19/2020


Sw

itzerland/1000477806/2020

Wales/PHW

C-24121/2020

Sw

itzerland/GR

3043/2020


England/NOTT-10E001/2020

Spain/Canarias201495/2020


Australia/V

IC139/2020

Net

herla

nds/

NA

235/

2020

US

A/N

Y-N

YU

MC

99/2

020



Iceland/69/2020



Sco

tland

/ED

B53

3/20

20

England/20146004804/2020


Algeria/G

0638 2264/2020

USA/NY-NYUMC73/2020

Switzerland/BE2536/2020

USA/WI-53/2020



US

A/W

A-U

W236/2020

US

A/W

A-S

46/2020

USA/W

A-UW

87/2020

USA/W

I-60/2020


Sco

tland

/CV

R17

/202

0


England/20142012304/2020

Spain/Andalucia201617/2020

England/S

HE

F-C

0725/2020

Hangz

hou/

HZCDC0001

/202

0



USA/WI-89/2020


HongK

ong/VB

20026565/2020

Wales/PH

WC

-270C7/2020


Singa

pore/3/2020

USA/W

A-S97/2020

US

A/N

Y-N

YU

MC

54/2

020

Wal

es/P

HW

C-2

5AF8

/202

0

Belgium

/ULG

-10061/2020

Australia/Q

LDID

922/2020

Wales/P

HW

C-2523C

/2020

Sco

tland

/CV

R26

4/20

20

Canada/ON P

HL6980/2020

Wales/PHW

C-26E55/2020


Aus

tralia

/WA

07/2

020

US

A/W

A-S

51/2020

Sco

tland

/ED

B06

8/20

20

IHUCOVID-0389

Wales/PHW

C-269CD/2020

IHUCOVID-0282

Finland/14M77/2020

USA/WA-UW-1853/2020

US

A/W

A-U

W212/2020

Iceland/509/2020

USA/W

A-S89/2020

USA/WI-1

3/2020


US

A/O

R 265

6/2020

US

A/W

A-U

W376/2020

France/H

F2039/2020



Can

ada/

ON

PH

L692

2/20

20

USA/WI-28/2020


IHUCOVID-0368

Nanchang/JX149/2020


Aus

tral

ia/N

SW

05/2

020

Wuh

an/W

HU02/2

020

USA/NH 0004/2020

Engla

nd/2

0132

0919

04/2

020

IHU

CO

VID-0

116


Wales/PHW

C-246AA/2020

Luxe

mbo

urg/

LNS

0641

910/

2020

Net

herla

nds/

NA

120/

2020


Scotland/CVR58/2020

Japan/TK

-20-31-3/2020

Eng

land

/201

3904

8804

/202

0


US

A/W

I-86/

2020

Bel

gium

/DH

WM

-030

41/2

020

USA/WI-3

0/2020


Japa

n/P

4-6

/202

0

Wales/PHWC-255AC/2020

Hon

gKon

g/H

KP

U28

300

1/20

20

Engl

and/

2013

4078

404/

2020

Netherlands/N

A 164/2020

Wales/P

HW

C-259A

0/2020

USA

/MA

3623

/202

0

Canada/ON PHLU8150/2020

US

A/M

N131-M

DH

131/2020

USA/NY-PV09029/2020

Net

herla

nds/

Zui

dHol

land

33/

2020

USA

/NC

003

1/20

20

France/IDF2412/2020

USA/WA-UW-1346/2020

Iceland/558/2020

Australia/V

IC228/2020

IHUCOVID-0136

Belgium

/VF

-030562/2020


Aust

ralia

/VIC

99/2

020


Australia

/VIC

227/2020

Wales/PHW

C-269DC/2020

IHUCOVID-0031

Eng

land

/201

3007

3404

/202

0

USA/WA-UW-2149/2020

Indi

a/16

44/2

020

Engla

nd/2

0132

0544

04/2

020

USA/NY-PV09129/2020

Jian/JX129/2020

Engla

nd/2

0132

0664

04/2

020

IHUCO

VID-0

336

USA/WI-37/2020

Eng

land

/201

3610

22/2

020


Wuhan/W

H04/2020



Finl

and/

13M

82/2

020

Iceland/220/2020

Eng

land

/201

3005

1704

/202

0

Australia/V

IC197/2020

Engl

and/

2013

6020

1/20

20


Can

ada

/BC

528

2984

/202

0

Eng

land

/201

3802

21/2

020

India/773/2020

Eng

land

/201

3202

0204

/202

0

Austra

lia/V

IC39

/202

0

IHUCOVID

-027

1

England/20132108704/2020

Canada/B

C 8622445/2020


USA

/WI-1

7/20

20



2/2020

Eng

land

/201

3110

4704

/202

0

Scotland/E

DB

513/2020

Wal

es/P

HWC-2

581C

/202

0

Eng

land

/201

3902

9104

/202

0

USA/IL 1293/2020

Eng

land

/NO

TT-1

0E14

A/2

020


Netherlands/U

trecht 15/2020


IHUCOVID

-045

6

England/20108003302/2020

Italy

/TE

5476

/202

0

Iceland/310/2020

Scotland/CVR62/2020

Iceland/56/2020


Bel

gium

/ULG

-100

11/2

020

IHUCOVID-0063

Engl

and/

2013

8002

8/20

20


Sco

tland

/CV

R17

1/20

20

Aus

tral

ia/V

IC66

/202

0

Nor

way

/145

5/20

20

Engl

and/

2013

2081

804/

2020

Net

herla

nds/

Noo

rdB

raba

nt 2

5/20

20

Kor

ea/K

CD

C20

01/2

020

Icel

and/

239/

2020

France/IDF2536/2020


Eng

land

/201

3402

1904

/202

0


USA/IL1/2020

Eng

land

/201

3210

3604

/202

0

DRC/KN-0054/2020

USA/IL-UW351/2020

Eng

land

/CA

MB

-738

D3/

2020

Slova

kia/

SK-BM

C5/20

20

US

A/W

A-S

38/2020

Iceland/399/2020


Netherlands/ZuidH

olland 102/2020

Engla

nd/2

0132

0357

04/2

020


Scotland/CVR54/2020

USA/NY-P

V0914

5/20

20

England/20142083804/2020

Canada/BC 41851/2020

Wal

es/P

HW

C-2

4BC

9/20

20

Italy/VR 20COV 21-37/2020

DRC/300/2020

Net

herla

nds/

Noo

rdB

raba

nt 6

/202

0

Taiw

an/C

GM

H-C

GU

-09/

2020

Belgium/UG

ent-22/2020

Shang

hai/S

H0066

/202

0

USA/WA-UW-1640/2020


IHUCOVID

-038

6

France/ID

F2075/2020

Iceland/232/2020

Icel

and/

229/

2020

Netherlands/NoordHolland 4/2020

Icel

and/

162/

2020

England/20144076204/2020

USA/UT-0

0022/2020



Scotla

nd/E

DB097/

2020



USA/NY2-PV08100/2020

Germany/N

RW-1

7/2020

Aus

tralia

/VIC

194/

2020


Thaila

nd/Bang

kok-CO

NI-0

017/2020

Iceland/211/2020

USA/GA 2118/2020

Sw

itzerland/GE

0199/2020

Aus

tralia

/VIC

152/

2020

Por

tuga

l/PT

0085

/202

0

Can

ada/

BC

407

8583

/202

0


Net

herla

nds/

NA

43/

2020

Ger

man

y/N

RW

-41/

2020


Iceland/1/2020

Iceland/104/2020

USA/NY-PV09128/2020

USA/WI-87/2020

DRC/396/2020

England/SHEF-BFD54/2020

Belgium/ULG-6

743/2020


Hangzhou/H

ZCDC0119/2020

Iceland/222/2020

US

A/W

I-84/

2020

Engl

and/

2013

2001

904/

2020

Eng

land

/201

3202

4004

/202

0

Belgium

/JL-03044/2020

Icel

and/

586/

2020

Iceland/198/2020

England/20139027304/2020


Taiw

an/C

GM

H-C

GU

-04/

2020

USA/WI-G

MF-00281/2020

Germany/NRW-42.3/2020

IHUCO

VID-0

172

Aust

ralia

/VIC

187/

2020

Shanghai/S

H0088/2020

Eng

land

/SH

EF

-C05

A3/

2020

Iceland/466/2020

USA/WI-85/2020

Iceland/173/2020

IHU

CO

VID

-050

2

France/HF2196/2020

Iceland/353/2020


Belgium

/QK

J-03015/2020

USA/N

Y-N

YUM

C32

/202

0

USA/WI-107/2020



Netherlands/Flevoland 1/2020

Japan/KY

-V-029/2020


US

A/N

Y-P

V09

152/

2020

Iceland/562/2020

USA/WA-UW-1715/2020


Portugal/P

T0009/2020

USA/NY-PV09041/2020

US

A/W

A-U

W68/2020

Cambodia/0012/2020

USA/MN36-M

DH36/2020

Iceland/560/2020

Net

herla

nds/

NA

46/

2020

Iceland/78/2020

England/NOTT-10EB6A/2020

Senegal/082/2020

USA/WA-UW-1774/2020


Wales/PHW

C-2675A/2020

Scotland/CVR

74/2020

Belgium/LY-030575/2020


Eng

land

/201

3209

1804

/202

0




IHUCOVID-0039

Sco

tland

/CV

R19

5/20

20

Aus

tralia

/VIC

299/

2020

Australia/Q

LDID

921/2020

USA/WI-32/2020

Australia/V

IC317/2020

US

A/V

A-D

CLS

-004

2/20

20

IHUCO

VID-0

247

Net

herla

nds/

Utr

echt

18/

2020

1

2

3

3b3a

4

Tree scale: 0.00001

Colored ranges

Africa

South America

Oceania

Europe

Asia

North America


https://doi.org/10.1101/2020.09.04.282616

Nb S

Nb NS

Deletion event

SARS-CoV-2 genome 126 178 6

nsp1 3 5 2

nsp2 11 17 1

nsp3 17 27 0

nsp4 6 7 0

nsp5 9 5 0

nsp6 1 5 0

nsp7 0 1 0

nsp8 3 5 0

nsp9 1 3 0

nsp10 2 0 0

nsp12a 0 1 0

nsp12b 12 11 0

nsp13 5 8 0

nsp14 9 7 0

nsp15 6 5 0

nsp16 4 2 0

S (spike protein) 12 18 1

ORF3a 4 13 1

E (envelope) 1 1 0

M (membrane) 4 4 0

ORF6 0 3 0

ORF7a 2 5 1

ORF7b 0 1 0

ORF8 3 4 0

N (nucleocapsid) 11 20 0

S: synonymous mutation; NS : nonsynonymous mutation

Table 1. Mutational events detected in the 309 SARS-CoV-2 genomes as compared to the reference Wuhan coronavirus.


https://doi.org/10.1101/2020.09.04.282616

Position

genome

Region/

gene

Mutation

nt

Mutation

aa

Nb

isolates

204 5UTR G>T 1

241 5UTR C>T 297

313 nsp1 CTC>CTT L 20

353 nsp1 GGC>AGC G30S 1

510 nsp1 GGT>X 1

511 nsp1 GGT>X 1

512 nsp1 CAT>X 1

513 nsp1 CAT>X 1

514 nsp1 CAT>X 1

515 nsp1 GTT>X 1

516 nsp1 GTT>X 1

517 nsp1 GTT>X 1

518 nsp1 ATG>X 1

537 nsp1 GAA>GGA E91G 1


625 nsp1 AAG>AAT K120N 1

639 nsp1 AAG>AGG K125R 1

683 nsp1 CTA>TTA L 2

686 nsp1 AAG>X 3

687 nsp1 AAG>X 3

688 nsp1 AAG>X 3

689 nsp1 TCA>X 3

690 nsp1 TCA>X 3

691 nsp1 TCA>X 3

692 nsp1 TTT>X 3

693 nsp1 TTT>X 3

694 nsp1 TTT>X 3

710 nsp1 CTT>ATT L149I 1

834 nsp2 TTC>TGC F10C 1

958 nsp2 TGC>TGT C 1

1059 nsp2 ACC>ATC T85I 68

1168 nsp2 CGA>CGG R 1

1198 nsp2 GAA>GAG E 6

1361 nsp2 GTT>TTT V186F 1

1427 nsp2 CAT>TAT H208Y 1

1440 nsp2 GGC>GAC G212D 2

1463 nsp2 GGT>AGT G220S 1

1493 nsp2 GTG>ATG V230M 1

1515 nsp2 CAT>CGT H237R 1

1605 nsp2 AAT>X 2

1606 nsp2 AAT>X 2

1607 nsp2 GAC>X 2

1736 nsp2 GTG>ATG V311M 1

1942 nsp2 GTG>GTT V 1

1959 nsp2 GCC>GTC A385V 1

2037 nsp2 GCT>GTT A411V 1

2062 nsp2 GCC>GCT A 2

2065 nsp2 TAC>TAT Y 1

2086 nsp2 CAG>CAT Q427H 1

2150 nsp2 GAT>AAT D449N 2

2189 nsp2 CTT>TTT L462F 4



2480 nsp2 ATT>GTT I559V 1

2537 nsp2 TTG>CTG L 1

2558 nsp2 CCA>TCA P585S 1

2578 nsp2 AGT>AGC S 3

2602 nsp2 GTT>GTG V 1

2676 nsp2 CCT>CTT P624L 2

2891 nsp3 GCA>ACA A58T 2

3037 nsp3 TTC>TTT F 303

3049 nsp3 GAT>GAC D 1

3094 nsp3 CCA>CCG P 1


3338 nsp3 GTG>TTG V207L 6

3429 nsp3 ACA>ATA T237I 1

3478 nsp3 GTT>GTC V 1

3619 nsp3 GTC>GTT V 2






4459 nsp3 ATA>ATC I 2


4655 nsp3 CGG>TGG R646W 1

4761 nsp3 GAA>GGA E681G 1

4800 nsp3 AAA>AGA K694R 1

4886 nsp3 CCT>TCT P723S 1

5336 nsp3 AAT>GAT N873D 1

5338 nsp3 AAT>AAC N 2


5570 nsp3 ATG>GTG M951V 1






6286 nsp3 ACC>ACT T 1


(which w

as not certified by peer review) is the author/funder. A

ll rights reserved. No reuse allow

ed without perm

ission. T

he copyright holder for this preprintthis version posted S

eptember 4, 2020.

; https://doi.org/10.1101/2020.09.04.282616

doi: bioR

xiv preprint

https://doi.org/10.1101/2020.09.04.282616

6531 nsp3 GAG>GTG E1271V 1

6539 nsp3 CAC>TAC H1274Y 1


6636 nsp3 ACT>ATT T1306I 1


7162 nsp3 GAC>GAT D 2

7393 nsp3 CCG>CCT P 1

7424 nsp3 TTT>GTT F1569V 1


7728 nsp3 TCT>TTT S1670F 1

8179 nsp3 CGG>CGT R 1

8371 nsp3 CAG>CAT Q1884H 100


8676 nsp4 AGT>ATT S41I 1

8782 nsp4 AGC>AGT S 2

8858 nsp4 GTC>ATC V102I 1


9166 nsp4 ACC>ACT T 1

9223 nsp4 CAC>CAT H 1


9286 nsp4 AAC>AAT N 5


9652 nsp4 ATG>ATT M366I 2



9996 nsp4 TCA>TTA S481L 1

10097 nsp5 GGT>AGT G15S 1


10252 nsp5 TTC>TTT F 1

10262 nsp5 GCT>TCT A70S 1

10277 nsp5 CTC>TTC L75F 1


10323 nsp5 AAG>AGG K90R 2

10369 nsp5 CGC>CGT R 1



10626 nsp5 GCA>GTA A191V 2

10642 nsp5 ACG>ACT T 1



11083 nsp6 TTG>TTT L37F 19

11117 nsp6 ATT>GTT I49V 1





11858 nsp7 GTA>ATA V6I 1

12318 nsp8 TCT>TAT S76Y 1

12334 nsp8 GCA>GCT A 2

12439 nsp8 CCC>CCT P 1

12496 nsp8 TAT>TAC Y 1

12513 nsp8 ACG>ATG T141M 3







13006 nsp9 GCT>GCC A 7


13132 nsp10 CAA>CAG Q 1

13458 nsp12a TCG>TTG S6L 11

13693 nsp12b ACA>TCA T76S 1

14228 nsp12b AAG>AGG K254R 2

14358 nsp12b TGC>TGT C 1

14391 nsp12b TTC>TTT F 5

14408 nsp12b CCT>CTT P314L 303

14599 nsp12b CTA>TTA L 1

14724 nsp12b TTC>TTT F 3

14805 nsp12b TAC>TAT Y 2

14857 nsp12b GTT>TTT V464F 1

15021 nsp12b CTT>CTC L 1

15120 nsp12b GTC>GTT V 1

15277 nsp12b CAC>TAC H604Y 1

15324 nsp12b AAC>AAT N 39

15327 nsp12b ATG>ATT M620I 1

15368 nsp12b ACA>ATA T634I 1

15438 nsp12b ATG>ATT M657I 6


15779 nsp12b AAG>AGG K771R 1


16020 nsp12b GTG>GTA V 1

16045 nsp12b CTT>TTT L860F 1

16089 nsp12b TTG>TTA L 1

16226 nsp12b ACA>ATA T920I 1


16377 nsp13 CCG>CCT P 1


16394 nsp13 CCA>CTA P53L 3

16816 nsp13 CAA>AAA Q194K 1

16968 nsp13 GAG>GAT E244D 2

16992 nsp13 TTA>TTG L 1

17104 nsp13 CAT>TAT H290Y 1

(which w



ed without perm

ission. T


eptember 4, 2020.

; https://doi.org/10.1101/2020.09.04.282616

doi: bioR

xiv preprint

https://doi.org/10.1101/2020.09.04.282616

17125 nsp13 CTC>TTC L297F 2

17247 nsp13 CGT>CGC R 1



17964 nsp13 ATG>ATA M576I 1

18175 nsp14 CCT>TCT P46S 3

18186 nsp14 ATG>ATT M49I 1

18259 nsp14 ATC>GTC I74V 1


18366 nsp14 CTA>CTG L 3


18495 nsp14 CTT>CTC L 1


18904 nsp14 CGT>TGT R289C 1


19167 nsp14 AAA>AAG K 1




19518 nsp14 TTG>TTT L493F 3





20125 nsp15 GGA>CGA G169R 1


20268 nsp15 TTA>TTG L 3

20294 nsp15 TAT>TGT Y225C 4

20310 nsp15 TAT>TAC Y 2

20382 nsp15 CTA>CTT L 1

20401 nsp15 TCA>CCA S261P 1

20628 nsp15 GGC>GGT G 1

20808 nsp16 CTG>CTA L 1


21015 nsp16 CAT>CAC H 2

21225 nsp16 TGG>TGT W189C 1


21369 nsp16 ATT>ATC I 1

21587 S CCA>TCA P9S 1

21597 S TCT>TTT S12F 6

21618 S ACA>ATA T19I 1

21622 S ACC>ACT T 1

21727 S TTC>TTT F 1

21765 S ATA>X 1

21766 S ATA>X 1

21767 S CAT>X 1

21768 S CAT>X 1

21769 S CAT>X 1

21770 S GTC>X 1

21778 S GGG>GGA G 1

21846 S ACT>ATT T95I 1

21938 S GTT>TTT V126F 1

22346 S GCT>TCT A262S 1

22458 S ACA>ATA T299I 2

22480 S TTC>TTT F 1

22550 S CCT>TCT P330S 1

22606 S GCA>GCT A 2

23042 S TCA>GCA S494A 1

23191 S TTC>TTT F 1

23248 S TTC>TTT F 3

23403 S GAT>GGT D614G 303

23415 S ACA>ATA T618I 1

23575 S TGC>TGT C 1

24050 S GAT>CAT D830H 1

24079 S GAT>GAC D 2

24095 S GCT>TCT A845S 2

24193 S CTG>CTA L 1

24771 S GCA>GTA A1070V 1

24794 S GCT>TCT A1078S 2

24867 S TGG>TTG W1102L 1

24904 S ATC>ATT I 1

24998 S GAC>TAC D1146Y 5

25012 S GAG>GAA E 1

25047 S CCA>CTA P1162L 1

25433 ORF3a ACT>ATT T14I 2

25440 ORF3a AAG>AAT K16N 2

25521 ORF3a TTC>TTT F 1

25563 ORF3a CAG>CAT Q57H 211

25606 ORF3a GCA>TCA A72S 1

25667 ORF3a TCA>TTA S92L 1

25688 ORF3a GCT>GTT A99V 8

25699 ORF3a GCC>TCC A103S 1

25710 ORF3a CTC>CTT L 1

25720 ORF3a GCT>TCT A110S 1

25731 ORF3a TAC>TAT Y 6

25776 ORF3a TGG>TGT W128C 1

25782 ORF3a TGC>TGT C 1

25825 ORF3a TAT>CAT Y145H 1

25909 ORF3a GAT>TAT D173Y 3

25933 ORF3a GAA>AAA E181K 1

26144 ORF3a GGT>GTT G251V 2

26161 ORF3a AAT>X 1

26256 E TTC>TTT F 1

(which w



ed without perm

ission. T


eptember 4, 2020.

; https://doi.org/10.1101/2020.09.04.282616

doi: bioR

xiv preprint

https://doi.org/10.1101/2020.09.04.282616

26314 E GTG>TTG V24L 1

26622 M CTT>TTT L34F 2

26718 M GTG>TTG V66L 4

26735 M TAC>TAT Y 1

26773 M ATG>ACG M84T 1

26885 M AAC>AAT N 1

26907 M CTG>ATG L129M 7

27002 M GAC>GAT D 1

27143 M AAC>AAT N 2

27208 ORF6 CAT>TAT H3Y 2

27256 ORF6 ATG>TTG M19L 1

27261 ORF6 AGG>AGT R20S 1

27403 ORF7a ATT>GTT I4V 1

27434 ORF7a ACT>X 1

27435 ORF7a ACT>X 1

27436 ORF7a TGT>X 1

27476 ORF7a ACA>ATA T28I 1

27494 ORF7a CCT>CTT P34L 2

27505 ORF7a GGA>TGA G38* 1

27641 ORF7a TCA>TTA S83L 1

27659 ORF7a AGA>ATA R89I 1

27684 ORF7a TAC>TAT Y 1

27741 ORF7a CTC>CTA L 2

27874 ORF7b ACT>ATT T40I 1

27942 ORF8 CAC>TAC H17Y 1

27996 ORF8 GAC>TAC D35Y 6

28027 ORF8 TGG>TTG W45L 3

28037 ORF8 AGA>AGG R 1

28144 ORF8 TTA>TCA L84S 2

28202 ORF8 TCG>TCA S 1

28253 ORF8 TTC>TTT F 2

28291 N CCC>CCT P 1

28311 N CCC>CTC P13L 1

28312 N CCC>CCT P 4

28326 N GGT>GTT G18V 1

28374 N GGG>GAG G34E 1

28519 N GAC>GAT D 1

28541 N GCT>ACT A90T 1

28628 N GCT>TCT A119S 2

28651 N AAC>AAT N 2

28682 N GGA>AGA G137R 2

28690 N TTG>TTT L139F 2

28775 N CCA>TCA P168S 1

28845 N CGC>CTC R191L 3

28851 N AGT>AAT S193N 2

28851 N AGT>ATT S193I 1

28878 N AGT>AAT S202N 2

28881 N AGG>AAA R203K 44

28882 N AGG>AAA R203K 44

28883 N GGA>CGA G204R 44

28886 N ACT>GCT T205A 1

28957 N AAC>AAT N 1

29167 N TAC>TAT Y 3

29171 N CAT>TAT H300Y 1

29179 N CCG>CCT P 1

29206 N GCT>GCA A 1

29218 N TTC>TTT F 1

29272 N TAC>TAT Y 1

29281 N GCC>GCT A 1

29363 N CCA>TCA P364S 2

29477 N GAT>TAT D402Y 1

29527 N CAG>CAT Q418H 1

29536 T>C 1

29555 C>T 1

29742 3UTR G>A 1

29744 3UTR G>T 1

29745 3UTR G>T 1

29747 3UTR G>T 1

29754 3UTR C>T 1

29779 3UTR G>C 3

Table 2. Detailed mutations detected in the 309 SARS-CoV-2 genomes from French patients

(which w



ed without perm

ission. T


eptember 4, 2020.

; https://doi.org/10.1101/2020.09.04.282616

doi: bioR

xiv preprint

https://doi.org/10.1101/2020.09.04.282616

genomic diversity and evolution of coronavirus (sars-cov-2 ... · 04/09/2020 · ten were in...

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