forest trees of maine

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CENTENNIAL EDITION

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The book contains information on 78 different tree species, including all of Maine's commercially important native tree species, as well as a few or the more common and important introduced trees.

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Page 1: Forest Trees of Maine

C E N T E N N I A L E D I T I O N

Page 2: Forest Trees of Maine

T M F S The Maine Forest Service was established in 1891 to ensure Maine’scitizens the greatest economic and social benefits from trees andforestlands of the State.

Part of the Department of Conservation since 1973, the primary responsibilitiesof the Maine Forest Service include:

to develop through information, education, and formal publicationsa greater public awareness and appreciation of forests as Maine’sbasic economy and renewable resource;

to provide advice and assistance in forest management towoodland owners;

to maintain and improve the scenic beauty, wildlife habitat,and recreational values of Maine;

to encourage and promote appropriate forest land managementpractices; and...

to protect the forest resource from fire, insects, diseases, andother natural enemies.

DThe Centennial Edition of the Forest Trees of Maine is dedicated to all the hardworking men and women who make their living in the Maine woods.

Forest Trees of MaineFirst published in 1908 Revised 2008

(Fourteenth Edition)Maine Forest Service

Department of ConservationMaine Forest Service Phone: (207) 287-2791

E-mail: [email protected]

Printed in Maine on certified paper.

Page 3: Forest Trees of Maine

T C Introduction............................................................................................ 2Forward by David Field .......................................................................... 3History ................................................................................................... 4A Few Precautions.................................................................................. 6Winter Key........................................................................................... 10Summer Key......................................................................................... 12Glossary................................................................................................ 14Tree Parts and Functions ...................................................................... 20Autumn Coloration .............................................................................. 22Forest Trees of Maine

Conifers ............................................................................................ 24Broadleaves....................................................................................... 60

Selected References ............................................................................ 174Index................................................................................................... 175

A A great many people contributed to the production of this publication. The Forest Trees of MaineCentennial Committee was instrumental in bringing this book into being. Members of the committeeincluded Peter Lammert, Dan Jacobs, Merle Ring, Judy Tyler, Kevin Doran, Greg Miller, Jan AmesSanterre, Andy Shultz and Keith Kanoti. Special credit is due to Peter Lammert for updating thewood uses, Dan Jacobs for researching and writing the History section, and Greg Miller for produc-ing the range maps. Others who contributed to the project include: Tom Whitworth, Scott Sawtelle,Tom Collins, Jen Wright, John Anastasio and Debbie Jacques.

Keith Kanoti revised the manuscript and acted as project manager.

The book was designed by David Deal and the manuscript was edited by Donna Stuart, both ofGlen Group Inc.

Tom Rawinski, botanist with the US Forest Service Northern Research Station, reviewed themanuscript for technical accuracy.

Cover photo by Arthur Rogers, presented to Forest Commissioner Austin Wilkins by GovernorPercival Baxter.

The glossary sketches were used with the generous permission of Dr. Michael A. Dirr.

The range maps are based on Atlas of United States Trees by the late Dr. Elbert L. Little.

The majority of the photographs were taken by the Maine Forest Service, Policy and ManagementDivision field staff. Through bugwood.org, several photographers generously let us use images that wecould not obtain ourselves. These include Paul Wray, Iowa State University (white oak acorns, swampwhite oak acorns, northern red oak acorns, black oak acorns, butternut fruit, shagbark hickory fruit,bitternut hickory fruit, balsam fir cones, black willow leaves); The Dow Gardens Archives, DowGardens (mountain laurel flowers, rhododendron flowers, striped maple fruit); Chris Evans, River toRiver CWMA (bitternut hickory leaves); Bill Cook, Michigan State University Extension (beechnuts,American hornbeam fruit); and David J. Moorhead, University of Georgia (flowering dogwood flowers).

Other photography sources were obtained through USDA-NRCS PLANTS Database. Theseinclude Robert H. Mohlenbrock 1995, Northeast wetland flora: Field office guide to plant species(Atlantic white cedar cones); D.E. Herman, et al. 1996, North Dakota tree handbook (Americanbasswood fruit, blue spruce cones/foliage, bur oak fruit, staghorn sumac fruit); and Steve Hurst(Sugar maple fruit, scarlet oak fruit, black ash fruit).

Page 4: Forest Trees of Maine

2

I In 1908 the Maine Forest Service released a booklet titled Forest Trees of Maine.

In his 1910 Commissioner’s report, Forest Commissioner Edgar Ring wrote of thepopularity of the new publication: “For the Forest Trees of Maine there has been a largeand constant demand which will very soon exhaust the edition. Possibly in order to meetthe demands for this pamphlet it will be considered wise and money well spent to issueanother edition.” Now, 100 years later and in its 14th edition, Forest Trees of Maineremains the Maine Forest Service’s most popular publication.

Since 1908, all editions of Forest Trees of Maine have had the same objective: torelate accurate information and to keep pace with new findings. As those who arefamiliar with Forest Trees of Maine will immediately notice, we have departed from thetraditional format for this edition. This has allowed us to include color photographs,which have long been requested. For those who prefer the tried and true Forest Treesof Maine format, it will still be made available.

For the first time, range maps have been included. The maps are based on thoseof those of the legendary US Forest Service dendrologist, Dr. Elbert Little, whoassisted with the 7th edition. The maps indicate the parts of the state where youare most likely to encounter each tree species. No map is perfect, and it is certainlypossible to find a species outside of its indicated range.

The keys have been revised and, for the first time, a winter key has beenincluded. To help you use the keys, sketches have been added to the glossary whichillustrate many of the terms used.The keys are limited to the trees in the publication.For information on more complete keys, see Selected References on page 174.

The book contains information on 78 different tree species, including all of Maine’scommercially important native tree species, as well as a few of the more common andimportant introduced trees. As with previous editions, no attempt has been made toinclude all the species in complicated groups, such as willows and hawthorns. Whendeciding which species to include in this edition, emphasis was placed on trees that occurin Maine’s forests.With a few exceptions (e.g., horsechestnut, blue spruce, black walnut),species limited to ornamental plantings were excluded. Other introduced species wereincluded if they have been commonly used in forest plantations (e.g., Norway spruce,Scots pine) or have escaped cultivation and are reproducing in forested areas (e.g., blacklocust, Norway maple). Several species are included that occasionally grow large enoughto be considered small trees (e.g., bear oak, witch hazel, rhododendron, mountainlaurel), but are more commonly found as shrubs.

Scientific names in this publication follow the Integrated Taxonomic InformationSystem database: www.itis.gov.

Historic photographs found throughout the book are from the Maine ForestService Archives and the Maine State Museum.

For more information about this publication or the Maine Forest Service,call 207-287-2791, e-mail us at: [email protected] or visit our website at:www.maineforestservice.gov

Page 5: Forest Trees of Maine

3F O R E W O R D

F I am privileged to be able to write a foreword for the centennial edition of The

Forest Trees of Maine, this wonderful gift that the Maine Forest Service has providedfor so long!

Suppose that someone invented a wonderful new machine. It can soak up the“greenhouse” gas carbon dioxide from the air and breathe out oxygen. It can pumphuge amounts of water from the soil to reduce floods, while holding the soil togetherand helping to clean water that flows into streams and ponds. It can produce strongmaterials for building and fiber for paper. Pick up a stick of firewood. You hold thepower of sunlight in your hands! This machine can store solar energy until werelease it through fire. If we take care of the land where it grows, then our machinecan be replaced by new machines just like it that will give us more of these wonderful things. Of course, the “machine” is not new at all—it is the tree—the tree that is beautiful in the forest and that forms beauty in the form of houses,furniture and the pages of a child’s book.

Maine’s forest trees are worth knowing. We don’t have as many species as dosome other states, but we have more acres of land covered with trees. Varioussources estimate that we have nearly 96% as much forest as when Maine was firstsettled. More than in any other state, this great forest is privately owned, yet is moreaccessible to the public than is true in most areas.

Today too many children grow up in a world of television and computer games.They know little about where the food they eat, the clothing they wear, or the paperthey write on comes from. Using this book to learn to identify trees, and to learnabout trees and forests, can help to keep your children physically and mentallyhealthy and can be a great family activity. What fun it can be to explore the woodsand see the diversity of trees! Can you identify one of the earliest to blossom in thespring—the serviceberry? Did you know that one shrub-like tree (witch hazel)does not blossom until the fall? Some—like the poplars (aspens), cherries, andwhite birch—are fast-growing “pioneers” that thrive in the full sun following aforest fire or timber harvest. Others, like sugar maple, can take root in deep shadeand grow slowly for hundreds of years.

I hope that you will take the time to enjoy Maine’s trees, and to use the wealthof information in this book to educate yourselves, your friends, and your childrenabout this marvelous, renewable resource that is essential to Maine’s quality of life.

—Dr. David Field, Professor Emeritus, School of Forest Resources, University of Maine

Page 6: Forest Trees of Maine

4 H I S T O R Y

H S 1908The history of the most recognizable publication by the

Maine Forest Service, Forest Trees of Maine, began in 1908.While Mainers were enjoying the first edition of the Forest

Trees of Maine that year, the nation as a whole was becomingincreasingly hungry for forest resources. In the nation’s capital,conservation issues and court rulings in Maine held the interestof the President.

In his 1908 State of the Union message, PresidentTheodore Roosevelt declared, “Thanks to our own reckless useof our splendid forests, we have already crossed the verge of atimber famine in this country…”

In a speech at the White House that same year, thePresident applauded the State of Maine (particularly the Supreme Court of Maine)for an “exceedingly important judicial decision.” The decision, which confirmed thelegislature’s authority to regulate timber harvesting, was viewed by PresidentRoosevelt as a step towards “the wise utilization of forests....Such a policy willpreserve soil, forests, waterpower as a heritage for the children and the children’s

children of the men and women of thisgeneration.”

In the conservation arena, 1908was an important year: It marked thearrival of this great companion that hasguided many people on journeysthrough the Maine woods.

1908

Region II forest management and operationsclass, Dyer Brook 2008.

Logging crew, Upper Kennebec River, 1908

Page 7: Forest Trees of Maine

5H I S T O R Y

T NA tremendous number of changes have occurred throughout Maine since the

first printing of the Forest Trees of Maine in 1908. These changes, too numerableto count, have impacted all our lives, as well as our forest products industry. Thetable below illustrates some of the changes that have occurred in Maine over thepast century.

T H E N A N D N O W 1 9 0 8 2 0 0 8

Forested Area (%) 751 89

Population of Maine 694,4662 1,274,923

Population of U.S. 92,228,496 281,421,906

Harvest Volume (cords) 2,879,807 6,742,351*

Stumpage Price of Spruce (MBF**) $5.491 $135.00

Forestry Students at UMO 313 50

Students at UMO 884 11,800±

Cost of Bangor Daily (year) $6.00 $180.00±

Most Harvested Hardwood Aspens3 Red Maple

Primary Use of Paper Birch Spools3 Pulp

Gypsy Moth Infested Area of Maine York and Cumberland All Counties

Maine Indian Basketmakers 400 100

Primary Use of Black Ash Indian Baskets

Agricultural & Industrial Collectables & Art

1 Kellogg, R.S. 1909. The Timber Supply of the United States, USDA Forest Service, Circular 166.2 1909. Maine Register, State Year Book and Legislative Manual 1909-1910, No. 40, G.M. Donham, Portland, 1051pp.3 Ring, E.E. 1910. Report of the Forest Commissioner, Maine, Kennebec Journal Print, Augusta, 110 pp.* Harvest volume listed is for 2005. **MBF: Thousand board feet. Note: All values listed in nominal dollars.

1917 1925 1932 1951

Page 8: Forest Trees of Maine

6 A F E W P R E C A U T I O N S

A F PThe Maine woods are a pretty safe place; however, there are a few hazards

anyone who is learning to identify trees should be aware of.

P O I S O N O U S P L A N T S

Maine has two species of poisonous plants you should learn to identify: poisonivy and poison sumac. Severe dermatitis can result when skin comes in contact withroots, stems, leaves, flowers, fruit or with implements or clothing that have comein contact with plant parts of either poison ivy or poison sumac. Smoke resultingfrom the burning of plant parts of either species is also poisonous.

POISON IVY, or mercury, is widely distrib-uted throughout the state. It grows as an aerially-rooted climbing vine on trees or as a smooth,trailing vine or erect shrub along stonewalls,fencerows, roadsides and near water bodies.

The leaves are alternate, compound, with 3very shiny, dark green leaflets. Leaflet margins arelobed, wavy, toothed or entire. The stalk of theterminal leaflet is much longer than those of the

2 lateral leaflets. Fall color is often a fiery red.The fruit is a creamy-white, ribbed, globular, BB-sized drupe that occurs in

axillary clusters.PO ISON SUMAC is an uncommon species that is found throughout the

southern part of the state and as far north as Penobscot County. It occurs as a smalltree in low, wet swamps. It is particularly common around Mt. Agamenticus insouthern Maine.

The leaves are alternate, 7–14 inches long, consisting of 7–13 leaflets along asmooth greenish-red rachis. Leaflets have entire margins, short stalks, are darkgreen and lustrous above with scarlet midribs, and paler and glabrous below. Twigsare without hairs.

Poison ivy (above) and poison sumac (below) are two plants everyone going into thewoods should know how to identify and avoid.

Page 9: Forest Trees of Maine

7A F E W P R E C A U T I O N S

The fruit is a globose, slightly compressed, thin-fleshed, ivory white or tawnywhite berry, about 1/5 inch in diameter; it is borne in loose, pendent axillary clustersthat ripen in September, but persist on the tree far into winter.

T I C K S

About 13 different species of ticks live in Maine. One of these species, the deertick (Ixodes scapularis) can transmit the bacterium that causes Lyme disease. Lymedisease frequently starts with a rash and flu-like symptoms, and if untreated mayprogress to neurological problems.

Ticks are most common in coastal and south-central Maine. When goinginto the woods in areas known to have high tick populations, it is wise to take someprecautions to help avoid tick bites. For example:

• Tuck your pant legs into your socks and your shirt into your pants.• Wear light-colored clothing so ticks can be seen more easily.• Use a repellent containing DEET according the label directions. Pay specialattention to treating shoes, socks and pant legs. Use caution in applying high-concentration products to the skin, especially on children.• To protect pets, consult your veterinarian about tick repellents.• Inspect yourself, your clothing, your children, your companion and your petswhen you get in from the field. Ticks often attach to body folds, behind theears and in the hair. If possible, shower and wash clothes immediately. Heatdrying is effective in killing ticks.Prompt removal of ticks from skin is very important. To remove a tick, grasp

it as close to the skin as possible, preferably with tweezers, and pull gently but firm-ly until the tick lets go. Do not handle the tick with bare hands. Apply antisepticto the bite. You should consult your physician if you develop a large rash at the siteof the tick bite or if you remove an engorged deer tick.

For more information on ticks, visit the Maine Medical Center ResearchInstitute Vector-borne Disease Laboratory’s Lyme disease research Website:www.mmcri.org/lyme/lymehome.html.

Dog Tick Deer Tick

Male

Female

Male

Female

Distr ibut ion of submitted deer t icks1989-2006

Page 10: Forest Trees of Maine

When supplies could notbe brought to loggingcamps by water, theywere transported overland on “tote” roads.

Page 11: Forest Trees of Maine

H O W T O U S E T H E K E Y S T O I D E N T I F Y T R E E S 9

H

This book contains keys to help you identify trees in both winter and summer.These are dichotomous keys; they work by giving you pairs of choices called couplets.Each couplet has the same number located on the left side of the key.

Begin at couplet number 1. Read both choices carefully to determine whichmatches the tree you are trying to identify. After you make your choice, the num-ber at the right tells you which couplet to go to next. Go to that couplet and decidewhich choice matches your tree; repeat the process until you arrive at a name orspecies group for your tree. Turn to the page indicated and compare your tree to thespecies in the table to figure out the individual species.

The final step is to compare your tree to the pictures and drawings in thebook. If they don’t seem to match your specimen, don’t be discouraged; return tothe key and check to see if you made an error. Remember that leaves and bark canvary a lot even on the same tree, but the photograph can only show one example.To help you with the terms in the key, a glossary is provided on page 14.

Example: We want to identify the tree these needles camefrom. Starting at the first couplet, choose the descriptions thatfit the specimen. The lines in red indicate the correct choicesin the key.

SEE GLOSSARY PAGE 14 GO TO

1. Leaves are needle-, awl- or scale-like; conifers 21. Leaves are broad and veined, not as above; hardwoods or broad-leaf trees 92. Leaves needle-like 32. Leaves awl- or scale-like, or both 73. Leaves flat, tips blunt, and occur singly 43. Leaves angular in cross section, tips pointed 54. Leaves taper, twigs limber; cones shorter than 1 inch Eastern Hemlock p. 484. Leaves parallel-sided, twigs stiff; cones over 2 inches Balsam Fir p. 465. Leaves occur singly, never clustered Spruce p. 375. Leaves occur in clusters, also singly in larch 66. Leaves in clusters of 2–5 with papery sheath at base1 Pine p.25 6. Leaves in clusters2 of 8 or more on spurs; papery sheath lacking Tamarack p. 50

We now know the tree is a pine. We then go to the species table to figure out what species of pine it is.

The tree is Eastern white pine.

Page 12: Forest Trees of Maine

10 W I N T E R K E Y

SEE GLOSSARY PAGE 14 GO TO

1. Leaves are evergreen 21. Leaves are deciduous (dead leaves may remain attached) 102. Leaves are needle- or scale-like; fruit is a cone; conifers 42. Leaves are broad and flat, often curled in winter 33. Leaves to 3 inches long Mountain laurel p. 1693. Leaves 4–8 inches long Rhododendron p. 1704. Leaves are needle-like 54. Leaves are awl- or scale-like or both 85. Leaves in clusters of 2, 3 or 5 with a papery sheath at the base (may be lacking in white pine) Pine p. 25-365. Leaves attached to the twig singly 66. Leaves angular in cross section, will roll easily between the fingers Spruce p. 37-456. Leaves flat in cross section, will not roll easily between the fingers 77. Leaves taper, attached to twig with tiny stem; bark thick, purple under scales Hemlock p. 487. Leaves parallel-sided, attached directly to twig with round base like tiny suction cup; bark thin with resin blisters Fir p. 468. Branchlets with awl-shaped leaves; leaves prickly Juniper, Red cedar p. 56-588. Branchlets with scale-like leaves; leaves not prickly 99. Twigs flat; cones oblong, up to 1⁄2 inch; common statewide Northern white-cedar p. 549. Twigs slightly flattened; cones 1⁄4 inch, rounded; rare tree of southern and midcoast Maine Atlantic white cedar p. 52

10. Older twigs with many short spur branches less than 1⁄4 inch long; fruit a cone Tamarack p. 5010. Spur branches lacking or if present are longer than 1⁄4 inch; fruit not a cone 1111. Leaf scars are opposite 1211. Leaf scars are alternate 1712. 3 bundle scars (may be obscured in flowering dogwood, see 15) 1312. More than 3 bundle scars 1613. More than 2 bud scales Maple p. 70-8513. 2 bud scales 1414. Buds long and narrow, base of terminal bud swollen; bud scales covered with minute, scale-like particles Nannyberry p. 16614. Buds not long and narrow; bud scales not covered with minute, scale-like particles 1515. Lateral buds hidden; rare small tree of southwestern Maine Flowering dogwood p.17115. Lateral buds conspicuous; common small trees Maple p. 70-8516. Buds shiny and sticky in spring before flowering Horsechestnut p. 15816. Buds not shiny; bud scales covered with minute, scale-like particles Ash p. 127-133 17. Twigs armed with spines, thorns or branches ending in a spine 1817. Twigs are unarmed 2018. Armed with paired spines less than 1 inch long Black locust p. 16218. Armed with thorns or branches ending in a spine greater than 1 inch long 1919. Armed with thorns that occur just above the leaf scar Hawthorn p. 14719. Armed with branches ending in a spine Canada plum p. 14620. Buds not visible Black locust/Honey locust p. 162-16420. Buds visible 2121. Leaf scar nearly encircling the bud 2221. Leaf scars extending less than 3⁄4 of the way around the buds 2322. Buds covered by scales; twigs not hairy; bark mottled in color Sycamore p. 15922. Buds naked; twigs very hairy; bark with prominent lenticels Staghorn sumac p. 16823. Pith chambered or diaphragmed 2423. Pith solid 2524. Pith chambered Butternut or Walnut p. 155-15724. Pith diaphragmed; uncommon tree of swamps in southern Maine. Black gum p. 160

W K

Page 13: Forest Trees of Maine

11W I N T E R K E Y

25. Buds naked 26 25. Buds covered by one or more scales 2726. Terminal buds scalpel-shaped Witch-hazel p. 16726. CAUTION POISONOUS Terminal buds ovoid Poison sumac p. 627. Buds covered with a single cap-like scale Willow p. 6827. Buds covered by 2 or more scales 2828. A single bundle scar; crushed twigs aromatic; rare tree of southwest Maine Sassafras p. 16528. More than 1 bundle scar 2929. Catkins present 3029. Catkins absent 3230. Buds stalked; 2 types of catkins present Alder p. 10230. Buds sessile; 1 type of catkin present 3131. Bud scales with tiny grooves; gray bark with loose vertical scales Eastern Hop-Hornbeam p. 9831. Bud scales without grooves; bark peeling or blocky, not with loose vertical scales Birch p. 86-9632. Terminal bud absent, the end bud is pseudo-terminal (except on spur shoots) 3332. Terminal bud present 3833. Up to 4 bud scales (except on spur shoots) 3433. 5 or more bud scales 3634. Bud scales deep red; fruit hard and round, borne in cymes attached to a bract American basswood p. 13434. Bud scales other than deep red; fruits not attached to a bract 3535. Pith star-shaped (rare tree) American chestnut p. 12435. Pith round, fruit borne in catkin-like cones Birch p. 86-9636. Buds four-angled, square in cross section; stem fluted, gun metal gray American Hornbeam p. 10036. Buds not four-angled, trunk and bark not as above 3737. Buds with small vertical grooves, yellow-green, round in cross section; grayish-brown bark peeling into vertical scales

Eastern Hop-Hornbeam p. 9837. Buds without small vertical grooves, brown, often laterally flattened; bark corky, ridged, often with alternating light and

dark layers in cross section Elm p. 136-13738. Leaf scars very long and narrow, several times longer than broad 3938. Leaf scars broader, at most 3 times longer than broad. 4039. Second bud scale more than 1⁄2 the length of the bud; buds always uniformly colored Mountain ash p. 15039. Second bud scale less than 1⁄2 the length of the bud; buds sometimes bi-colored—reddish and greenish Serviceberry p. 14840. Pith star-shaped 41 40. Pith round or if angled without 5 points. 4541. Buds clustered toward the tip of the twig Oak p. 106-12341. Buds not clustered toward the tip of the twig 4242. Terminal bud more than 3⁄8 inch long; bud scales loose; older trees with gray shaggy bark Shagbark hickory p. 15242. Terminal buds less than 3⁄8 inch long; bud scales not loose; bark of older trees fissured and ridged or smooth, not shaggy 4343. Buds sulfur yellow (rare tree of southwestern Maine) Bitternut hickory p. 15443. Buds brown to reddish-brown 4444. Lowest bud scale centered over the leaf scar; wood diffuse porous Aspen/Poplar p. 61-6744. Lowest bud scale not centered over the leaf scar; wood ring porous (rare tree) American chestnut p. 12445. Buds long and narrow, several times longer than broad, diverge from the twig at a wide angle; bark smooth gray or often

pockmarked with small cankers American beech p. 10445. Buds not long and narrow, do not diverge from twig at wide angles. 4646. Nodes often clustered toward the ends of twig; bark of dead branchlets yellowish-orange Alternate-leaved dogwood p. 17246. Nodes not clustered; dead branchlets not yellowish-orange 4747. Buds stalked; pith triangular Alder p. 10247. Buds sessile; pith round Cherry p. 139-145

Page 14: Forest Trees of Maine

S U M M E R K E Y12

S K SEE GLOSSARY PAGE 14 GO TO

1. Leaves are needle-, awl- or scale-like; conifers 21. Leaves are broad and veined, not as above; hardwoods or broad-leaf trees 92. Leaves needle-like 32. Leaves awl- or scale-like, or both 73. Leaves flat, tips blunt, and occur singly 43. Leaves angular in cross section, tips pointed 54. Leaves taper, twigs limber; cones shorter than 1 inch Eastern Hemlock p. 484. Leaves parallel-sided, twigs stiff; cones over 2 inches and upright Balsam Fir p. 465. Leaves occur singly, never clustered Spruce p. 37-455. Leaves occur in clusters, also singly in larch 66. Leaves in clusters of 2–5 with papery sheath at base1 Pine p. 25-36 6. Leaves in clusters2 of 8 or more on spurs; papery sheath lacking Tamarack p. 507. Branchlets with prickly, awl-shaped leaves; cones are berry-like Juniper/Eastern Redcedar p. 56-58 7. Branchlets with scale-like leaves; leaves not prickly; cones un-berry-like 88. Twigs flat; cones oblong, woody, up to 1⁄2 inch; wood slightly aromatic Northern White Cedar p.54 8. Twigs slightly flattened; cones 1⁄4 inch, rounded, leathery; wood strongly aromatic Atlantic White Cedar p. 529. Leaves opposite, trees only 109. Leaves alternate 15

10. Leaves simple 1110. Leaves compound 1311. Leaf margin serrate Nannyberry p. 16611. Leaf margin lobed or entire 1212. Leaf margin lobed Maple p. 70-8512. Leaf margin entire Flowering Dogwood p. 17113. Leaves palmate Horsechestnut p. 15813. Leaves pinnate 1414. 3–5 leaflets, lobed, coarse teeth Boxelder p. 8414. 5–13 leaflets Ash p. 127-13315. Leaves simple 1615. Leaves compound 3616. Leaf margin entire, wavy, or lobed 1716. Leaf margin toothed or serrate 2417. Leaf margin entire 1817. Leaf margin wavy or lobed 2118. Leaves thin, veins parallel Alternate Leaf Dogwood p. 17218. Leaves thick and leathery, net-veined 1919. Pith diaphragmed; leaves 2–5 inches long Black Tupelo p. 16019. Pith not diaphragmed 2020. Leaves to 3 inches long Mountain Laurel p. 16920. Leaves 4–8 inches long Roseberry Rhododendron p. 17021. Leaf margin wavy toward tip; base of leaf one-sided Witch-Hazel p. 16721. Leaf margin lobed or wavy throughout 2222. Leaf petiole hollow and covers bud; numerous main leaf veins radiate from base American Sycamore p. 15922. Leaf petiole neither swollen nor hollow; leaves with one main vein 23

1 Papery sheath on white pine drops in late August2 Larch leaves are borne singly on elongating shoots

Page 15: Forest Trees of Maine

13S U M M E R K E Y

23. Twigs angular; pith star-shaped Oak p. 106-12323. Twigs round, spicy odor and taste; leaves 0–3 lobed Sassafras p.16524. Leaf margin singly toothed or serrated 2524. Leaf margin doubly serrated 3125. Teeth hooked, prominent; fruit a bur 2625. Teeth not hooked, fruit not a bur 2726. Pith star-shaped; buds blunt; bark brown American Chestnut p. 12426. Pith round; buds long, pointed; bark gray American Beech p. 10427. Leaf base one-sided, leaf cordate; pith not symmetrical American Basswood p. 13427. Leaf base even; pith symmetric in cross section 2828. Leaves long and narrow; petioles short without glands; buds with a single, cap-like scale Willows p. 6828. Leaves broad, or if narrow with glands on petiole; buds with several scales 2929. Leaf petiole usually long, flat, except rounded in balsam poplar; pith star-shaped Aspen/Poplar p. 61-6729. Leaf petiole short, not flat; pith round 3030. Twigs pungent when broken; glands on petiole Cherry, Plum p. 139-14530. Twigs odorless; leaf petiole glandless; buds slender, twisted at tip, silky within Serviceberry p. 14831. Leaf base one-sided, surface sand-papery Elm p. 13631. Leaf base even, surface smooth 3232. Branches with thorns 1 inch or more long Hawthorn p. 14732. Branches without thorns 3333. Pith triangular; buds stalked, smooth Speckled Alder p. 10233. Pith not triangular; bud scales overlapping 3434. Leaves hairy on both surfaces; pith green Eastern Hop-Hornbeam p. 9834. Leaves if hairy only so on one surface; bark smooth 3535. Stem fluted; bark smooth, gun-metal gray American Hornbeam p. 10035. Stem not fluted; bark white, yellow, or red to dark brown Birch p. 87-9636. Leaflets with margin entire 3736. Leaflets with serrated margin 3837. Twigs with paired spines; 7–19 leaflets Black Locust p. 16237. Twigs spineless; 7–13 leaflets; leaflets poisonous Poison-Sumac** p. 638. Leaflets 1⁄2 inch long with fine, rounded teeth Honeylocust p. 16438. Leaflets over 1 inch long 3939. Pith chambered or diaphragmed Black Walnut/Butternut p. 155–15739. Pith solid 4040. 5–7 leaflets; pith star-shaped Shagbark/Bitternut Hickory p. 152–154 40. 11–31 leaflets 4141. Twigs smooth; 11–17 leaflets; buds large Mountain Ash p. 15041. Twigs densely hairy; 11–31 leaflets; buds small Staghorn Sumac p. 168

** See also Poison ivy, page 6

Page 16: Forest Trees of Maine

G L O S S A R Y14

G Structure in brackets indicates part to which the term applies.

Abortive [fruit] Not developed completely.

Alternate [arrangement of leaves or buds] Not opposite on sides of twig.

Appressed Pressed close or lying flat against something.

Awl-shaped [leaf ] Narrow and tapering to a sharp point.

Axillary Growing from the Axil. The angle between the upperside of a leaf or stem and the supporting stem or branch.

Basal disc [fruit] A plate-like structure on the base of a fruit.

Bloom A whitish covering; usually on new shoot growth or fruit.

Bole The main stem of a tree; usually the part that is commercially useful forlumber or other wood products.

Bract A leaf-like structure which is attached to a flower, a fruit or to its stalk.

Branchlet Shoot growth of the latest growing season.

Broadleafed Having relatively broad rather than needle-like or scale-like leaves.

Bur [fruit] A prickly or spiny husk enclosing the seed.

Capsule [fruit] A dry fruit enclosing more than one seed and splitting freelyat maturity.

Catkin A compact, cylindrical cluster of flowers of the same sex.

Chambered [pith] With hollow cavities separated by discs or plates.

Compound [leaf ] A leaf composed of smaller leaf units or leaflets.

Conical Wide at the base and gradually tapering to a point;circular in cross section.

Conifer Cone-bearing trees; the “evergreens.”

Cordate [leaf ] Heart-shaped at the petiole end or base.

Corymb A flat-topped floral cluster with outer flowers opening first.

Cup [fruit] The scaled, concave basal portion of oak fruit.

Cyme A flattened flowering structure, center flowers bloom earliest.

Deciduous [leaves] All leaves drop in the autumn; not evergreen.

Diaphragmed [pith] Solid but divided into sections by firmer discs.

Drupe [fruit] Fleshy outside, hard and stone-like inside.

Alternate

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15

Ellipsoid Tapers equally at both ends; more than twice as long as broad.

Elliptical Like an ellipse; flat and tapering equally at both ends.

Entire [leaf ] Margin of leaf without teeth, lobes, or divisions.

Fascicle [leaf ] A cluster of conifer leaves.

Fluted [stem] With alternating, rounded depressions and ridges.

Fruit The seed-bearing part of a tree.

Glabrous Smooth, without hairs

Glands Generally raised structures at the tips of hairs, or on a leaf, petiole, or twig.

Globose Spherical or globe-shaped.

Habitat The place where a plant usually grows, e.g. rocky, moist, well-drained, etc.

Hardwood Term used to describe all broad-leaved trees. These tree species aretypically deciduous, retaining their leaves only one growing season. Despite theterm, some “hardwoods,” such as the aspens, have wood that is relatively soft.

Head A compact aggregate of flowers or fruit on a common stalk.

Husk [fruit] The somewhat leathery, outer covering of a fruit sometimes capableof splitting along well-defined lines.

Invasive Not native to and tending to spread widely in a habitat or environment,sometimes displacing native species.

Lance-shaped Long and tapering; several times longer than broad; broadestat the base.

Leaf Stalk (petiole) and blade of hardwoods; needles and scales of conifers.

Leaflets Smaller leaf units which together form a compound leaf.

Lenticel [bark] Corky, raised pores on woody parts with openings for air-gasexchange.

Linear [leaf ] Much longer than broad with parallel margins.

Lobed [leaf ] With large, rounded or pointed projectionsalong the leaf margin. Projection formed by indentations ofthe leaf margin.

Margin [leaf ] The edge, perimeter, or portion formingthe outline.

Midrib [leaf ] The large central vein.

Oblong Longer than wide with nearly parallel sides.

G L O S S A R Y

Lobed

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16

Obovate Egg-shaped in outline; broadest above the middle.

Opposite [arrangement of leaves or buds]Directly across from one another on a common axis, or twig.

Oval Somewhat elliptical; less than twice as long as broad.

Ovate Egg-shaped in outline; broadest below the middle.

Ovoid An egg-shaped solid.

Palmate [leaf or veins] Compound, with leaflets originatingat the same point on a common stalk. Veins originating at acommon point at base of leaf blade.

Panicle A loosely branched, pyramidal cluster of flowers.

Pendulous Drooping or hanging downward.

Petiole [leaf ] The stalk that supports the leaf blade.

Pinnate [leaf or vein] Compound, with leaflets along a commonrachis or stalk. Veins originating along a common mid-vein.

Pistillate Containing female portions of flowers, or the pistils.

Pith The central, soft part of the stem.

Prickle A small spine-like growth.

Pseudo-terminal [bud] When the bud on the end of a twig has a leaf scarlocated directly below.

Pubescent Covered with hairs.

Raceme Numerous stalked flowers or fruit along a common axis.

Rachis The common stalk in a compound leaf to which the leaflets are attached.

Ranked [leaves] Arranged in rows or files.

Samara A winged fruit, e.g. ash, maple.

Scales [bud] Small, modified leaves on the outer surface ofbuds.

Scales [cone] The basic structures that enclose the seeds.

Scale-like [leaf ] Small, generally overlapping, triangular-shaped leaves of some conifers.

Seed That part of the fruit capable of germinating and producing a new plant.

Serrate [leaf ] Margins with a saw-tooth outline. Doubly serrate: with smallteeth on the larger teeth.

G L O S S A R Y

Scale- l ike

PinnateVeins

Opposite

PalmatelyCompound

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17

Sessile Attached directly by the base without an intervening stalk.

Shrub A woody, many-stemmed plant, usually under 15 feet in height at maturity,which branches from its base.

Simple [leaf ] A single leaf composed of a single blade. Not compound.

Smooth Without hairs, glands, or any roughness.

Softwood Term used to describe all needle-leaved trees. These species are typicallyevergreen, retaining their leaves through two or more growing seasons. Larches,including tamarack, are exceptions, being deciduous “softwoods.”

Solid [pith] Without cavities or sections separated by discs.

Spike A flower stalk.

Spinescent Having a spine or spines; or terminating in a spine.

Spur A short, extremely slow-growing, woody twig projection.

Staminate Containing male portions of flowers, or the stamens.

Stipule A tiny, leafy, sometimes spiny projection arising at the base of a petiole.

Stomate Plural stomata Small pore on a leaf used for gas exchange.

Stone The “bony” or stony pit of drupes.

Style The usually slender part of a pistil, situated between the ovary and the stigma.

Toothed [leaf ] With moderate projections along the margin.

Tree A woody plant, generally single-stemmed, that reaches a height of morethan 15 feet at maturity and a diameter of 3 inches or more measured at 4½ feetabove the ground.

Umbel A group of flowers or fruit whose stalks have a common point of attachment.

Unequal [leaf base] Base parts of blade on either side of midrib are uneven.

Valve-like [bud scales] Meet at their margins and do not overlap.

Wavy [leaf margin] Undulating but smooth; not toothed nor lobed.

Whorl [leaves or branches] More than two originating at the same level on a common axis.

G L O S S A R Y

Whorled

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18 G L O S S A R Y

M A R G I N S

L E A F S H A P E S

Ovate Lanceolate Cordate El l ipt ical(Lance-shaped) (Heart-shaped)

Obovate Oblong L inear

Crenate (Round toothed)

Doubly-Serrate Dentate(Doubly toothed) (Coarsely toothed)

Ent ire Serrate (Toothed)

Page 21: Forest Trees of Maine

19G L O S S A R Y

F L O W E R T Y P E S

Awl-shaped

L E A F S T R U C T U R E S

Simple Leaf Pinnately Compound Leaf Palmately Compound Leaf

N E E D L E T Y P E S T W I G S T R U C T U R E

Scale-shapedNeedle-shaped

Raceme Spike Panicle (of Racemes)

Cyme Umbel Corymb

Terminal Bud

Leaf Scar

Lateral Bud

Lenticel

Bud Scale

Vascular Bundle Scar

Terminal Bud Scale Scar

Pith

Stem TipLeaf Scar

Pseudoterminal BudPeduncle

Pedicel

Leaflet

BudPetiole

Petiolule

Rachis

Blade

Leaf

Bud

Stem

Stipule

Petiole

Midrib

Page 22: Forest Trees of Maine

20 T R E E P A R T S A N D F U N C T I O N S

T P FA tree has three major parts: roots, trunk and crown.

Large roots anchor the tree and store foods which are manufactured in theleaves. Small roots and root hairs absorb water and dissolved mineral salts from thesoil. These raw materials are conducted upward to the leaves where they are utilizedin the synthesis of necessary plant food. Air must be present in the soil for the rootsto live, although some species can endure several months of flooding.

The trunk is the main body of the tree. In the center of the trunk is the pith.Next to the pith is the heartwood which is composed of dead cells and serves assupport. On the outer side of the heartwood is the sapwood, which contains thesap conducting tubes. Sapwood is usually lighter in color, but it darkens with ageand becomes heartwood. Heartwood and sapwood together comprise the xylem.Outside the sapwood is the cambium, a thin layer of cells, which annually producesnew sapwood inwardly and new inner bark outwardly. The cambium producesdiameter growth, and callus growth around open wounds. The inner bark orphloem is outside the cambium and carries food from the leaves downward tonourish the cambium and growing parts. The outer bark is the outer-most part ofthe tree. Essentially, it is composed of dead cork cells and protects the inner barkfrom mechanical injury, drying or disease; it also insulates the phloem fromextremes of heat and cold. Damage to the phloem causes interference with foodmovement to growing parts below the injury. Girdling of a tree through its innerbark will kill the tree. Wood or medullary rays radiate out from the center, and servein lateral conduction and as food storage areas. They are most conspicuous in across-sectional view.

The crown is composed of branches, twigs, buds, leaves, flowers and fruit.The process of photosynthesis occurs in the leaves. Using energy produced bysunlight, the leaves combine carbon dioxide from the air and water from the soilto produce carbohydrates. Oxygen is released in the process. Carbohydrates plusfats and proteins are the plant foods necessary for growth and respiration of thetree. Flowers and fruit are important in reproduction.

A N N U A L R I N G S

The yearly growth of a tree can be compared to the annual placement of hol-low wooden cones, one on top of the other. Each cone would represent a singleyear’s growth over the entire stem. At the beginning of each new growth period,new wood cells are large and thin-walled, and form the springwood or early wood.As the growing season progresses, the smaller, thicker-walled cells of the summer-wood or late wood are produced. The darker appearance of the late wood delineatesthe annual ring of growth put on by a tree. The age of a tree, at any desired pointalong the trunk, can be determined by counting these annual rings.

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21T R E E P A R T S A N D F U N C T I O N S

The relationship between tree age and size can be very deceptive. Many treespecies can survive in a shaded understory for years, with nearly microscopic growthrings. These tree species are called shade tolerant. When the canopy is partially orwholly removed by harvest, mortality or weather, increased available sunlight allowsaccelerated growth. This process is known as release. Some long-lived species, suchas red spruce, can survive for decades in the understory, then for centuries in theoverstory. Others are shade tolerant but short lived; for example, balsam fir rarelyexceeds 100 years in total age. Still other species such as quaking aspen and paperbirch are intolerant of shade and relatively short-lived. An observer with knowledgeof tree species and growth characteristics can deduce the history of a forest withoutcutting or boring holes in stems to count rings.

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22 A U T U M N C O L O R A T I O N

A C Autumn foliage coloration, one of Maine’s

greatest aesthetic assets, is enjoyed every year, gen-erally with little appreciation for the processesresponsible for it. Most hardwoods produce dra-matic leaf coloration if climactic conditions arefavorable, while conifers usually produce only weakcoloration of yellow and brown.

Hardwoods contain green, yellow and orangepigments in their leaves. Chlorophyll allows thegreen to be the most prominent of the pigments;however, the green pigment is also the least stable.It is repeatedly produced and destroyed throughoutthe summer and masks the carotenoid pigments(xanthophyll and carotene) which give the yellow

and orange shades. As autumn approaches, chlorophyll is destroyed faster than itis produced. As the chlorophyll disappears, the carotenoid pigments begin toshow. The purple and brilliant red shades become visible from the production ofanthocyanin pigments, which are also capable of masking the carotenoids.Tannins cause brown shades in some species.

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23A U T U M N C O L O R A T I O N

Certain conditions favor maximum autumn coloration. They include adequatesummer rainfall, adequate sugar accumulations in the leaves and prolonged periodsof cool, bright, sunny weather without severe frosts. Frost is not an essential elementfor leaf coloration. In fact, weakened trees occasionally color in mid-summer.

Although variations are numerous, a general guideline to autumn tree coloration is listed below.

Y E L L O W Stamarack, green ash, black ash, basswood, beech, birch, butternut, elm,boxelder, mountain maple, silver maple, striped maple, sugar maple,mountain-ash, poplar, serviceberry, willow, witch-hazel.

R E D / S C A R L E Thornbeam, red maple, mountain maple, sugar maple, black oak,red oak, scarlet oak, white oak, sumac, tupelo.

O R A N G E sugar maple.

B R O W N S black oak, beech.

P U R P L E white ash.

Although much less appreciated than fall color, the subtle color of Maine’s trees in early spring canbe just as dramatic.

Page 26: Forest Trees of Maine

C

Often referred to as “softwoods”, conifers belong to the group of plants

known as the gymnosperms. Conifers are cone bearing trees and shrubs

that have needle or scale like leaves and resinous wood. All of Maine’s

conifers except for tamarack are evergreen. There are 16 species of conifer

native to Maine, 13 of these are trees and three others are more commonly

found as shrubs. Several other species of conifer native to other parts of

the world are commonly planted in Maine for both ornamental and timber

production purposes. Photo location: Penobscot Experimental Forest, Bradley, Maine.

Page 27: Forest Trees of Maine

Eastern White PinePinus strobus

Red PinePinus resinosa

Pitch PinePinus rigida

N E E D L E S

NUMBER/CLUSTER 5 2 3

DESCRIPTION Slender, flexible, 3–5 inches Straight, flexible, 4–6 inchesStout, not flexible, usually twisted,

grow at right angles to the branchlets, 3–5 inches

COLOR Bluish-green Dark green Dark yellow-green

SHEATH Shed in late August Persists Persists

C O N E S

LENGTH 4–8 inches 11⁄2–21⁄4 inches 11⁄2–31⁄2 inches

DESCRIPTIONBorne on a long stalk; thin smooth

scales without prickles

Borne on short stalks; scaleswithout prickles. Several basal

scales remain on branches when cone drops.

Borne on a short stalk, havingprickles on the cone scales, flat-

based when completely open.Often remain on branches

for 10–12 years.

Jack PinePinus banksiana

Scots PinePinus sylvestris

N E E D L E S

NUMBER/CLUSTER 2 2

DESCRIPTIONStout, flat, twisted,

3⁄4–11⁄2 inchesStout, stiff, twisted,

11⁄2–3 inches

COLORLight yellow-green, later

becoming dark greenDull blue-green

SHEATH Persists Persists

C O N E S

LENGTH 11⁄2–2 inches 1–2 inches

DESCRIPTION

Much curved inward, without stalk. Prickles minute.

Often remain on branches for many years.

Egg shape, borne on a short stalk,scales with occasional prickles.

25

PI

NE

P I N E S

P The Important Distinctions

Page 28: Forest Trees of Maine

26 E A S T E R N W H I T E P I N E

Eastern white pine has been animportant tree for the people of

what is now the State of Maine forhundreds, if not thousands, of years.

Therefore, it is no coincidence thatMaine has come to be known as the“Pine Tree State.” Recognizing itsimportance, in 1895 the Maine legisla-ture designated the “Pine Cone andTassel” as Maine’s official floralemblem. In 1945 the legislatureResolved: “That the white pine tree be,and hereby is, designated the officialtree of the State of Maine.”

The availability and high qualityof white pine lumber has played animportant part in the development andeconomy of Maine since 1605, whenCaptain George Weymouth of theBritish Royal Navy collected sampleshere and brought them back to Englandfor display.The shortage of ship masts inEurope led to England’s Broad ArrowPolicy in 1691, whereby pines 24 inchesor more in diameter within 3 miles ofwater were blazed with the mark of the

E W P Pinus Strobus L.

Maine is known asthe “Pine Tree State”and the Eastern white pineis the official tree of theState of Maine.

E A S T E R N W H I T E P I N E

Page 29: Forest Trees of Maine

27E A S T E R N W H I T E P I N E

broad arrow; such trees to be reservedfor use in the Royal Navy. The termKing’s Arrow Pine originated from thispolicy. Most of the accessible virgin pinewas cut by 1850. Lumber productionreached its peak in 1909, but white pineis still a valuable species that contributesgreatly to the economy of the state.

White pine occurs in all localitiesin the state in moist situations, onuplands and on sandy soil, but developsbest on fertile, well-drained soils. Onsandy soil it often becomes establishedin pure or nearly pure stands. It is one ofthe major species planted in the state.The tree grows rapidly both in heightand diameter, making an average growthin height of 1 foot or more each year.

When growing in the open, theyoung tree is symmetrical and conical inoutline except when deformed by whitepine weevil. White pine weevil is aninsect that kills the topmost shoot, andoften causes the tree to have multiplestems and a round profile. In the forest,a white pine tree has a narrow head; and

the trunk is commonly free of livebranches for a considerable portion of itslength. Old forest trees have a broad andsomewhat irregular head. The branchesare horizontal and in regular whorls,usually of 5 each. Very old trees oftenbecome very irregular and picturesque.The trunk tapers gradually, and the treeoften attains a height of 100 feet.Commonly it is from 70–80 feet tall,and has a diameter of 1–3 feet.

The bark of young trees is smoothand thin, green with a reddish-browntinge overall, or brown in spots. On oldtrees, it is from 1–2 inches thick, verydark, and divided into broad, flat ridgesby shallow fissures.

Leaves are in clusters of 5, flexible,3–5 inches long, bluish-green butwhitish on one side. The papery sheathat the base of the new needle clustersfalls in late August.

The cones are 4–8 inches long,cylindrical and borne on a long stalk.They take 2 years to mature, and open todischarge the seed shortly after ripening

E A S T E R N W H I T E P I N E

PI

NE

Page 30: Forest Trees of Maine

28 E A S T E R N W H I T E P I N E

in late August through September of thesecond season.

The wood is light in color anddurable, except when in prolonged con-tact with moisture. It is soft, not heavyand is easily worked. The wood is usedextensively for interior trim, doors, win-dows, cabinetmaking, sash and doormanufacture, patternmaking, furniture,small building construction, interior andexterior finish, and boat planking.

Pine furniture is always popular inNorth America. Lumber from Maine is

Eastern white pine leaves (needles) are 3–5 inches longand in clusters of 5.

Young bark (left) and old bark (right).

N AT I O N A L A N D M A I N E R E G I S T E R O F

B I G T R E E S 2 0 0 8Eastern White Pine

Circumference: 229''Height: 125'

Crown Spread: 72'Location: Morrill

Nominator: C. Brown 2004

sold from Newfoundland to Washing-ton state and south into Mexico. Lowergrade boards have clear sections cut tosize for sale. These clear short piecesmay also be finger-jointed to createlonger lengths of clear wood. Any part ofa pine not making log grade is used forpulp. Ceiling tiles and paper are madefrom this pulp.

Page 31: Forest Trees of Maine

White pine has been an important timber tree in Maine for more than 300 years.

Page 32: Forest Trees of Maine

30

Red or Norway pine,though common,is found only locally throughout

the state, growing on dry, rocky ridges,or light, sandy soil. Stands are usuallyscattered through forests of otherspecies.The beautiful “Cathedral Pines”occur near Eustis.

Young trees often have branchesextending to the ground and form aconical outline. Later, the head is round-ed and picturesque. Branches are gen-erally horizontal. It attains a height of60–80 feet, and a diameter of 2–3 feet.The trunk is straight and tapers slowly.Red pine is not tolerant of shade.

The reddish-brown bark is dividedinto broad, flat ridges by shallow fissures.

The leaves are arranged in clustersof two. They are 4–6 inches long, darkgreen, soft and flexible. When doubledbetween the fingers, they break cleanly,at a sharp angle.

The cones are egg-shaped and areabout 2 inches long. They lack pricklesand are borne on short stalks. The base

R P Pinus Resinosa Soland.

The red pine is named forits reddish-brown bark andpale red heartwood.

R E D P I N E

Page 33: Forest Trees of Maine

R E D P I N E 31

of fallen cones is hollow.They mature inthe fall of the second season and usuallyremain on the branches until the follow-ing summer. Cones may be collected forseeds from September throughout thefall and winter, due to their gradualrelease of seed.

The wood is a little heavier andharder than white pine, close-grained,and fairly strong. It is used for lumber,poles, piles, building construction andpulp. It is treated readily with woodpreservatives, and therefore is a locally-produced alternative to southern yellowpine. Older stands produce large, high-value poles.

Owing to the reddish bark andthe pale red heartwood, the name “redpine” is appropriate.The name “Norwaypine” refers to its original finding nearNorway, Maine. Since it implies thatthe tree is foreign in origin, use of thisname is discouraged.

The reddish-brown bark of the red pine is dividedinto broad, flat ridges by shallow fissures.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Red Pine Circumference: 88'' Height: 96' Crown Spread: 36' Location: Weld Nominator: M. Gill 1988

PI

NE

Page 34: Forest Trees of Maine

32 J A C K P I N E

J P Pinus banksiana Lamb.

Jack or gray pine grows on sandy,rocky, shallow acidic soils. It is known

to occur naturally at Alamoosook Lakein Orland, Schoodic Point in WinterHarbor, Great Wass Island in Beals,Matagamon Lake, Cliff Lake, LobsterLake, and in the areas south and westof Jackman.

The spreading branches are longand flexible, and form an open headsymmetrical in outline. At maturity thetree is about 50–60 feet tall and 8–10inches in diameter. Trees in the coastalpopulations tend to be much shorterand usually have a picturesque, gnarledlook. Cones are often produced whenthe trees are only a few years old.

The cones of thejack pine usually remain closed for several years unlessexposed to extreme heat, andoften do not fall for 12–15 years.

Page 35: Forest Trees of Maine

33J A C K P I N E

The bark is thin with irregularrounded ridges. It is dark brown with aslight tinge of red. The leaves are inclusters of two, and are ¾ to 1½ inch-es long. They are stout, yellow-greenat first, dark green later, rather flat, andtwisted at the base. The cones require 2years to mature, are rather slender,1½–2 inches long, lack a stalk and arecurved. The scales have minute prick-

les that are often deciduous. The conesusually remain closed for several yearsunless exposed to extreme heat, andoften do not fall for 12–15 years.

The wood is moderately hard,heavy, and close-grained. It is usedmostly for pulp; historically it was usedfor firewood and box boards.

Jack pines growing on thecoast in eastern Maine areoften stunted and gnarled.

Jack pine cones are curved and persist onthe tree for many years.

PI

NE

Page 36: Forest Trees of Maine

34 P I T C H P I N E

P P Pinus rigida P. Mill.

P itch pine grows on sandy barrensor plains, and on gravelly soil of

the uplands. It is quite common in thesouthern part of the state, on the sandplains near Brunswick and Oxford andon Mt. Desert Island.

Branches are horizontal, rigid,contorted and form an open crown.Pitch pine attains a diameter of 1–2feet, and a height of only 30–40 feet.The trunk tapers rapidly and generallyis straight. Often the tree producescones when small. It is the only nativepine that will resprout when damagedby such factors as fire.

The bark is rough, even on youngstems and branches. On old trees, it isirregularly divided into continuousbroad flat ridges, and is deep gray orreddish-brown.

The leaves are in clusters of three,and are 3–5 inches long. They are darkyellow-green and stiff, standing atright angles to the branch.

The cones require 2 years tomature, are 1½–3½ inches long, borne

Pitch pine wood is used forconstruction lumber, pulp andfire-starting “fat wood.”

Page 37: Forest Trees of Maine

35P I T C H P I N E

on short, hardly-noticeable stalks, andare often produced in clusters. A sharp,rigid, curved prickle is produced onthe tip of each scale. The cones opengradually during midwinter. Seeds arereleased over a period of several years.Cones often remain on the trees 10–12years. Fresh cones are used in wreathdecorations.

The wood is moderately heavy,strong, hard and stiff. It is used forconstruction lumber, pulp and fire-starting “fat wood.” In the past, con-siderable quantities of pitch and tur-pentine were obtained from this tree;these commodities were referred to as“naval stores,” a term originally appliedto the resin-based components used inbuilding and maintaining wooden sail-ing ships. Today naval stores are usedin the manufacture of soap, paint, var-nish, shoe polish, lubricants, linoleumand roofing material.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Pitch Pine Circumference: 97'' Height: 80' Crown Spread: 44' Location: Poland

Pitch pine cones have a sharp prickle atthe end of each scale.

Pitch pine often has needles growing directlyout of the trunk. This plus its clusters of3 needles make it easy to recognize.

PI

NE

Page 38: Forest Trees of Maine

mature trees is gray to red-brown andhas irregular ridges and furrows.

The leaves are needles in clustersof two. They are 1½–3 inches long,stout, stiff, twisted, dull blue-greenwith distinctive lines of stomata.

The cones are 1½–2 inches longand numerous, even on comparativelyyoung trees; the scales are armed withsmall, blunt spines.

The wood is similar in characterto red pine; however, due to its poorform, in Maine it is seldom used forlumber. It is occasionally used for pulpand fuel.

36 S C O T S P I N E

S (S ) P Pinus sylvestris L.

Scots pine is the most widelydistributed pine in the world but

is not native to Maine. A native ofnorthern Europe and Asia, it growsnaturally from Scotland almost to thePacific Ocean and from above theArctic Circle in Scandinavia to theMediterranean. In parts of its nativerange, Scots pine grows to be a talltimber tree in dense stands. There aremany strains of this species; the treesthat have been planted in Maine oftenhave very poor growth habits. This,plus its susceptibility to snow, porcu-pine and bird injury, makes it undesir-able for timber production here. Scotspine will grow on very poor soils. Somestrains are planted for Christmas trees,and it has been widely used in orna-mental plantings.

The bark in the crown region ofmedium to large trees has conspicuousorange coloration. The lower bark of

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Scots Pine Circumference: 124'' Height: 60' Crown Spread: 45' Location: Falmouth

Scots pine, as the name suggests,is not native to Maine.

Page 39: Forest Trees of Maine

37S P R U C E

S The Important Distinctions

SP

RU

CE

Black SprucePicea mariana

Red SprucePicea rubens

White SprucePicea glauca

Norway SprucePicea abies

N E E D L E S

COLOR Blue-green Dark yellow-greenBlue-green to dark

greenDark green

LENGTH 1⁄4–1⁄2 inches 1⁄2–5⁄8 inches 1⁄2–3⁄4 inches 1⁄2–1 inch

DESCRIPTION Dull with waxy bloom Very shinyDull with waxy bloom,

strong, unpleasant odorwhen crushed

Shiny, sharp pointed

C O N E S

LENGTH 1⁄2–11⁄2 inches 11⁄4–2 inches 2 inches 4–7 inches

RETENTIONRemain on tree for

many yearsFall first year Fall first year Falls first year

SHAPE Spherical Wide in middle Cylindrical Cylindrical

SCALESStiff and rigid when

ripe; margin irregularlynotched

Stiff, with marginentirely or slightly

notched

Flexible at maturity,margin entire

Stiff, irregularly notched

T W I G S

COLOR Yellow-brown to brownReddish to orange-

brownLight gray to yellow-

brownOrangish-brown

HAIRSShort, rusty to black

hairs; some hairs tippedwith globose glands

Short, rusty to blackhairs; tips lack glands

Without hairsWithout hairs

(twigs droop from mainbranch)

Seed of all spruce is winged; cones are pendant; bare twigs are roughened by persistent leaf bases

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38 B L A C K S P R U C E

B S Picea mariana (P. Mill.) B. S. P.

In the past, spruce beer wasmade by boiling the branches ofthe black spruce.

Black spruce occurs statewide; itgrows on cool upland soils, but is

more commonly found along streams,on the borders of swamps and insphagnum bogs. It is also often foundon the sandy soils of eastern Maine. Itcan grow to a height of 50–70 feet anda diameter of 6–12 inches, but is nor-mally smaller than the maximum size.On a good site, it will grow rapidly. Insphagnum bogs, trees 50–80 years oldmay be only 6–8 feet tall and about oneinch in diameter. The branches areshort, pendulous and have a tendencyto curve up at the ends. It forms anopen, irregular crown. The lowerbranches often touch the ground, androot to form new trees. This methodof reproduction is known as “layering.”

The bark on the trunk is grayish-brown and the surface is broken intothin scales. The leaves are ¼–½ inch-es long, dull blue-green, blunt-point-ed, flexible and soft to the touch.

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39B L A C K S P R U C E

The cones, which usually stay onthe trees for many years, are ½–1½inches long, ovoid, and become nearlyspherical when open. The cone scalesare stiff and have toothed margins.

The twigs have many hairs, someof which are tipped with glands. Theinner bark is olive-green.

The wood is soft and light, butstrong. It is used for pulp, framing andconstruction lumber, and planking.Historically, spruce beer was made byboiling the branches.

Black spruce cones persist on the tree for manyyears. Look for clumps of old. gray, weatheredcones high in the tree.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Black Spruce Circumference: 47'' Height: 66' Crown Spread: 20' Location: Camden

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40 R E D S P R U C E

Red spruce is commonly foundthroughout the state. It grows on

well-drained, rocky upland soils, andparticularly on the north side of moun-tain slopes where it may be the majorspecies present. The spreading branch-es form a somewhat conical, narrowhead in young trees. The trunk is long,with a slight taper. It grows to consid-erable size, and is capable of attaining aheight of 60–80 feet and a diameter of1–2 feet, but occasionally exceeds thesemeasurements. Red spruce is shade-tolerant and will become established inthe understory of mixed stands.

The bark on mature trees is thickand is broken into thin, reddish-brownscales of irregular shape. The leaves aredark green, often with a yellow tinge,and are very shiny. They are about ½inch long, sharp-pointed, stiff, pricklyto the touch, and point toward the tipof the branch. The cones are oblongand usually 1½–2 inches long. Whenripe, they are reddish-brown and quiteshiny. The cone scales are stiff like the

R S Picea rubens Sarg.

Red spruce is one of our mostvaluable trees for the productionof building lumber.

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41R E D S P R U C E

black spruce, but the margins are gen-erally without conspicuous notches.The cones begin to drop in autumn orearly winter, and are all gone from thebranches by the next summer.

The twigs have hairs, none ofwhich have a gland at the tip. Theinner bark is reddish-brown. The woodis fairly soft, light, close-grained andstrong, but is not as durable as pinewhen exposed to the weather.

Red spruce is one of our mostvaluable trees for the production ofbuilding lumber. It is used for joists,sills, rafters, pilings, weir poles andheavy construction timbers. It is a prin-cipal wood used in the manufactureof paper pulp, and is valuable forthe sounding boards of musical instru-ments. Pitch for spruce gum is obtainedlargely from this tree.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Red Spruce Circumference: 103'' Height: 87' Crown Spread: 35' Location: T15 R9 WELS

Red spruce is the characteristic tree of the“Acadian forest” of northern New Englandand the Canadian Maritimes.

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42 W H I T E S P R U C E

W S Picea glauca (Moench) Voss

White or cat spruce occursstatewide except in York

county. It is widely distributed, but notas abundant as red spruce. It grows onshallow, rocky sites from the coast tothe tree line in the mountains, and isalso commonly found in old pasturesand on cleared land. It does not toler-ate shade and does not grow as anunderstory tree. The long and ratherthick branches, densely clothed withstout, rigid lateral branches, are curvedupward and form a somewhat open,irregular head with a broad base. Itcommonly grows to a height of 60–90feet and to a diameter of 2 feet.

The bark on old trees has lightgray, plate-like scales, which are thinand irregular, with a somewhat brown-ish surface. Younger trees have smooth,light gray bark.

The leaves point straight out fromthe branch. On the lower half of thetwig the leaves are often bent upwardin such a manner as to bring them allon the upper side. They are pale blue-

The wood of the white spruceis used for pulp, paddles, oars,piano sounding boards anddimension lumber, while itscones are used to makedecorative wreaths.

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43W H I T E S P R U C E

green at first, later becoming a darkblue-green. The foliage emits a pecu-liar and characteristic odor, which is aready means of distinguishing it fromthe other spruce species and is the rea-son for the alternate name.

The cones are slender, cylindrical,pale brown and shiny when ripe, andusually about 2 inches long. They ripenin August and September, and may becollected for seed until October. Conesusually fall off the first year. The conescales are thin and flexible, so that theygive easily when the cone is clasped inthe hand. The twigs are without hairs.The inner bark is silvery and glistens.

The wood is fairly light, soft,finishes well and is moderately strong.It is used for pulp, paddles, oars, pianosounding boards and dimension lumber,while its cones are used to make decora-tive wreaths. It shouldn’t, however, beused as a Christmas tree; when it isbrought indoors, the reason for itsnicknames—cat spruce and skunk

spruce—become evident. White andblack spruce produce long, tough,pliable roots which were used byAmerican Indians to tie togetherpieces of birch bark for canoes andother purposes.

White spruce cones are cylindrical and the scalescan be easily broken apart. This distinguishes itfrom red and black spruce, which have globe oregg-shaped cones with stiff scales.

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Page 46: Forest Trees of Maine

are ½–1 inch long, deep shiny green,four-sided in cross section and slightlyflattened. The needles lack the tenden-cy to bend upward on the twigs as inwhite spruce. Twigs are orangish-brown and without hairs.

In its native Europe, Norwayspruce is a very important lumber andpulpwood species. In Maine, the woodis primarily used for pulp and occa-sionally for lumber.

44 N O R W A Y S P R U C E

N S Picea abies (L.) Karst.

Anative of Europe, Norway spruceis of great economic importance

in its natural range. In Maine it is com-monly planted both in forest planta-tions and as an ornamental tree. Itrarely reproduces in the wild.

It is very symmetrical and gracefulin its growth habit; open-grown treesoften carry branches clear to theground. The tips of branches on largertrees have an upward sweep; and later-al branchlets are long and pendent.Norway spruce grows more rapidlythan any of our native species of spruce,and has been frequently planted forpulpwood, particularly in old fields inAroostook County. It is very suscepti-ble to attack by the white pine weevil.

The bark of younger trees is red-dish-brown; older trees have grayishbark with flaking scales. The cones arelarge, 4–7 inches long, and cylindricalwith stiff, notched scales. The leaves

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Norway Spruce Circumference: 143'' Height: 116' Crown Spread: 68' Location: Freeport

As its name implies, Norwayspruce is not native to Maine.

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45B L U E S P R U C E

Anative of the Rocky Mountainregion, blue spruce will grow on

a variety of sites and tolerate a widerange of growing conditions. Thesefactors, plus the striking color of itsfoliage, contribute to its popularity asan ornamental species, particularly inthe East where it is planted as a deco-rative tree. It does not readily becomenaturalized in Maine, and therefore isnot likely to be found growing in forestsettings. It can grow to be a large tree1–2 feet in diameter and to about 80feet in height. It is pyramidal in shape.

Foliage coloration varies from sil-very-blue to blue-green; the intensityof blue varies between individual spec-imens. The bark is gray to red-brownand scaly. The leaves are ½–1½ incheslong, stiff, very sharp-pointed, andstrongly incurved and covered with awaxy coating that gives the blue color.

B S Picea pungens Engelm.

Cones are light brown, oblong, 2½–4inches long, with thin, flexible, notchedscales. The twigs are stouter than theother spruces, hairless and tan.

Wild trees growing in the RockyMountains seldom have the intense col-oration of the cultivated varieties plant-ed here. Even in its native range, thewood is not often used commerciallybecause of its limited availability and itstendency to be brittle and full of knots.

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

Blue SpruceCircumference: 100''

Height: 91' Crown Spread: 27'Location: Auburn

Blue spruce is not native toMaine and is not likely to befound growing in forest settings.

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46 B A L S A M F I R

B F Abies balsamea (L.) P. Mill.

Balsam fir occurs statewide and isthe most abundant tree in the

state. It is frequently found in dampwoods and on well-drained hillsides,and often occurs in thickets. The treenormally forms a sharp spire to aheight of 60–70 feet and grows to12–20 inches in diameter. On youngtrees, the branches are horizontal, slen-der, and produced in regular whorls toform a strikingly symmetrical crown.In old age, the top is often slim, regu-lar and spire-like.

The bark on young trees is palegray, smooth, thin and has prominentblisters that are filled with a resinousliquid known as “Canada balsam.” Onold trees the bark gets rougher andblisters are absent.

The aromatic leaves are about1 inch long, dark green, and shinyabove with 2 rows of white stomatabelow. The tips are occasionallynotched. On branches in full sun, leavesturn up, but on lower branches theyspread out at right angles to the branch,giving it a flattened appearance.

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

Balsam FirCircumference: 78''

Height: 104' Crown Spread: 23'

Location: T4 R3 WELS

Balsam fir is the most abundant tree

in the state.

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47B A L S A M F I R

The cones are 2–4 inches long,erect and dark purple before maturity.Cones ripen in August and Septemberof the first year, and disintegrate short-ly thereafter, leaving only the centralspike-like stalks. The twigs are smoothafter the leaves have shed. Winter budsare covered with clear resin.

The wood is soft, light and moder-ately limber. It is sawed into dimensionlumber chiefly for light and mediumbuilding construction, and is usedextensively for pulp. Balsam fir isfavored for Christmas trees and greens.Each fall many tons of branch tips are collected for making Christmaswreaths. In the past, the branches weresteamed in a retort to produce oil ofbalsam. Also, the clear pitch formed inthe blisters of relatively young bark wasused to mount microscope slides and toattach theatrical costumes to bare skin.

The smooth bark with resin blisters distinguishesbalsam fir from the rest of our conifers.

Like all true firs, balsam fir conespoint upward and disintegratewhen they are mature.

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48 E A S T E R N H E M L O C K

E H Tsuga canadensis (L.) Carr.

Eastern hemlock is found in scat-tered stands in nearly every part

of the state. Best growth is attained onmoist, cool sites. It generally attains aheight of 60–70 feet, and a diameter of2–3 feet. The terminal shoot droopsand bends away from the prevailingwinds, quite often toward the east. Thetrunk usually tapers rapidly from thebase. This species can withstand con-siderable shading.

The bark is divided into narrow,rounded ridges covered with thickscales, and varies in color from cinna-mon-red to gray. Inner bark exposed bycuts or bruises shows a purplish tinge.

The leaves are flat, tapering,generally rounded at the apex, from⅓–⅔ inch long, with a distinct shortpetiole and so arranged that the twigappears flat. Leaves become progres-sively shorter towards the tip of thetwig. They are dark yellow-green witha lustrous upper surface, and a whitishundersurface.

The wood of the Eastern hemlock is used for

framing, sheathing, roof boards,timbers, bark mulch and pulp.

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49E A S T E R N H E M L O C K

The cones are about ¾ inch long,oblong, light brown, pendant and sus-pended on short, slender stalks. Conesmature during the first autumn andgenerally remain on the branches untilthe next spring. Seeds are winged andfall during the winter. The twigs arevery fine, limber and are not pitchy.

The wood is coarse, brittle whenvery dry, light, strong and difficult towork as it is likely to separate at one ormore of the annual growth rings. It isused for framing, sheathing, roof boards,timbers and pulp. The bark was oncevaluable for tanning but has been replacedby chemicals; now it is prized for its pur-ple color when made into mulch.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Eastern Hemlock Circumference: 127'' Height: 88' Crown Spread: 32' Location: North Yarmouth

When cut with a knife,Eastern hemlock barkwill show a purple color.

HE

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50T A M A R A C K

T Larix laricina (Du Roi) K. Koch

Tamarack, eastern larch or hack-matack is most commonly found

in cool, swampy places, although it alsogrows on well-drained soil. It is foundin scattered stands throughout thestate. It can grow rapidly and is not tol-erant of shade.

In the forest, the tree grows to aheight of 50–60 feet and a diameter of20 inches. It has a regular, narrow,pyramidal head with small, stiff hori-zontal branches.

In northern Maine, the name“juniper” is quite commonly applied tothis tree, but since juniper is the truename of another tree, its use for tama-rack is discouraged.

Tamarack is our only nativeconifer that sheds all its leaves

every fall.

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51T A M A R A C K

The bark separates on the surfaceinto small, thin, irregular reddish-brown scales.

The leaves are linear, about 1 inchlong, triangular in cross section, andborne in clusters of 8 or more on spurs,except on elongating new shoots,where they occur singly. They arebright green and turn a beautiful yellow just before they fall. Tamarackprovides some of the last color of thefall, as its needles turn color after mosttrees have already shed their leaves. Itis our only native conifer that sheds all its leaves every fall.

The cones are small, nearly spher-ical, about ¾ inch long, light brown,and borne erect on stout stems. Theyopen in fall to liberate the small

N AT I O N A L A N D M A I N E R E G I S T E R O F

B I G T R E E S 2 0 0 8Tamarack

Circumference: 143''Height: 92'

Crown Spread: 31' Location: T13 R8 WELS

winged seeds and usually remain onthe tree until the following year.

The wood is rather coarse-grained, hard, heavy and strong, withdurable heartwood. It is used forplanking, timbers, ties, poles, signposts,pilings and pulp. Historically, tamarackknees (the buttresses formed by largeroots) were used in shipbuilding.Tamarack was also used for mud sills inhome construction.

On older twigs, tamarack needles occur in clusters of up to 50 on short spur branches.

TA

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52 A T L A N T I C W H I T E C E D A R

Atlantic or coast white cedar isfound in bogs or low areas along

ponds or streams. It has a scattered dis-tribution from the mid-coast south. InMaine it rarely reaches a height of over40 feet. The short branches come outfrom a gradually tapering trunk, givingthe tree a conical appearance. Thetwigs are only slightly flattened.

The bark is fibrous, grayish to red-dish-brown, often with twisted spirals; onyoung trees it is easily pulled off in strips.

The leaves are bluish-green, scale-like, and arranged in somewhat fan-shaped clusters. When crushed, theygive off an aroma.

The cones are small, round,smooth and purplish before maturity,about ¼ inch in diameter with tack-like scales. They persist through thewinter, but are inconspicuous.

While Atlantic white cedar wood is of limited use,

small trees are cut for fenceposts and shavings are used for

pet bedding.

A W CChamaecyparis thyoides (L.) B. S. P.

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53A T L A N T I C W H I T E C E D A R

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Atlantic White Cedar Circumference: 68'' Height: 63' Crown Spread: 19' Location: Alfred

Atlantic white cedar is rare in Maine andoccurs only in a few isolated bogs in the southand mid-coast.

The wood is light, close-grained,strongly fragrant, and light browntinged with red. It is brittle and there-fore of limited use, though small treesare cut for fenceposts. The shavings areused for pet bedding.

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54 N O R T H E R N W H I T E C E D A R

N W C Thuja occidentalis L. Northern white cedar or eastern

arborvitae is generally found inswamps, along streams, on mountainslopes and in old pastures where thesoil is moist. Dense stands are widelydistributed statewide. It is most abun-dant in the northern and eastern sec-tions, and grows best on alkaline soils.It is widely used as an ornamental. Thehead is compact, narrow and pyrami-dal. The branches are horizontal, shortand turned upward. Trees grow to 60feet in height and to 3 feet in diameter.The trunk is often strongly buttressed.

The bark has shallow fissures,which divide it into flat narrow ridges.It is reddish-brown and often tingedwith orange.

The leaves are opposite or two-ranked, usually only about ⅛ inchlong, scale-like, blunt, and so arrangedas to make the small branches flat in

Cedar has emerged as a viablealternative to pressure-treated wood.

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55N O R T H E R N W H I T E C E D A R

shape. They have a pleasant aroma anda rather pleasing taste, and are a majorsource of food for deer in the winter.

The cones are erect, small, about½ inch long, with only a few pairs ofscales. They mature in one season. Theseed is small and winged.

The wood is soft and light, coarse-grained, brittle, has very durable heart-wood and a fragrant odor. It is usedprimarily for shingles, slack cooperage(barrels for dry, semi-dry or solid prod-ucts), poles, posts and rustic fencing;and it is sawed into lumber for hopechests (since the wood is said to repelmoths), siding, canoes and boats. Morerecently, cedar has emerged as a viablealternative to pressure-treated wood.Naturally weather-resistant, it is usedfor decks, post and rail fencing, out-door furniture, roof shakes, and peltstretchers.

Northern white cedar cones are about 1⁄2 inch long and often occur in large numbers.

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56 E A S T E R N R E D C E D A R

E R Juniperus virginiana L.

Eastern redcedar is not common inMaine. It grows on poor soils,

gravelly slopes, rocky ridges and onmoist, sandy ground. It is found inter-mittently in southern Maine and inBridgton, Porter, Denmark and WestGardiner. It gets the name “redcedar”from the color of the heartwood.

It is variable in its habit. Youngtrees have slender horizontal branchesand a narrow, compact, conical head.The crown of old trees becomes broadand rounded. In Maine, trees attain adiameter of 8–12 inches, and a heightof 30 feet.In Maine, Eastern redcedar is

not sufficiently plentiful to be ofcommercial importance.

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57E A S T E R N R E D C E D A R

The bark on the trunk is lightbrown, tinged with red; it separatesinto long, narrow shreds on old trees.

The leaves are scale-like, overlap-ping, about 1⁄16 inch long, dark green,and remain on the tree 5–6 years, grow-ing hard and woody the third season.Branchlets appear square in cross section.Current growth and vigorous shootscontain sharp-pointed, awl-shaped leaves—the so-called “juvenile” growth.

The fruit is berry-like, globose,with 1–2 seeds, pale green at first, darkblue when ripe, and is about the sizeof a small pea.

The wood is brittle, fine-grained,light, easily worked, durable, and veryaromatic. The heartwood is a dull red.It is valuable for fence posts and panel-ing for moth-proof closets, but inMaine it is not sufficiently plentiful tobe of commercial importance. Theshavings are used as bedding for pets.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Eastern Redcedar Circumference: 62'' Height: 53' Crown Spread: 38' Location: Hebron

Eastern redcedar invades old pasturelandand quickly dies out when other treesbegin to shade it.

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58 C O M M O N J U N I P E R

C J Juniperus communis L.

Common juniper is found prima-rily as a shrub in pastures and

open spaces on shallow, rocky soil. Itoccurs infrequently, primarily in thesouthern half of the state. It is occa-sionally found as a tree. Specimens upto 25 feet in height have been record-ed, but are extremely rare.

The bark is grayish-brown andoccurs in thin, longitudinal, shreddedlayers. The inner portion has a reddishtinge. The leaves occur in whorls ofthree. They are sharp, stiff, dagger-likeand persist for several seasons. They are¼–¾ inch in length. The upper surfaceis concave and marked with a broad,white line. The underside, which due

Common juniper is usually foundas a shrub rather than as a tree.

to the bending of the twigs usuallyappears uppermost, is dark green.

The fruit is dark blue, coveredwith a thin bloom and is slightly small-er than a pea. Fruits remain on thetrees during the winter, and have astrong resinous taste. The fruit is usu-ally found only on select trees sincemale and female flowers are generallyproduced on separate trees. This trait iscommon to most junipers.

The wood is hard, close-grainedand very durable.The heartwood is lightbrown. Large stems make long-lastingfence posts if the bark is removed.

Juniper shavings can be used forpet bedding. In Europe, the fruits areused to make gin.

Page 61: Forest Trees of Maine

Horses were stillcommonly used tohaul logs in theMaine woods until the 1950s.

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60

B

Often referred to as “hardwoods”, broadleaf trees belong to the group of

plants known as the angiosperms or flowering plants. Almost all of Maine’s

hardwood species are deciduous, meaning they loose all of their leaves

each autumn and grow new ones in the spring. Maine has over 50 species

of native hardwood trees; only about half of these are considered important

timber trees. The name “hardwood” is somewhat misleading since some

hardwood species have wood that is soft. Photo location: T1 R6 WELS

Page 63: Forest Trees of Maine

Quaking AspenPopulus tremuloides

Bigtooth AspenPopulus grandidentata

Balsam PopularPopulus balsamifera

B A R K

TEXTURESmooth in younger trees, often

with horizontal bands of circularwart-like outgrowths

Smooth in young trees; furrowed in older trees

Smooth or roughed by dark outgrowths; older trees furrowed

with scaly ridges

COLOR Light or grayish-green Dark or olive green Reddish-brown on younger trees

TASTE Very bitter Not bitter Not bitter

L E A V E S

LENGTH 11⁄2–3 inches 3–4 inches 3–5 inches

SHAPE Circular Broad egg-shaped Egg-shaped

MARGIN Finely toothed Coarsely toothed Finely toothed

SURFACEShiny upper,

not rusty beneathNot shiny upper,

not rusty beneathVery shiny upper,

rusty beneath

PETIOLE Flattened Flattened Flattened

B U D S

TEXTURE Not sticky; shiny Not sticky; dull Very sticky; shiny

SHAPE Conical Broad egg-shaped Egg-shaped

SCALES No hairs Covered with white hairs No hairs

ODOR Not fragrant Not fragrant Sweet balsam fragrance

PO

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P O P L A R S & A S P E N S

P A The Important Distinctions

61

The pith of poplar twigs is star-shaped in cross section. Poplars belong to the willow family and resemble willows in flower andfruit characteristics. The nodding, “woolly bear” caterpillar-like staminate and pistillate catkins are borne on different trees.They open before the leaves are out and are conspicuous in the early spring. Poplars, like willows, have a transcontinental range.They can be propagated very easily from cuttings.

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62 Q U A K I N G A S P E N

Quaking aspen, popple or trem-bling aspen is found statewide

and is an abundant, rapid-growing treeoccurring in either pure stands or inmixture with other species. It is foundon many different kinds of soil, butmakes the best growth on sandy, moistsoils. Frequently it is the first species,with paper birch, to become estab-lished following heavy cuttings orburns. Intolerant of shade, it does notpersist in dense woods. It is a gracefultree with slender branches that are farapart and often contorted. It has around and narrow head. It grows to aheight of 60–75 feet and a diameter of10–16 inches.

The bark is smooth, often rough-ened by horizontal lines of wart-likeoutgrowths. It is a pale green with darkbrown patches. The pale green areasfeel waxy when rubbed. Bark on oldtrees is ash gray and dark at the basewhere it is divided into broad, flatridges. It has a very bitter taste similarto quinine.

Q A Populus tremuloides Michx.

In the past, the quakingaspen was ground up and cooked for cattle feed.

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63Q U A K I N G A S P E N

The leaves are alternate, roundedand short-pointed, with finely roundedteeth; dark green and shiny above and1½–3 inches long. The flattened peti-ole causes the leaves to tremble in abreeze, resulting in a rustling sound.

The flowers are in catkins thatappear before the leaves. The fruit,which ripens about June, is a capsule.The seeds are very small, light and cot-tony, and are carried long distances bythe wind. The buds are dark brown,have a varnished appearance and maybe slightly sticky. Flower buds are usu-ally larger than the leaf buds.

The wood is close-grained, softand rots very easily. It is used increas-ingly for trim, lumber, pallets, and for themanufacture of oriented strand board,landscape ties, plywood, core stock andexpendable turnery items. It is usedextensively for pulp. In the past, it wasground up and cooked for cattle feed.Sometimes referred to as “biscuit wood,”it was also used as firewood for cooking.

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Above left: young bark.Above right: old bark.Left: Quaking aspen buds are dark brownand very shiny.

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

Quaking AspenCircumference: 43''

Height: 69' Crown Spread: 21'

Location: Richmond

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64 B I G T O O T H A S P E N

B A Populus grandidentata Michx.

Bigtooth aspen, poplar or poppleoccurs statewide and commonly

grows with quaking aspen. It is a rapidgrower in various soils and in differentsituations. It grows best in a rich, sandyand fairly moist soil. It is more shade-tolerant, and therefore more competi-tive, than quaking aspen and growswith other species in either scatteredor small groves. Bigtooth aspen tendsto have better form than quakingaspen. It attains a height of 60–80 feetand a diameter of 10–20 inches.

When first emerging in spring, the bigtooth aspen leafis a distinctive silvery-green.

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65B I G T O O T H A S P E N

The bark is smooth, and olive togray-green. At the base of old trees, itis dark and divided into broad, irregu-lar, flat ridges.

The leaves are alternate, 3–4 inch-es long, broadly egg-shaped in outline,and have a dark green upper surface.When first emerging in spring, they area distinctive silvery-green. The edgesare coarsely and irregularly toothed.The petiole, or leaf stalk, is flat.

The flowers are in catkins, andappear before the leaves.

The fruit ripens in May about thetime the leaves begin to come out. Theseeds are small, light and are carried longdistances by the wind.The buds are dullgray, slightly hairy and not sticky.

The wood is like that of the quakingaspen and is used for the same purposes,as well as rails for apple-picking ladders.

N AT I O N A L A N D M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

Bigtooth Aspen Circumference: 151'' Height: 76' Crown Spread: 45' Location: Appleton

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66 B A L S A M P O P L A R

B P Populus balsamifera L.

Balsam poplar, or Balm-of-Gilead,inhabits the borders of swamps

and the low bottomlands along riversthroughout the state, except in YorkCounty. It gets its name from the fra-grance of the resinous, sticky buds.

The tree is somewhat different fromthe two preceding poplars. The branch-es are stout, erect, more or less contortedat the ends and form an open, rather nar-row head. It reaches a height of 30–70feet, and a diameter of 15–30 inches.

The bark on young trees issmooth, or sometimes roughened bydark outgrowths, and is greenish toreddish-brown. On the trunk of oldtrees, it is gray and separated intobroad, rough ridges.

The balsam poplar gets its name from the fragrance

of the resinous, sticky buds.

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67B A L S A M P O P L A R

The leaves are alternate, ovate, 3–5inches long and 2–3 inches wide. Theyare deep dark green and shiny on theupper surface, light green and usuallywith rusty blotches on the under side.The edges are lined closely with small,rounded teeth. The petioles are roundin cross section. In late summer theentire tree can have a rusty appearance.

The flowers are in catkins that appearearly in spring just before the leaves.

The fruit ripens the end of May orearly in June. Each seed is attached to acottony mass, so that it is often carriedlong distances by the wind.

The wood is somewhat like that ofquaking and bigtooth aspen, but it isnot as strong. The wood is prone todecay while growing. Larger logs aresawed into landscaping ties. OSB—ori-ented strand board, a structurally engi-neered wood product—can include asmall percentage of balsam popular.

Balsam poplar has largesticky buds that have asweet fragrance.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Balsam Poplar Circumference: 99'' Height: 93' Crown Spread: 50' Location: Yarmouth

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68 W I L L O W S • B L A C K W I L L O W

B W Salix nigra Marsh.

Black willow occurs primarily insouthern and western Maine. It

grows to a height of 45–65 feet, and isfound along streams and ponds. Thestout, upright, spreading branches givethe tree a broad, irregular outline. It is probably our largest native willow.The bark on old trees is shaggy anddark brown. The leaves are very nar-row, sometimes sickle-shaped, finely-toothed, 3–6 inches long and green onboth sides. The wood is soft, light,weak and is used occasionally for farmlumber and pulp.

W I L L O W S Salix spp. L.

Maine has many willows, but this is a large

and difficult group to identify. The Revised

Checklist of the Vascular Plants of Maine,

1995 (see Appendix Four, p. 105) shows 58

native and exotic species, varieties, and

hybrids known to be present in the state.

Maine’s willows range in size from large

trees to small prostrate shrubs found in the

alpine tundra. With the exception of black

willow, most of Maine’s native willows are

small trees or shrubs. Some of the exotic

species can grow to be very large. All wil-

lows share the following characteristics:

Buds are covered with a single, cap-like

scale with silky, gray hairs beneath the

scale. Leaves are alternate, mostly narrow,

and the petioles are short or lacking. Flowers

occur in catkins. Fruits consist of small,

usually two-valved capsules filled with silky

hairs that are attached to the seeds.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Black Willow Circumference: 266'' Height: 84'

Crown Spread: 64' Location: Yarmouth

WI

LL

OW

Top right: A typicalwillow twig.

Page 71: Forest Trees of Maine

Much of Maine’saccessible, large whitepine had been cut bythe 1850s. After that,spruce, seen here,became an increasinglyimportant timber tree.

Page 72: Forest Trees of Maine

Red MapleAcer rubrum

Sugar MapleAcer saccharum

Silver MapleAcer saccharinum

B A R K

TEXTUREOlder trees: ridged and broken

into plate like scales. Young trees: smooth.

Older trees: rough and deeply furrowed. Young trees: smooth

and slightly fissured.

Older trees: somewhat furrowed,separates into thin plates.

Young trees: smooth.

COLORDark gray on older trunks, light gray on young trees

Gray on older trunks, light gray on young trees

Dark gray on older trunks, gray on young trees

L E A V E S

LOBES/SHAPE3–5 lobes. sides of terminal lobeconverge, notches between lobes

V-shaped

3–5 lobes, sinuous, sides of terminal lobes flare outward,

notches between lobes rounded

5 lobes, rarely 3; lobes long andnarrow like fingers on a hand

MARGIN Irregularly doubly toothed Sparsely toothed Irregularly and sharply toothed

SURFACE Under-surface slightly white Under-surface pale green Under-surface silvery white

F L O W E R

APPEARANCEScarlet or yellow-red, appears

before the leavesGreenish-yellow, appears with

the leavesGreenish-yellow or pinkish,

appears long before the leaves

B U D S

LENGTH Terminal bud—1⁄8 inch Terminal bud—1⁄4 inch Terminal bud—1⁄8 inch

SHAPEBlunt-pointed, as long

as broadSharp-pointed, many

scales showingBlunt-pointed, slightly ridged

COLOR Dark red Purplish-brown to gray Bright red above, green below

F R U I T

SHAPE Paired, slightly divergent Paired and slightly divergentPaired, but with

one usually abortive

SEED BODY Oval in outline Round Football-shaped

WING Reddish; 3⁄4 inch long 1 inch longStrongly divergent,

2 inches long and hooked

70 M A P L E S

MA

PL

EM * The Important Distinctions

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71M A P L E S

Striped MapleAcer pensylvanicum

Mountain MapleAcer spicatum

Norway MapleAcer platanoides

B A R K

TEXTUREMarked with whitish stripesrunning lengthwise on trunk

Smooth when young; shallowlyfurrowed when older

Smooth when young; regularlyfurrowed on older trees

COLOR Reddish-brown or dark green Reddish-brown to grayGray when young,

gray-brown on older trees

L E A V E S

LOBES/SHAPE3 lobes, shaped;

like a duck’s foot, thinUsually 3 lobes, sometimes 5

5–7 lobes, blade wider thantall, stems exude a milky sap

when broken

MARGIN Edges finely and sharply-toothed Coarsely-toothed Sparsely toothed

SURFACEUnder-surface pale green;

pubescent Prominently sunken veins on

the upper surfaceVery dark green; some cultivars

red or deep purple

F L O W E R

APPEARANCEBright yellow, appears after

leaves are full grownYellow-green in long clusters

after the leaves are full grownYellow-green, appear before

the leaves

B U D S

LENGTH Terminal bud—1⁄2 inch Terminal bud—1⁄4 inch >1⁄4 inch

SHAPEDistinctly stalked with

2 scales showingSlender and pointed,

slightly stalkedTurban-shaped, blunt-pointed,

large scales

COLOR Bright red Green to red Green to purple

F R U I T

SHAPE Paired and moderately divergentPaired, slightly divergent,

ascending clustersPaired, flattened

SEED BODYLarge smooth depression in

seed bodyWrinkled depression on

seed bodyFlattened

WING Reddish-brown; 3⁄4 inch long Slightly divergent; 1⁄2 inch longStrongly divergent, leathery;

2 inches long

MA

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*Key does not include boxelder. Boxelder (page 84–85) is the only maple in Maine with compound leaves.

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72 R E D M A P L E

R M Acer rubrum L.

Red maple—also known as soft,white or swamp maple—occurs

throughout the state. A rapid growerand the most abundant of the maples,it is typically found in swamps andpoorly drained sites, but also occurselsewhere. The red maple is a medium-sized, slender tree that becomes 50–60feet high, and 1–2 feet in diameter.The branches are upright, forming asomewhat narrow head. Usually thetrunk is not divided.

The bark on young trees is smoothand light gray. On old trunks, it is darkgray, ridged and broken into plate-like scales.

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73R E D M A P L E

The leaves are opposite, 3–5 inch-es long, with 3–5 lobes and marginsthat are irregularly double-toothed.The upper surface is light green; lowersurface is white. The sides of the termi-nal lobe converge toward the tip; andthe notches between lobes are V-shaped. In fall, the leaves turn scarletand orange.

The flowers are produced in clus-ters on stalks before leaf buds open.Males are yellowish-red while femalesare bright scarlet. The red maple is oneof the first trees to flower in spring.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8

Red Maple Circumference: 183'' Height: 69'Crown Spread: 67' Location: Richmond

Red maples produce bright red flowers followed by abundant seeds in the springtime.

The fruit is winged, ripens inspring or early summer, and germi-nates as soon as it falls. Wings are onlyslightly divergent, about ¾ inch long.The seed body lacks a depression.

The twigs are straight, stiff, do nothave a rank odor when broken, and arered on both surfaces. Buds are red andoften clustered.

The wood is close-grained, heavy,moderately strong, easily worked butnot durable, although it will take agood polish. It is used mainly for pulpand firewood, but also for pallets, fur-niture stock, canoe paddles and turneryproducts. As sugar maple becomesmore expensive, more mills are usingred maple. It is also commonly used forlandscape plantings.

MA

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74

Sugar, rock or hard maple is foundabundantly throughout the state

on moist, rocky slopes, but grows beston moist, upland soils. In the forest, itgrows to 60–70 feet and a diameter of20–30 inches. The top is short andspreading. In the open, the branchesbegin 8–10 feet up, forming an egg-shaped head when the tree is young anda broad, rounded top when older. Itmakes an attractive street or ornamen-tal tree, but it is sensitive to road salt.Maple sugar and syrup are made largelyfrom the sap of this tree, although sugaris present in the sap of all maples.

S M Acer saccharum Marsh.

Historically, sugar maple was used to make parts for sleighs, sleds,pungs (low, one-horse box sleighs) and buggy shafts.

S U G A R M A P L E

Page 77: Forest Trees of Maine

S U G A R M A P L E 75

Bark on young trees and largebranches is smooth or slightly fissuredand pale. Some trees have oval light-colored blotches on the bark. Oldertrees are deeply furrowed and light to darker gray. Leaves are opposite,with 3–5 lobes, sparingly-toothed, 3–5inches long, dark green above, palegreen below. Sides of the terminal lobeare parallel or divergent; and notchesbetween lobes are u-shaped. In autumn,leaves turn various shades of red, scar-let, orange or yellow.

Flowers are greenish-yellow, pen-dulous, appear on long, slender, hairystalks and in clusters, with the leaves.The fruit is paired, round with wingsthat are about 1 inch long and slightlydivergent. It ripens in the fall. Thetwigs are brown with sharp-pointedbrown buds.

The wood is heavy, close-grained,strong and hard. It is used for furniture,flooring, tool handles, veneer, railroadties, novelties, dowels, woodenware,canoe paddles, firewood and pulp.“Birds-eye” and curly-patterned mapleis in high demand in the furniture andveneer industry. Historically, sugarmaple was used to make parts forsleighs, sleds, pungs (low, one-horsebox sleighs) and buggy shafts.

Sugar maple buds are sharp-pointed and have scales thathave a dark margin.

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

Sugar MapleCircumference: 213''

Height: 80' Crown Spread: 64'Location: Palermo

MA

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76 S I L V E R M A P L E

S M Acer saccharinum L.

Abundant in some localities, silvermaple is a common tree, found

throughout the state except along thecoast. It grows largely on sandy banksalong streams, usually attaining aheight of 60–80 feet and a diameter of2–3 feet. The trunk normally separatesinto 3 or 4 upright secondary stems,devoid of branches for some distance.The branches are long and slender,often pendulous.

The bark on young trees issmooth, gray, slightly tinged with red.On old trees, it is reddish-brown, fur-rowed, and separated into large thinscales that are loose at the bottom.Twigsare chestnut brown and shiny.

The leaves are opposite, deeplyfive-lobed; and the edges are irregularand sharply toothed. The upper surfaceis pale green, the lower, silvery white.They turn a pale yellow in fall.

In Maine, silver maple is mostcommon along major rivers.

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77S I L V E R M A P L E

The flowers are on very shortstalks and in clusters. They are green-ish-yellow or sometimes pinkish, open-ing early, long before the leaves appear.

The fruit is paired, winged andripens in spring. Frequently, one of thepair does not fully develop. The twigsare curved upward at the tip, orange orred-brown above and green below,slender, with a bitter taste and a rankodor when broken.

The wood is softer than that of the hard maple, close-grained, notdurable and easily worked. It is used toa limited extent for pulp.

Silver maple has large globe-shaped flowerbuds and smaller vegetative buds.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Silver Maple Circumference: 316'' Height: 89'

Crown Spread: 75' Location: Leeds

MA

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Page 80: Forest Trees of Maine

78 S T R I P E D M A P L E

S M Acer pensylvanicum L.

Striped maple or moosewood iscommon throughout the state. It

is a shade-loving tree that is foundgrowing with other hardwoods, oroccasionally with conifers, on rich,moist soils or rocky slopes. Of littlevalue except for its beauty, it rarelyexceeds a height of 25 feet and a diam-eter of 8 inches. The branches are slen-der and upright, and the top narrowand often short.

The bark on the trunk is reddish-brown or dark green, and marked by whitish lines running lengthwise,which turn brown after a time. Theleaves are three-lobed toward the apex,resembling a goose foot, opposite, fine-ly toothed, pale green, 5–6 inches longand about as broad. In fall they turnlight yellow.The striped maple is a shade-

loving tree that is usually foundgrowing with other hardwoods.

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79S T R I P E D M A P L E

The flowers are bright yellow inslender drooping racemes that openthe end of May or early June, when theleaves are fully grown. The fruit ispaired, with wings moderately diver-gent, fully grown in late summer. It hasa smooth, oval depression in the seedbody. The twigs are smooth, reddish orgreenish; the buds are valve-like, stout,stalked and without hairs.

The wood is close-grained, lightand soft. During spring when the cam-bium layer is active, it is easy to make awhistle from the smaller branch sections.

The white and green-striped bark of thestriped maple distinguishes it from any othernative tree.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Striped Maple*

Circumference: 30"/31" Height: 45'/50' Crown Spread: 21'/20'

Location: Harpswell /Monhegan Island *Tie

MA

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80 M O U N T A I N M A P L E

M M Acer spicatum Lam.

Mountain maple occurs through-out Maine and is especially

common in the northern part of thestate. It grows as a small bushy tree, sel-dom over 30 feet in height. At times,the tree forms fairly dense thickets, dueto its habit of growing in clumps. Itgrows best in a wet habitat or on damp,northern slopes. The slender twigsgrow in a somewhat upright position.

The mountain maple growsas a small bushy tree, seldomover 30 feet in height.

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81M O U N T A I N M A P L E

The bark is reddish-brown togray, thin and somewhat furrowed.

The leaves are opposite, three-lobed, shiny above, somewhat hairybelow. They have rather coarse teethand prominently sunken veins on theupper surface.

The flowers appear in June inlong, hairy, yellow-green clusters afterthe leaves are full grown.

The fruit is paired, with wings

slightly divergent, and occurs inascending clusters. It has a wrinkleddepression on the seed body and ripensin early fall.

The twigs are hairy, green, red orreddish-brown, not striped; and thepith is brown. The buds are hairy,valve-like, green, and only slightlystalked, slender and pointed.

The wood is close-grained, soft,light and not used commercially.

Facing page, far left: Mountain maple leaveshave deeply impressed veins on the upper surface.

MA

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82 N O R W A Y M A P L E

N M Acer platanoides L.

Norway maple is not native toMaine. Because of its aggressivenature, it is considered to bea serious potential threat toour native flora and furtherplanting of it is discouraged.

Native to continental Europe,Norway maple thrives in a wide

variety of conditions, grows rapidly andcasts a deep shade. Because of its aes-thetic appeal and ease of propagation,it has been planted across Maine as astreet and shade tree. It has escapedinto the wild around many of our citiesand towns, particularly in the southernhalf of the state. Because of its aggres-sive nature, Norway maple is consid-ered to be a serious potential threat toour native flora and further planting ofit is discouraged.

The bark of young trees is grayand smooth. Bark of older trees is gray-brown to almost black, and broken intolong, interlacing vertical furrows.

Norway maple drawing by Anna Anisko, used with thepermission of the Pennsylvania Flora Project, MorrisArboretum of the University of Pennsylvania.

Page 85: Forest Trees of Maine

83N O R W A Y M A P L E

The leaves are opposite, with 5–7lobes, and very dark green. The leafblade is usually broader than it is long.Early in summer, the petiole will exudea white milky sap when broken. Thischaracteristic is not shared by any ofour native maples.

The flowers are bright yellow-greenand appear in spring before the leaves.

The fruits are paired and diverge ata wide angle from each other. Each fruithas a leathery wing attached. The fruitand wing are about 2 inches long, flat-tened, light brown and mature in the fall.

The twigs are stout and brownish.The buds are large and green to purplewith large bud scales.

The wood is occasionally used forfirewood.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Norway Maple Circumference: 166'' Height: 70' Crown Spread: 86' Location: South Berwick

Norway maple fruithas a flattened seedbody and a leathery,2 inch wing.

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84 B O X E L D E R

Boxelder, or ashleaf maple, isapparently not native to Maine,

but has been planted as an ornamentaltree throughout the state and hasescaped in localized areas near habita-tion. It was introduced along the St.John River in Aroostook County. Itreaches a maximum height of 50 feetand diameter of about 2 feet in Maine.It is a short-lived, fast-growing, brittletree, prone to wind and ice damage. Itcan become invasive.

B Acer negundo L.

Boxelder is a short-lived,fast-growing, brittle tree,prone to wind and ice damage.It can become invasive.

BO

XE

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ER

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85B O X E L D E R

The bark is light gray and smoothon young stems, becoming roughenedand shallow-fissured on older trees.

The leaves are opposite, compound,usually 3–7 leaflets per leaf, rarely nine.The leaflets vary greatly in shape, oftenlobed and unlobed leaflets are found onthe same leaf. Leaflets are occasionallydivided into individual blades.

The flowers open just before theleaves in the spring and are yellow-green. They have no petals.

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

BoxelderCircumference: 115''

Height: 85' Crown Spread: 66'Location: Wilton

Boxelder twigs arepurple and coveredwith a whitishwaxy bloom.

MA

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The fruit attains mature size insummer, ripening in autumn. It con-sists of a double-winged pair of seeds.Wings are only slightly divergent; andthe seed body is wrinkled, three timeslonger than broad.

The twigs are smooth, ratherstout, green or maroon, and coveredwith a white, chalky bloom. The barkyields a pungent odor when bruised.

The wood is light, soft, creamywhite, often tinged with green, weakand close-grained. Occasionally it isused for pulp.

Page 88: Forest Trees of Maine

Paper BirchBetula papyrifera

Gray BirchBetula populifolia

Yellow BirchBetula alleghaniensis

B A R K

TEXTURESeparates into thin, horizontal,

papery layersDoes not separate into papery

layersSeparates into thin, horizontal,

ribbon-like strips

COLOROuter, chalky or grayish white;

inner bark orangeOuter, chalky or grayish-white,dirty-looking; inner bark orange

Bright silvery gray or light yellow

ODOR No odor No odorWintergreen odor when young

branches are scraped

L E A V E S

LENGTH 2–4 inches 21⁄2–3 inches 3–41⁄2 inches

OUTLINE Egg-shaped Triangular Egg-shaped

MARGIN Doubly toothed Coarsely and doubly toothed Coarsely and doubly toothed

SHAPE Tip short pointed; base rounded Tip long pointed; base truncated Base unevenly rounded

SURFACE Upper dark green, dull Upper dark green and glossy Upper dark green, dull and hairy

F L O W E R

STRUCTURE 3 catkins Single or paired catkins 3–4 catkins

ARRANGEMENT Clustered Not clustered Not clustered

B U D S

TEXTURE Sticky when squeezed Not sticky Smooth

SHAPE Long, tapered Short, globose Long, sharp-pointed

SCALES Without hairs Without hairs Hairy

COLOR Reddish-brown Red-brown to greenish-brown Reddish-brown

T W I G S

TEXTURE Hairy, with spur shootsVery fine, warty but not hairy,

without spur shootsSomewhat hairy, with spur shoots

COLOR Depends on age Dull gray or brown Greenish or yellow-brown

ODOR No wintergreen odor No wintergreen odor Slight wintergreen odor

86 B I R C H E S

B The Important Distinctions

BI

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Page 89: Forest Trees of Maine

87B I R C H E S

Sweet BirchBetula lenta

Mountain Paper BirchBetula cordifolia

B A R K

TEXTURESmooth on young trees;

broken into irregular plates on older treesSmall portions peal away in thin sheets

COLOR Dark to almost blackWhitish with a pink to salmon- colored tinged to reddish-brown

ODORWintergreen odor when young branches

are scrapedNo odor

L E A V E S

LENGTH 3–5 inches 2–4 inches

OUTLINE Egg-shaped Egg-shaped

MARGIN Singly and sharply toothed Doubly toothed

SHAPE Base heart-shaped Tip short pointed; base heart-shaped

SURFACEUpper dark green, dull; lower light yellow-green

Upper dull green

F L O W E R

STRUCTURE 3–4 catkins 2–4 catkins

ARRANGEMENT Not clustered Clustered

B U D S

TEXTURE Smooth Sticky when squeezed

SHAPE Long, sharp-pointed Long, tapered

SCALES Without hairs Without hairs

COLOR Chestnut brown Brown

T W I G S

TEXTURE Smooth with spur shoots, no hairsSparsely hairy, often warty,

with spur shoots

COLOR Reddish-brown Yellowish-brown to dark brown

ODOR Strong wintergreen odor No wintergreen odor

BI

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Page 90: Forest Trees of Maine

88 P A P E R B I R C H

P B Betula papyrifera Marsh.

Paper, white or canoe birch is acommon tree in all parts of the

state; it occurs in pure stands or in mix-ture with other species. It reaches60–70 feet in height and 1–2 feet indiameter. It grows along streams andon the borders of lakes and ponds,thriving best in a rich, moist soil.

When young, the branches areshort, slender, spreading, somewhatdrooping, and form a narrow, regularhead. In the forest, the trunk is freefrom branches well up from theground; and the tree forms an open,narrow and round-topped head.

The bark is a protective layer andshould never be removed from livingtrees. On the trunk and limbs, it sepa-rates freely and easily into thin, paperysheets. The outer surface is white, theinner part bright orange. Seedlings or

Paper birch is used to make toothpicks

and golf tees.

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89P A P E R B I R C H

very young trees have a darker coloredbark, which gradually changes to acreamy-white.

The leaves are alternate, ovate,short-pointed, 2 –4 inches long, thick-er than those of gray birch, doubly-toothed, with the upper surface darkgreen and dull.

The flowers are in catkins. Theyopen in early spring before the leaves.Those appearing in fall are dormant,staminate catkins and occur mostly inclusters of three.

The twigs are usually hairy and,unlike yellow birch, without a winter-green taste. The buds are slightly sticky.

The wood is close-grained, mod-erately hard, and strong. It is used for woodenware, flatware and turnedproducts including toys, dowels, furni-ture parts, pulp and firewood.

The tree gets the name of “paperbirch” from how the bark was used by early settlers, and that of “canoebirch” because the bark was used to

make canoes. In the early spring paperbirch sap contains considerable sugar.Historically paper birch was one of themost valuable tree species in Maine. Inthe past, the wood was used to makeshoe pegs (used instead of nails in themanufacture of shoes) as well as anumber of products that used to bemade in Maine, but are now manufac-tured off-shore. These include clothes-pins, yarn spools, toothpicks, paper rollplugs and plywood.

Paper birch bark will peel off inlarge sheets, but it should neverbe removed from living trees.

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

Paper BirchCircumference: 148"

Height: 72' Crown Spread: 22'

Location: Alton

BI

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90 G R A Y B I R C H

G B Betula populifolia Marsh.

Gray birch is a short-lived and not particu-larly valuable tree. It occurs to some

extent statewide, but is only abundant in thesouthern and eastern sections of the state. It isfrequently found in old fields, burns and heavi-ly-cut areas. This is a small tree that commonlyreaches 20–30 feet in height and 4–8 inches in

Gray birch is a short-lived and notparticularly valuable tree that is usedprimarily for pulp and firewood.

Page 93: Forest Trees of Maine

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

Gray BirchCircumference: 71"

Height: 65' Crown Spread: 27'

Location: Richmond

91G R A Y B I R C H

diameter. It usually occurs in clumpsand often leans.The branches are short,slender, frequently pendulous and con-torted, and bend toward the groundwhen the tree is not crowded. The headis long, narrow, pointed and open.

The bark is close and firm, anddoes not easily separate into thin lay-ers. The outer part is dull grayish-whiteor chalky. The inner portion is orange.

The leaves are 2½–3 inches inlength, thin, long-pointed, triangular,alternate and doubly toothed. Theupper surface is dark green and glossy.The slightest breeze causes them toflutter like those of the poplars, hencethe scientific name Betula populifoliawhich means “birch with poplar leaves.”

The flowers are produced incatkins. They open in early springbefore the leaves. Those that appear infall are male and usually solitary.

The twigs are the most slender ofour native hardwoods. They are toughand wiry, dull gray or brown, hairless,and have a rough, warty surface. Deadtwigs tend to stay attached to the trunk.This, plus the dirty appearance of thebark, makes this tree easy to recognize.

The wood is light, soft, oftencoarse-grained, and decays rapidlywhen exposed. It is occasionally usedfor pulp and firewood; in the past itwas used for paper roll plugs.

Gray birch has single or paired catkins inwinter and spring.

BI

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92 Y E L L O W B I R C H

Y B Betula alleghaniensis Britt.

Yellow birch is the largest of thenative birches, growing to a

diameter of 3 feet and a height of70–85 feet. The spreading branches aresomewhat pendulous; they form abroad, round-topped head in the open,but an irregular head in the woods. Itgrows well statewide on cool, moistsites, and is frequently mixed with beechand sugar maple, or with hemlock.

The bark on the branches and onthe stems of young trees is very shiny,silvery-gray or yellowish-brown, sepa-rating into loose, thin, horizontal, oftenribbon-like layers. On old trees, it isdivided into large thin plates and is dullgray or black.

The leaves are 3–4½ inches long,ovate or nearly oblong, alternate; theedges are doubly toothed, the upper

The yellow birch is one of ourmost valuable timber trees andmakes excellent firewood.

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93Y E L L O W B I R C H

side dull, dark green and hairy. Leavesclosely resemble those of easternhophornbeam.

The flowers are in catkins. In win-ter there are 3–4 pre-formed staminatecatkins on the shoots, but not in clus-ters. They open in the early spring.

The twigs are yellowish to darkbrown and somewhat hairy. The youngtwigs are aromatic like sweet birch,although to a lesser degree. Both thebuds and twigs have a pronouncedwintergreen taste.

The wood is hard, strong, heavyand will take a good polish. It is close-grained and evenly textured. The

heartwood, which makes up the bulkof the wood, has a pleasing reddishcolor; this is why it is sometimes calledred birch. It takes stains easily, makesexcellent veneer wood, and does noteasily warp. It is also used for furniture,flooring, woodenware, lumber forinterior finish, plywood, railroad ties,pallets, pulp, gunstocks and dowels.The yellow birch is one of our mostvaluable timber trees and makes excel-lent firewood. As with sweet birch,wintergreen oil wasformerly distilled fromtwigs and branches.

Yellow birch’s bark peels in small curls. Very old trees have platy bark.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Yellow Birch Circumference: 200" Height: 48' Crown Spread: 91'

Location: Deer Isle

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94 S W E E T B I R C H

S B Betula lenta L.

Sweet, black or cherry birch isfound, though uncommonly, in

the southern third of the state. Itinhabits the banks of streams or moist,rich upland soil. It is a handsome treewith a tall dark stem, and spreading,slender, horizontal branches that arependulous at the ends. It has a graceful,open, narrow head, which in full sunbecomes round and symmetrical. Itgrows to a height of 60–70 feet and adiameter of 1–2 feet.

The bark on the trunk of old treesis dark to almost black, and separatesinto large, thick, irregular plates. Onyoung trees and branches, it is smooth,shiny, dark brown tinged with red, aro-matic, and has a very pronounced win-tergreen flavor.

The leaves are alternate, 3–5 inch-es long, aromatic, ovate or somewhat

The name “cherry birch” is applied to this tree because of theresemblance of the bark on old trunks to that of the black cherry.

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95S W E E T B I R C H

oblong, and sharply toothed. Theupper surface is dark green and dull;the lower surface is light yellow-green.

The flowers are produced incatkins. The winter shoots support 3to 4 staminate catkins. They open justbefore the leaves unfold in the spring.

The wood is hard, heavy, strongand can be beautifully polished. It isprized for use in the manufacture offurniture and it makes excellent fire-wood. Limited amounts are used aspulpwood. Historically, oil with somemedicinal value was obtained from thebranches and bark by distillation, andwas generally known as wintergreen oil.

The name “cherry birch” is appliedto this tree because of the resemblanceof the bark on old trunks to that of theblack cherry.

Sweet birchtwigs havesmall spurbranches andtaste likewintergreen.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Sweet Birch Circumference: 139" Height: 82'

Crown Spread: 48' Location: Gorham

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96 M O U N T A I N P A P E R B I R C H

M P B Betula cordifolia Regel.

Mountain paper birch is closelyrelated to paper birch, and has

been designated as a variety of thespecies by some authors (Betula papy-ifera var. cordifolia (Regel) Fern.). It isknown at many points in Maine, par-ticularly on mountain slopes, coastalheadlands and islands east of MountDesert Island. It often grows as aclump of several stems. It can grow toabout 60 feet in height and 1 foot ormore in diameter.

The bark of young trees andbranches is dark reddish-brown anddoes not peel. The bark of older treeswill separate into thin, papery layers.In mature trees, bark color ranges fromwhitish with a pinkish or salmon-col-ored tinge to reddish-brown or bronze.

The leaves are egg-shaped withheart-shaped bases, abruptly pointed,and coarsely doubly-toothed. Theflowers are borne in catkins. Dormantmale catkins in clusters of 2–4 arevisible during winter. Both male andfemale catkins expand in spring.

The fruit is a tiny nutlet with 2small lateral wings. It matures in fall andis dispersed during the fall and winter.Large numbers of birch seed can oftenbe seen on the surface of the snow.

The wood is similar to that ofpaper birch; and the two species are usu-ally not separated. It is used for turneryproducts, cabinetry, pulp and fuel.

Mountain paper birch is most often foundalong the coast and at high elevations.

Page 99: Forest Trees of Maine

Maine’s trees have beenused to make hundredsof different products,many of which are nowobsolete, including elmhub stock, shown herein an early photo fromCumberland, Maine.

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98 E A S T E R N H O P H O R N B E A M

E H Ostrya virginiana (P. Mill.) K. Koch

Eastern hophornbeam or ironwoodis a small tree with either an open

or rounded crown. It reaches a height of20–30 feet and a diameter of 6–10 inch-es. The branches are long and slender,with ends that are somewhat drooping.

It is a fairly rapid grower, especiallyin good soil. It grows on slopes andridges having a dry, gravelly soil, and isoften found in the shade of other species.

The bark is gray, and separates easilyinto thin, narrow, vertical scales, becom-ing finer and stringy on older trees.

The leaves are 2–3 inches long,egg-shaped to nearly oblong in outline,widest in the middle, hairy on both sur-faces, alternate and sharply toothed.Theyare somewhat like those of yellow birch.

The flowers occur in catkins, whichopen with the leaf buds. The malecatkins are pre-formed in the fall andare usually in clusters of three.

The fruit is bladder-like, enclosesa ribbed nutlet and occurs in clusters.It ripens in September. The name

The name “hophornbeam” refersto the fruit, which closelyresembles the true hops.

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99E A S T E R N H O P H O R N B E A M

“hophornbeam” refers to the fruit,which closely resembles the true hops.

The twigs are light brown, fine, toughand wiry, and have a small green pith.

The wood is very close-grained,heavy, very strong and is exceedinglyhard when seasoned. It is used for toolhandles, wedges for directional fellingof trees, and firewood. In the past, itwas used to make trip stakes on loghauling trucks (which contain, thenrelease the logs from the truck beds),wagon tongues (the shaft where thehorses are hitched) and other parts.

Eastern hophornbeambuds have tiny verticalgrooves that can be seenwith a magnifying glass.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Eastern Hophornbeam*

Circumference: 77"/70" Height: 63'/67' Crown Spread: 38'/42'

Location: Livermore Falls/Pownall*Tie

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100 A M E R I C A N H O R N B E A M

A HCarpinus caroliniana Walt.

American hornbeam, blue-beechor musclewood occurs west of

the Penobscot River in the southernthird of the state. The American horn-beam is most commonly found inhab-iting wet woods and the borders ofswamps and streams. It is a small,slow-growing tree 10–25 feet tall and4–10 inches in diameter. The branchesare crooked. The trunk is characteristi-cally ridged, or fluted longitudinally.

The bark is smooth and grayish-blue. The leaves are alternate, egg-shaped or oval, 2–3 inches long,sharply toothed, smooth above andhairy below. They turn a brilliant scar-let in autumn.

Close-grained, strong, toughand durable, Americanhornbeam wood is used forlevers, handles and wedges.

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101A M E R I C A N H O R N B E A M

The flowers are produced in catkinsthat open in spring before the leaves.

The fruit is a ribbed nutlet, whichis attached to the base of a three-lobedbract, and is borne in open clusters.

The twigs are reddish-brown, slen-der and tough. Buds are also reddish-brown and slender, and sharp-pointed.

The wood is close-grained, com-pact, strong, tough and durable. It isused for levers, handles, and wedges.

American hornbeam bark has asinewed appearance that gives it itsalternate name, “musclewood.”

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102

Speckled alder is very common inMaine, usually growing in wet

areas along brooks, in swamps and inpastures. It sprouts readily and is a nui-sance on pasture land. Alder usuallyoccurs as a shrub, rarely as a small tree.It is seldom more than 4 inches indiameter and 20 feet in height.

The bark is smooth, dark chocolatebrown, and marked with white, hori-zontal, elongated spots called lenticels.

The leaves are alternate, 2–3 inch-es long, usually broadly ovate; and thetexture is rough or rugose as the scien-tific name implies. The edges areunevenly or doubly-toothed.

The flowers are in catkins, and openbefore the leaves in spring.The purplish,wax-like male catkins are pre-formed theprevious fall. The fruit is woody andcone-like, with a very short stalk.

The winter buds are short-stalkedand maroon, with few scales showing.

The twigs are reddish-brown; the pithis triangular in cross section.

The wood is light and soft, andhas very little commercial use. Thewood discolors very rapidly on expo-sure to air. Baskets for the florist indus-try are made from small diameterstems. In the past, the wood was usedin hand forges, because of the intenseheat it produces when burned.

Two other species, green ormountain alder—Alnus viridis (Vill.)Lam. & DC. Spp. Crispa (Ait.) Tirrill)—and hazel alder (Alnus serrulata(Ait.) Willd.) occur as shrubs.

S AAlnus incana ssp. Rugosa (Du Roi) Clausen

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Page 105: Forest Trees of Maine

Mechanical equipment,including the Lombard log hauler, began to replace horses in the Maine woods in the early 1900s.

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104 A M E R I C A N B E E C H

A B Fagus grandifolia Ehrh.

American beech occurs statewide,growing up to 70 feet in height

and 1–3 feet in diameter. Although itgrows best on rich upland soil, beech iscommon and sometimes forms nearlypure stands, with shoots often spring-ing up from the trees’ roots. Beech barkdisease causes significant mortality inMaine. The disease results when bark,attacked and altered by the beech scaleinsect, Cryptococcus fagisuga is invadedand killed by fungi, primarily Nectriacoccinea var. faginata.

The bark is light gray and smoothunless affected by beech bark disease.The bark of trees affected by the dis-ease is rough and pockmarked withsmall cankers.

American beech nuts aresweet and are an importantfood source for wildlife.

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105A M E R I C A N B E E C H

The leaves are alternate, 3–5 inch-es long, elliptic, acutely pointed, withcoarse and hooked teeth. The marginbetween the teeth is nearly straight.Dead leaves are light tan and tend toremain on trees into winter.

The fruit consists of a bur, whichusually contains 2 triangular ediblenuts. These nuts are sweet and are animportant food source for wildlife.Trees that bears have climbed to eatbeech nuts show claw marks on thebark. The winter buds are long, slen-der, many scaled, and sharp-pointed.

The wood is strong, hard andtough, but not durable. Current usesinclude for pulp, pallet stock and fire-wood. In the past, it was used forclothespins, furniture, handles, wooden-ware, railroad ties, dowels and flooring.

European beech (Fagus sylvaticaL.), Purple (Fagus sylvatica var. atrop-unicea Weston) and Copper beech(Fagus sylvatica var. cuprea L.) are speciesof European origin planted in southernand central Maine as ornamentals.

Smooth barked beech (left) have become a rarityin Maine. The bark of most trees (right) isroughened by cankers caused by beech bark disease.

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

American BeechCircumference: 104"

Height: 87' Crown Spread: 58'Location: Hallowell

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Page 108: Forest Trees of Maine

Northern Red Oak

Quercus rubra

Black OakQuercus velutina

Scarlet OakQuercus coccinea

Bear OakQuercus ilicifolia

Black oak group: leaves sharp-tipped, acorns mature in two years and are hairy inside.

B A R K

TEXTURE Slightly ridgedDeeply fissured blocky,

ridges very darkRidges small and

irregularSmooth, with a few raised

lenticels

COLORDark gray to black;

inner reddishBlack; inner—deep

orange or bright yellowDark gray to black;

inner—pale red or grayGray to dark brown

L E A V E S

LENGTH 5–8 inches 5–6 inches 3–6 inches 3–6 inches

COLORSurface—dull, dark

green; below—yellow-green

Surface—dark green and shiny

Surface—bright greenand shiny; below—paler

Surface—dark green;below—white or gray

SURFACE Smooth beneath Hairy beneath Smooth beneath Hairy beneath

A C O R N S

SIZE2–4 times longer

than cupTwice as long as cup Twice as long as cup

Small, nearly hemispherical,striped above middle

CUPSaucer-like,

with scales fusedBowl-like, with dull scales

Bowl-like,with shiny scales

Shallow

B U D S

SHAPE Conical, smooth Large, decidedly angled Small, rounded Small, short, blunt-pointed

COLOR Chestnut brown Yellowish-gray Light brown Reddish-brown

SCALES Silky at tipCoated with matted

wool-like hairsHairy at tip only Loose scales

O A K S

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B O The Important Distinctions

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107

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O A K S

W O The Important Distinctions

White OakQuercus alba

Chestnut OakQuercus prinus

Bur OakQuercus macrocarpa

Swamp White OakQuercus bicolor

White oak group: leaf lobes rounded, acorns mature in one year and lack hairs inside.

B A R K

TEXTURE Ridges broad, flat, flaky Deeply furrowed Deeply furrowed, flakyDeeply fissured; broad,

flat ridges, flaky

COLOR Light grayReddish-brown to

dark brownGrayish

Grayish-brown,inner bark orange

L E A V E S

LENGTH 4–7 inches 4–8 inches 6-12 inches 4–6 inches

COLOR Surface—bright green Surface—yellow green Surface—dark green Surface—dark green

SURFACE Upper—dull Upper—shiny Upper—shiny Upper—dull

SHAPELobes rounded; cleft to midrib

Narrowly elliptical; shallow rounded lobes

Violin-shaped; lobes rounded

Slightly lobed

A C O R N S

SIZE Very fineThree times as long ascup; long and wrinkled

Twice as long as cupThree times

as long as cup

CUP Short-stalkedHairy, moderately

long-stalked

Margin fringed with long,hair-like scales, short

stalked

Margins fringed withscales, long-stalked

BB UU DD SS

SHAPE2–4 times longer

than cupBroadly ovoid, sharp-pointed

Broad ovoid, blunt or sharp-pointed

Roundish, blunt-pointed

COLOR Dark red-brown Yellowish-brown Reddish-brown Brown

SCALES Without hairs Without hairs Coated with soft hairs Small, without hairs

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108 N O R T H E R N R E D O A K

N R O Quercus rubra L.

Northern red oak is the mostcommon oak species in Maine.

It occurs state-wide but is most abun-dant in the southern half of the state.Best growth is attained on rich uplandsoils. It grows to a height of 60–80 feetand a diameter of 2–3 feet, forming eithera narrow or broad head. The branchesare stout, horizontal or upright.

The bark on the trunks of oldtrees is dark gray or nearly black, and isdivided into rounded ridges. Onyounger trees and branches, it issmooth and greenish-brown or gray.The inner bark is reddish. The leavesvary in shape, are 5–8 inches long,alternate, are dull, dark green above

Northern red oak is the most

common species ofoak in Maine.

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109N O R T H E R N R E D O A K

and yellow-green below, and have bris-tle-tipped lobes. Some dead leaves mayremained attached in winter.

The flowers appear in May, whenthe leaves are only partly grown. Thefruit ripens the second year. The acornis broad, large, 1–1¼ inches long, andup to four times longer than the shal-low cup. Red oak acorns are a majorsource of food for many species ofwildlife. The inside lining of the acornis densely hairy. Its tannic acid contentmakes it bitter. The twigs are smooth,greenish to reddish-brown, and have astar-shaped pith.

The wood is hard, strong and rel-atively heavy. It is used for furniture,interior finish, planks and frames, lob-ster trap runners, flooring, piling,cross-ties, timbers, pallets, dowels andfirewood. Historically, it was used forshipbuilding (ribs, beams and timbers),weir poles (some 60 feet long), as kilnwood and fence posts (when split).

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Northern Red Oak Circumference: 265'' Height: 84' Crown Spread: 71' Location: Vienna

Northern red oak oftenhas a reddish colorationin the bark fissures.

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110 B L A C K O A K

B O Quercus velutina Lam.

Black oak is used to a limited extent for interior finish, shipbuilding,flooring, piling, cross-ties, timbers, pallets, dowels, and firewood.

Black or yellow oak is found insouthern Maine from Lincoln

and southern Oxford counties south-ward; it is common near Fryeburg.It grows on dry ridges and graveluplands. The branches are slender; andthe head is narrow and open. It growsto a height of 50–60 feet and a diame-ter of 1–2 feet.

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111B L A C K O A K

The bark is smooth and dark grayor brown on young stems. On oldtrees, it is divided by deep fissures intobroad, rounded ridges and is dark,almost black. It is rougher than redoak. The inner bark, which is charac-teristically bright orange or bright yel-low, was used in tanning.

The leaves are alternate, 5–6inches long, varying much in shape andgeneral outline, usually seven-lobedwith bristle points. The upper surfaceis glossy and dark green; the under sur-face is generally hairy with more obvi-ous, rusty hairs in axils of veins.

The flowers appear in May whenthe leaves are only partly grown.

The fruit, which is bitter, maturesthe second season. The acorn is ½–¾inch long, almost twice as long as thecup and one-half enclosed by it.

The twigs are smooth; buds aredensely hairy, angled and yellowish-gray.

The wood is hard, heavy, strongand coarse-grained. It is used for thesame purposes as scarlet oak

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Black Oak Circumference: 181'' Height: 73' Crown Spread: 73' Location: Fryeburg

Black oak buds andtwigs are coveredwith tan fine hairs.

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112 S C A R L E T O A K

S O Quercus coccinea Muenchh.

Scarlet oak occurs rarely in thesouthern tip of Maine. It grows on

the dry soil of ridges and uplands inYork, Cumberland and Androscoggincounties.

In Maine, scarlet oak is a smallertree than red oak, growing to a heightof 30–50 feet and 1–2 feet in diameter.The branches are slender and form anopen, narrow head.

In Maine, scarlet oak is a smaller tree than red oak.

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113S C A R L E T O A K

The bark on the trunks of oldtrees is separated into irregular ridgesby shallow fissures, and is dark graywith a reddish inner bark.

The leaves are alternate, 3–6 inch-es long with a variable outline. Theupper surface is bright green and shiny;the lower is paler and less shiny. Lobesare sharp-tipped. In fall, the leaves turna deep scarlet, which accounts for thecommon name of the tree.

The flowers appear in May whenthe leaves are only partly developed.

The fruit ripens the second year.The acorn is about ½ inch long, abouttwice as long as the cup, and is fromone-third to one-half enclosed by thecup. It is quite bitter.

The wood is hard, strong andheavy, but coarse-grained. It is used toa limited extent for interior finish,shipbuilding, planks and frames, floor-ing, piling, cross-ties, timbers, pallets,dowels, and firewood.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Scarlet Oak Circumference: 73'' Height: 73' Crown Spread: 40' Location: Yarmouth

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Scarlet oak acorns havesmall rings around the tip.

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114 B E A R O A K

B O Quercus ilicifolia Wangenh.

Bear oak wood is occasionally used as fuel, but isgenerally not considered to be of commercial value.

Bear or scrub oak is a small, thick-et-forming, shrubby tree usually

less than 20 feet high. It is common onthe sand barrens of southern Maine,extending into Oxford County whereit is common on the barrens surround-ing the Saco River near Fryeburg. Italso occurs in eastern Hancock County,and on rocky ridges and barren ledgesites along the coast.

The bark is smooth, gray-brown,and has a few raised lenticels. On larg-er trees the bark may become roughand scaly.

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115B E A R O A K

The leaves are the primary distin-guishing feature. They have 5–9 bris-tle-tipped lobes, are whitened on theunderside, alternate and 2–4 incheslong. The second set of lobes from thebase tends to be much larger than others.

Male flowers are pale reddish-green catkins that appear in May whenthe leaves are only partially grown. Thefruit is a dark brown acorn 2⁄5–4⁄5 inchlong that matures in the fall of the sec-ond season.

The twigs are slender and denselyhairy during the first year. The termi-nal buds are clustered, chestnut brownand blunt-pointed. The lateral buds areof similar size to the terminal.

The wood is occasionally used asfuel, but is generally not considered tobe of commercial value.

Bear oak twigs are finerthan the other native oaksand covered with hairs.

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116 W H I T E O A K

White oak, which gets its namefrom the color of the bark,

occurs naturally in southern and cen-tral Maine as far north as Oakland innorthern Kennebec County. It growson sandy land, gravelly ridges andmoist bottomland, but makes the bestgrowth on rich, heavy, upland soils. Ingood situations, it attains a height of60–70 feet and a diameter of 3–4 feet.When not crowded by other trees, the

W O Quercus alba L.

White oak wood is strong, heavy, hard and durable, makingit ideal for use in flooring, furniture and boatbuilding.

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117W H I T E O A K

bole (trunk) is short, the limbs are largeand diverging, and the head is broadand rounded. In the forest, it has a longbole and a narrow head.

The bark on the trunk is separat-ed into thin, irregular flakes and variesfrom light to ashy-gray.

The leaves are usually nine-lobed.The lobes are rounded, slightly cleft orcleft nearly to the midrib, alternate, 4–7inches long, bright green above, palegreen or whitish beneath. They some-times remain on the tree during winter.

The flowers come out in Maywhen the leaves are half grown.

The fruit ripens in September ofthe first year. The acorn is about ¾inch long, 2–4 times longer than thecup, and about one-quarter enclosed byit. The fruit is edible. AmericanIndians pounded it into a flour andbleached out the tannin with hot water.

White oak acorns mature in one year. Theycontain less tannin than red oak acorns andare preferred by wildlife.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8White Oak Circumference: 239'' Height: 64’ Crown Spread: 95' Location: Pittston

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The twigs are gray to purple; buds areblunt-pointed, and scales are withouthairs.

The wood is strong, heavy, hardand durable. It is used for ship andboatbuilding, railroad ties, piling, agri-cultural implements, interior finish,furniture, flooring, pulp, and firewood.In the past, it was used for deck plank-ing on ships, tight cooperage (tightcasks capable of holding liquid such aswhiskey), and spokes and rims ofwooden wheels.

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118 C H E S T N U T O A K

C O Quercus prinus L.

Chestnut oak only occurs in thesouthern tip of Maine. It is

found on Mt. Agamenticus in thetown of York and has been reportedfrom Oxford County. In Maine, treesgrow 12 inches or more in diameterand about 40 feet in height.

The gray-brown bark is smoothon young trees, but becomes thick andvery deeply furrowed on older trees.The leaves are similar to those of theAmerican chestnut. They are yellow-

Due to chestnut oak’s rarity in Maine,it is not used commercially here.

Page 121: Forest Trees of Maine

green above, hairy below, narrowlyelliptical with shallow rounded lobeswithout bristle tips. They are oftenwidest above the middle.

Male flowers are yellow-green,borne on catkins and appear in May.Female flowers are reddish, borne inspikes with the leaves in mid-spring.The edible fruit is a large, 1½ inchlong, ellipsoid acorn that matures inone season. Its cup encloses about halfof the acorn. The twigs are hairless andorange-brown to gray. The chestnutbrown buds are clustered toward theend of the twig, pointed, and quitelong and narrow in shape.

119C H E S T N U T O A K

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Chestnut Oak Circumference: 191'' Height: 90' Crown Spread: 72' Location: Yarmouth

They gray bark of the chestnut oakis very deeply furrowed.

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The wood is similar in characterto white oak and has similar uses.In areas where chestnut oak is moreabundant, it is sold as white oak. Dueto its rarity in Maine, it is not usedcommercially here.

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120 B U R O A K

B O Quercus macrocarpa Michx.

Bur oak is found in the southerntwo-thirds of the state and is

locally plentiful in central Maine. It isquite common along the SebasticookRiver, the lower Penobscot basin, andeast into Hancock County. It grows inlow, rich bottomland, and is rarelyfound on dry soil. It has a broad top ofwide, spreading branches. The trunk isoften clear of limbs for two-thirds ormore of its length. It attains a height of60–70 feet, and a diameter of 2–3 feet.

Very durable, hard, heavy and strong, bur oak is used forfor cabinetry, barrels, hardwood flooring and fence posts.

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121B U R O A K

The bark is grayish, deeply fur-rowed and broken into plate-like irreg-ular scales.

The leaves are roughly violin-shaped in outline, with rounded lobesthat are not generally as deeply cut asthe white oak.The upper end of the leafis widest. They are alternate, dark greenand shiny on the upper surface, palegreen or silvery-white on the lower.

The flowers appear in May whenthe leaves are partly formed.

The fruit, which is edible, maturesthe first year and is usually solitary. Itvaries in size and shape. The acorn isabout ¾ inch long, and about halfenclosed by the cup. The margin of thecup is fringed with long, hair-like scales.The twigs have corky wings or ridges.

The wood is very durable, hard,heavy and strong. It is used for thesame purposes as white oak.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Bur Oak Circumference: 176'' Height: 86' Crown Spread: 74' Location: West Gardiner

Bur oak twigs have ridges of corky bark,a characteristic that is not shared by anyof our other native oaks.

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122 S W A M P W H I T E O A K

S W O Quercus bicolor Willd.

Swamp white oak is not abundant,but occurs in small, widely scat-

tered groves in York and Androscoggincounties. It grows in moist, fertile soilon the borders of swamps and alongstreams.

Swamp white oak grows to aheight of about 50 feet and a diameterof 2–3 feet. The limbs are small, usual-ly pendulous. The head is narrow, openand round-topped.

The bark on old trees is deeplyfurrowed, divided into broad, flatridges, flaky and grayish-brown. On

Swamp white oak is not abundant, but occurs in small,widely scattered groves.

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123S W A M P W H I T E O A K

young trees and branches, it is smoothand separates into papery scales thathang loosely. The inner bark is orange.

The leaves are alternate, 4–6 inch-es long and slightly lobed. The uppersurface is dark green and shiny; thelower, pale white or tawny.

The flowers appear in May whenthe leaves are not more than half-grown.

The fruit matures the first season.The acorn has a long stalk, is about 1inch long, three times as long as the cupand about one-third enclosed by it.

The twigs have a yellowish or alight orange to reddish-brown bark.

The wood is strong, heavy, hardand used for the same purposes as thewhite oak.

M A I N E R E G I S T E RO F B I G T R E E S

2 0 0 8Swamp White Oak*

Circumference:122''/126''

Height: 74'/75'Crown Spread: 65'/60'

Location:Wiscasset/Paris Hill

*Tie

Swamp white oak twigs have small roundedbuds. The bark of the twigs and small branchestends to peel and flake.

OA

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A R E A O FO C C U R R E N C E

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124 A M E R I C A N C H E S T N U T

A C Castanea dentata (Marsh.) Borkh.

Most of the chestnut trees inMaine have been destroyed by

the chestnut blight fungus, Endothiaparasitica. There are national effortsunderway to develop blight-resistanttrees of this once very important treespecies.

There are national efforts underwayto develop blight-resistant trees of thisonce very important tree species.

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125A M E R I C A N C H E S T N U T

The natural range of Americanchestnut only extended into southernand central portions of the state.Chestnut now occurs infrequently, usu-ally as sprout growth, in the southernhalf of the state on rich, well-drainedsoil. It has been planted occasionally asfar north as Orono. The tree grows rap-idly. In the forest, it has a tall, straighttrunk free of limbs, and a small head.When not crowded, the trunk dividesinto 3 or 4 limbs and forms a low, broadtop. It reaches a height of 60–70 feetand a diameter of 15–30 inches.

The bark on the trunks of oldtrees is dark brown and divided intobroad, flat ridges by shallow, irregularfissures. On young stems, it is smoothand dark gray with a green tinge.

The leaves are coarsely-toothedand hooked, with the leaf marginrounded between the teeth. Leaves arealternate, 5–8 inches long, yellow-green and smooth on both surfaces.

The fruit is a prickly bur contain-ing 2–3 nuts tipped with hairs. Theinner lining of the bur is plush-like.

The fruit of American chestnut is contained ina very prickly bur.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8American Chestnut Circumference: 117'' Height: 45' Crown Spread: 44' Location: Orono

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The nuts contain a sweet meat; theywere once gathered in large quantitiesfor the market.

The wood is soft, very durable,strong and splits easily. It is used forinterior finishing and was once inmuch demand—prior to the blight—for telephone poles, railroad ties,beams and timbers up to 50 feet inlength, furniture stock and fence posts.The durability of the wood is due tothe tannic acid that it contains.

Page 128: Forest Trees of Maine

To keep logs sorted as theywere transported to mills,each company choppeda brand into its logs.A Great Northern PaperCompany brand can beseen in the foreground.

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127A S H E S

White AshFraxinus americana

Green Ash*Fraxinus pennsylvanica

Black AshFraxinus nigra

L E A V E S

LEAFLETS 5–9, usually 7 7–9 7–11

DESCRIPTIONLeaflets are mostly entire, borneon stalks, without hairs below.

Turn purple in autumn

Leaflets borne on stalks. Hairy below and on rachis. Turn

yellow or bronze in autumn.

Toothed leaflets which are without stalks except the one at theend. Hairs lacking below except forbuff-colored hairs at the junction of

the leaflets and the rachis. Turn yellow in autumn.

B U D S

SIZE 1⁄8 inch 1⁄8 inch Less than 1⁄4 inch

SHAPE Blunt-pointed Cone-shaped Sharply-pointed

COLOR Brown Brown with rusty or dull red hairs Black or very dark

F R U I T

WINGS Wing terminalSeed body grading gradually

into wingFlat, completely surrounds

seed body

SEED BODY Cigar-shaped Funnel-shapedSlightly twisted, less than half

the length of the fruit

T W I G S

TEXTURESmooth and shiny, often with

slight bloom, very brittleSomewhat covered with

downy hairsSmooth, not shiny

COLORGray or greenish-brown,

inner bark bright brick redGreenish-gray, inner bark

cinnamon-coloredPale gray,

inner bark dirty white

AS

H

A The Important Distinctions

*Specimens of green ash which lack hairs on the twigs or leaflets, but otherwise fit the above description, were formally designated as var. lanceolata. They are now designated under the species due to the many gradations of the hairiness character.

Page 130: Forest Trees of Maine

White ash isone of Maine’svaluable timber trees and isfound commonlythroughout the state.

128 W H I T E A S H

W A Fraxinus americana L.

White ash is one of Maine’svaluable timber trees and is

found commonly throughout the state.Best growth occurs on rich, rathermoist soil of low hills. It grows to aheight of 60–70 feet and a diameter of15–30 inches. The branches areupright or spreading, forming a narrowtop in the forest.

The bark pattern resembles awoven basket; it is broken into broad,parallel ridges by deep furrows, and isdark brown or deep gray.

The leaves are opposite, 8–12inches long and consist of 5–9 (usually7) leaflets. The leaflets are 3–5 incheslong, oval to lance-shape, borne onshort stalks, edges remotely toothedtowards the tip, dark green and oftenshiny on the upper surface. In fall, theyturn to a soft, velvety purple.

The fruit is a single samara occur-ring in clusters. The seed body is cigar-shaped and has a terminal wing.

Page 131: Forest Trees of Maine

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

White AshCircumference: 244'’

Height: 95'Crown Spread: 70'

Location: South Waterford

129W H I T E A S H

The twigs have a smooth, shinybark which is grayish, greenish ormaroon on the surface. The inner layerof the bark is brick red. The terminalbuds are rounded or dome-shaped.

The wood is hard, strong andtough. It is used for agricultural imple-ments, tool handles, oars, furniture, inte-rior finish, dowels, pulp and firewood,and sporting goods including baseballbats, hockey sticks and snowshoe frames.

White ash twigs arehairless and have deeplynotched leaf scars.

AS

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A R E A O FO C C U R R E N C E

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130 G R E E N A S H

G A Fraxinus pennsylvanica Marsh.

Green or red ash occurs overmuch of the state, particularly

along the major rivers. It is not asabundant as the white and black ash,but is fairly common in central Maine.Sometimes mistaken for black ash, itgrows near the banks of streams andlakes on rich, moist soil. It has stoutbranches that bend downward on oldertrees and form an irregular, compacthead in the forest. It seldom exceeds aheight of 50–60 feet and a diameter of16–20 inches.

The quality of green ash woodis not as good as white ash.

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131G R E E N A S H

The bark on the trunk of old treesis dark gray or brown, and firm andfurrowed like that of the white ash.

The leaves are 10–12 inches long,opposite, with 7–9 leaflets borne perstalk. Leaflets are 4–6 inches long,entire or wavy, or sometimes toothed,particularly on the upper-half of theleaflets, yellow-green on the upper sur-face, hairy below and on the rachis, andoval to elliptical.

The fruit has a funnel-shaped seedbody gradually blending into the ter-minal wing.

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

Green AshCircumference: 115''

Height: 63'Crown Spread: 65'

Location: Mechanic Falls

The current year twigs are green-ish-gray and covered with numeroushairs, although sometimes there are nohairs. Inner bark is cinnamon red.

The wood is hard, heavy, fairlystrong, coarse-grained and brittle. It isused in the same ways as white ash.

Green ash twigs are oftenhairy and do not havedeeply notched leaf scars.

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A R E A O FO C C U R R E N C E

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132 B L A C K A S H

Black ash wood is used for interior finishing, cabinet work, basketsand, to a limited extent, pulp.

B A Fraxinus nigra Marsh.

Black or brown ash occurs statewide.It grows almost entirely on rich,

moist ground or in cold, wet swampsand along the banks of streams.

It is a tall, slender tree with ashort, narrow head. It grows to aheight of 50–60 feet and a diameter of10–20 inches. The trunk is often with-out branches for a considerable dis-tance from the ground.

The bark is gray to dark gray,corky and spongy, with more or lessparallel ridges. It rubs off freely withthe hand.

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133B L A C K A S H

The leaves are 12–15 inches long,opposite, and have 7–11 leaflets thatare 4–5 inches long, and without stalksexcept the one at the tip. Leaflets arelance-shape and have remotely-toothedmargins. The upper surface is darkgreen. There are buff-colored hairs atthe junction of the leaflets and rachis.

The fruit is a single samara occur-ring in clusters. The seed is flattenedand completely surrounded by the wing.

The twigs are smooth, gray toolive-green. The buds are black orbrown and pointed at the tip. Theinner layer of the bark is dirty white.

The wood is coarse-grained,heavy, tough, durable and pliable. It isused for interior finishing, cabinetwork, baskets and, to a limited extent,pulp. It the past it was used to makebarrel hoops.

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

Black AshCircumference: 139''

Height: 82'Crown Spread: 56'Location: Waterboro

The wing of black ash fruit completelysurrounds the seed body.

AS

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A R E A O FO C C U R R E N C E

Page 136: Forest Trees of Maine

134 A M E R I C A N B A S S W O O D

A B Tilia americana L.

American basswood or lindenoccurs as scattered specimens

throughout the state. It grows to aheight of 50–70 feet and a diameter of2–3 feet. The branches are slender,somewhat pendulous, comparativelysmall and numerous, forming a broadand rounded head.

The bark on the trunk of old treesis deeply and irregularly furrowed. Onyoung trees, it is smooth or slightly fis-sured and has a grayish appearance.

The leaves are alternate, 5–6 inch-es long with uneven bases. They arebroadly egg-shaped to heart-shaped inoutline, and toothed; the upper surface

Light, soft, easily worked and carved, American basswood is used formolding, yardsticks, veneer, dowels, furniture, carvings and pulp.

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135A M E R I C A N B A S S W O O D

is dark green, while the lower is yellow-green and shiny.

The flowers are greenish-yellow,borne on a slender stalk that is attachedto a rather long, yellowish, leaf-likebract. They are fragrant, contain anabundance of nectar and open in July.

The fruit is clustered, spherical,covered with short buff-colored hairs,woody and about as large as a pea. Itremains attached to the leaf-like bractwhen it falls.

The twigs have a zigzag patternand bright red buds.

The wood is light, soft, easilyworked and carved. It is used for mold-ing, yardsticks, veneer, dowels, furni-ture, pattern stock, carvings and pulp.Traditionally it was used to make but-ter box molds, dough bowls and otherkitchen items that touched food.

In Germany, basswood is calledthe bee tree. Bees make an excellentgrade of honey from the flowers. Theyoung fruit and flowers ground into apaste make an excellent substitute forchocolate.

The European linden (Tiliaeuropaea L.) and Little-leaf linden(Tilia cordata Mill.) are commonlyplanted as shade trees. They are small-er in height than our native species andwith smaller leaves. Baxter Boulevardin Portland is lined with both of thesespecies.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8American Basswood Circumference: 224'' Height: 85' Crown Spread: 63' Location: Strong

The fruit ofAmerican basswood

is attached to a distinctive

leaf-like bract.

BA

SS

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A R E A O FO C C U R R E N C E

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136 A M E R I C A N E L M

A E Ulmus americana L.

American elm is one of our largestand most graceful trees; it occurs

throughout the state, although itsnumbers have been severely reduced byDutch elm disease. It is found mostoften on rich bottomland and moistsoil along streams, but sometimesgrows on higher ground. It growsquickly, attaining a height of 60–70feet and a diameter of 2–4 feet.

The trunk often divides intonumerous limbs, which form a vase-shaped or spreading, round-toppedhead with graceful, drooping branches.

The number of American elms in Maine hasbeen severely reduced by Dutch elm disease.

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137A M E R I C A N E L M

The bark on the trunk is separat-ed into broad ridges by deep fissuresand is ashy-gray on the surface. Itshows alternate layers of chocolatebrown and buff coloration beneath.

The leaves are alternate, 3–6 inch-es long, with coarsely doubly-toothedmargins and uneven bases. The uppersurface is dark green and sandpaper-like.

The flowers appear in April beforethe leaves.

The fruit consists of a small,winged seed which ripens about theend of May, before the leaves have fullydeveloped. It has a wide, open notch atthe apex and a hairy margin.

The wood is spiral and coarse-grained, hard, heavy, strong, tough andhard to split. It is used for flooring, rail-road ties and pulp. In the past it was usedto make barrel hoops, barn stall flooring,door thresholds and wheel hubs.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8American Elm Circumference: 244'' Height: 110' Crown Spread: 120' Location: Yarmouth

American elm twigs have a zigzag pattern and slightly flattened buds.

EL

M

Slippery elm, Ulmus rubra Muhl,has been recorded in Franklin and Yorkcounties, but these records are histori-cal. A few specimens have been foundin association with cultural settings,but it is not known if these populationsare native or escaped. If it still occursnaturally in the state, it is undoubtedlyquite rare. Slippery elm is most easilydistinguished from American elm bythe winter buds which are covered withrusty hairs. In the past, the inner barkof the slippery elm was chewed torelieve sore throats.

A R E A O FO C C U R R E N C E

Page 140: Forest Trees of Maine

Loads of logs were“snubbed” when goingdownhill to preventthe horses from beingovertaken by the load.

Page 141: Forest Trees of Maine

C H E R R I E S A N D P L U M S

Pin CherryPrunus pennsylvanica

Black CherryPrunus serotina

CommonChokecherryPrunus virginiana

Canada PlumPrunus nigra

B A R K

TEXTURENearly smooth. Large

horizontal lenticels showorange when rubbed.

Young trunks: prominentwhite lenticals.

Older trunks: fissured and ridged.

Smooth with a pungent,disagreeable odor.

Lenticels less prominentthan on other

Prunus species.

Lenticels yellowish

COLOR Reddish-brown Young trunks are blackGrayish-brown, with

light-colored fissuresDull reddish-brown to black

L E A V E S

GENERALDESCRIP-T ION

Long and tapering frombase to tip. Widest in thelower 1⁄3; thin and firm

textured with round teeth.Glands on stalk, and no

hairs on midribs.

Elliptic/oblong, widest inthe center, thick leatheryand shiny. Underside of

midrib near stalk end cov-ered with rusty, brownhairs. Glands on stalknear blade. Margin has

rounded teeth.

Obovate, widest in theterminal 1⁄3, sharply saw-toothed and without hairs,

medium leathery in tex-ture, glands on stalk andno brown hairs on midrib.

Ovate or obovate taperingabruptly into a long thin

point. Teeth rounded.Glands on stalk.

T W I G S

SHAPE Very fine Waxy Medium slenderThorns common on

older twigs

COLOR Red and reddish-brownRed-brown with a lighter

or greenish marginGray or purplish-brown

Current growth gray, oldergrowth darkening to black

ODOR Slight cherry odorSharp, pungent smell

when brokenStrong, pungent bitter-

almond odorNone

B U D S

SHAPEFootball-shaped with a

longitudinal furrowOvate, flattened

Cone shaped, slender –pointed, side buds

not flattenedCone shaped, pointed

COLOR Red-brownRed-brown with a lighter

or greenish marginPurple to tan pattern Gray-brown

F R U I T

COLOR Bright Red Black Deep red to purple Light red to yellow

SIZE 1⁄4 inch diameter 1⁄2 inch diameter 1⁄4–1⁄2 inch diameter 1 inch diameter

ARRANGE-MENT

Hang in umbellate or corymbose clusters

Produced in a raceme,the individual fruit havea persistent basal disc

Produced in racemes,basal disc not persistent

Football-shaped with alongitudinal furrow

The twigs and branches of cherry and plum trees may be distorted by the‘black knot’ fungus (Dibotryon morbosum (Schw.)Th. & Syd.).

C P The Important Distinctions

139

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140 P I N C H E R R Y

P C Prunus pensylvanica L. f.

Pin or fire cherry is a small tree, sel-dom growing taller than 25–30

feet in height and 6–10 inches indiameter. It has slender, horizontalbranches and a narrow, somewhatrounded head. It is common through-out the state, but has little value exceptas a protection and cover for the soilon recent clearings or burned areas.

The bark on the trunk of old trees isdark red-brown and broken into thinplates. Bark on young trees is smoothand reddish-brown. The inner bark isslightly aromatic and very bitter. Thelarge lenticels show orange when rubbed.

The leaves are alternate, narrow tooblong, widest in the lower third,rather sharp pointed, finely and sharplytoothed on the edges, bright green andshiny above, without hairs below, 3–4inches long, bitter and aromatic, withglands on petiole.

The flowers are white, in clustersof four or five, and appear in Maywhen the leaves are only half grown.The fruit is bright red, almost translu-

Pin cherry is not usedcommercially and has littlevalue except as a protectionand cover for the soil on recentclearings or burned areas.

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141P I N C H E R R Y

cent, pea-sized, globular and ripens fromthe first of July to August. In the past, itwas used to make jams and jellies.

The twigs are shiny, reddish-brown and very slender. The buds aresmall, ovoid, reddish-brown and clus-tered at the end of twigs. They arecommonly distorted by a black, warty,fungus growth called “black knot.”

The wood is coarse-grained, softand light. It is not used commercially.The gum is edible.

Pin cherry produces abundant5-petaled flowers every spring.

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142 B L A C K C H E R R Y

B C Prunus serotina Ehrh.

Black cherry is widely distributedthroughout the state. In some

parts of the country it is an extremelyvaluable timber tree, but in Maine doesnot often grow to sufficient size. It growson a variety of soils, but makes rapid andbest growth on rich, moist land. It has anarrow head, small horizontal branches,and attains a height of 40–50 feet and adiameter of 10–20 inches.

The bark on the trunk of youngtrees is red-brown to black and rather

Black cherry is one of our most valuable timber trees, although not abundant in sufficient size.

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A R E A O FO C C U R R E N C E

143B L A C K C H E R R Y

shiny with prominent white lenticels.On older trees, the bark is broken intosmall irregular plates.

The leaves are alternate, elliptic,oblong, widest at the center, finelytoothed, dark green, shiny, thick,somewhat leathery and 2–5 incheslong. The underside of the midrib nearthe stalk end is covered with rustybrown hairs.

The flowers are produced in manyflowered racemes 4–5 inches long thatappear at the end of May or in earlyJune when the leaves are half-grown.The fruit is in drooping racemes, darkpurple or almost black when ripe, ¼–½inch in diameter, and globular in shape.It ripens from June to October and isan important wildlife food.

The twigs give off a pungent odorwhen broken, and the bark has a bittertaste. As with pin cherry, twigs andbranches are commonly distorted by ablack, warty, fungus growth called“black knot.”

The wood is rather hard, close-grained, light, strong and easily pol-ished. It is used for furniture and cabi-netmaking, interior finishing, wooden-ware, veneer and plywood. It is valuedas firewood due to its high heat valueand fragrance.

The rusty hairs alongthe midvein on the backof the leaf distinguishblack cherry.

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

Black CherryCircumference: 148''

Height: 52'Crown Spread: 60'Location: Falmouth

Black cherry twigshave a bitter tasteand give off a pungentodor when broken.

CH

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144 C O M M O N C H O K E C H E R R Y

C C Prunus virginiana L.

Common chokecherry is a shrubor small tree that occurs through-

out the state, especially along fencerowsin farming communities. It occasion-ally is 25 feet high and 6 inches indiameter.

The smooth, grayish-brown toblack bark is usually marked by long,light-colored fissures and has a dis-agreeable scent.

The leaves are alternate, dull,widest at the terminal one-third, 2–4inches long, finely-toothed, medium-leathery in texture and at maturity arewithout hairs.

The flowers appear from the firstof May to June on slender stalks inracemes.

Common chokecherry wood isheavy and hard, but not strong,and is not used commercially.

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145C O M M O N C H O K E C H E R R Y

The fruit ripens from July toSeptember, and is about ¼–⅓ inch indiameter, at first bright red, turning atmaturity to dark red or nearly black. Itis slightly astringent, but edible.

The winter buds are strictly cone-shaped, slender and pointed with adefinite purple and tan pattern on thescales. Side buds are not flattened as inblack cherry.

The twigs have a strong, pungent,skunk-like odor when broken and, aswith pin cherry and black cherry, are

Chokecherry twigs havecone-shaped buds andgive off a strong odorwhen broken.

CH

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RY

frequently distorted by a black, warty,fungus growth called “black knot.”

The wood is heavy, hard but notstrong, and is not used commercially.

A R E A O FO C C U R R E N C E

Page 148: Forest Trees of Maine

black knot fungus, Apiosporina morbosa.The bark is thin, dull reddish-

brown to black; it peels in thin paperyscales, exposing the shiny reddish-brown, inner bark.

The leaves are alternate, obovate,and taper at the apex to a long, sharppoint. Leaves are dark green on theupper surface, lighter below; the mar-gin has glandular, rounded teeth.

The flowers are white, and appearearly in spring before the leaves ingroups of three or four on slenderstalks. The edible fruit ripens the latterpart of August, is football-shaped andfurrowed along one side, and has anorange-red skin and yellow flesh. Thesingle stone is flattened and slightlygrooved on the edges.

The twigs and branches oftenhave thorns. The buds are brown togray and are without hairs. The wood isnot used commercially.

146 C A N A D A P L U M

C P Prunus nigra Ait.

Canada or red plum, has beenplanted as an ornamental and is

found occasionally throughout much ofthe state. It does not occur in denselyforested areas; rather it usually occurs inthickets along field edges. It is seldomover 8 inches in diameter and 30 feethigh. The twigs and branches of cherryand plum trees are distorted by the

The Canada plum’s fruit,which ripens in the latterpart of August, is edible.

PL

UM

A R E A O FO C C U R R E N C E

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147H A W T H O R N

Hawthorn, or thorn-apple, occursin Maine as a low spreading tree

or shrub that rarely reaches a height of more than 15–18 feet. There areapproximately 22 different speciesfound in the state. Hawthorns canusually be recognized by the smallapple-like fruits and the thorns on thebranches. In the past, hawthorns wereplanted as hedges in place of fencing.

The bark is dark brown to ashygray and somewhat scaly.

The leaves are alternate, doubly-toothed, and usually somewhat lobed,thin and dark green.

The flowers appear about the firstof June in flat, showy white clusters.

The fruit, which is ¾ inch indiameter, resembles a small apple. Theflesh is thin, mealy and encloses 1–5rounded nutlets. It is used for jelliesand bird food.

The twigs are slender, rigid andusually armed with long thorns. Theyform a compact crown due to theirzigzag method of growth.

H Crataegus spp. L.

The wood is heavy, hard andclose-grained. It is used to some extentfor handles and other small articles.

The fruit of the hawthornis used for jellies and bird food.

HA

WT

HO

RN

A R E A O FO C C U R R E N C E

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148 S E R V I C E B E R R Y

Approximately seven species ofserviceberry or shad bush grow

as shrubs or small trees in Maine. Ofthese, two species—Allegheny service-berry Amelanchier laevis Wieg. anddowny serviceberry Amelanchier arborea(Michx. f.) Fern.—commonly grow to besmall trees 30–40 feet in height and 6 to8 inches in diameter. Allegheny service-berry, is the more common of the two.They are both found in open hardwoodstands or along the margins of open areasthroughout much of the state.

The bark of serviceberry is smooth,gray to light violet-brown with darkervertical stripes; older bark is slightly fis-sured longitudinally and twisted.

The leaves of Allegheny service-berry are half grown at flowering time,and have a reddish or purplish tinge.When downy serviceberry leaves arejust unfolding, they are green anddensely hairy beneath. Mature leavesof both species are alternate, dark green

S Amelanchier spp. Medik.

Serviceberry wood isoccasionally used for toolhandles, small implementsand fishing rods.

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149S E R V I C E B E R R Y

The flowers are white and sweet-smelling with 5 petals. The serviceber-ry flowers before other trees and is veryeasy to spot along the edges of fieldsand streams in spring.

Serviceberry fruit is berry-like,ripens in early summer, is ⅓–½ inch indiameter, and red to dark purple whenmature and edible. Serviceberry twigsare slender, red-brown and finely hairywhen young, becoming smooth as thetwigs grow. The buds are long, sharppointed, reddish or pinkish, and filledwith silky hairs.

The wood is occasionally used fortool handles, small implements andfishing rods. It is heavy, hard, strong,close-grained and dark brown tingedwith red.

Serviceberry buds are longand sharp-pointed; thelateral buds hug the twig.

above and lighter green below, 1½–3inches long, 1–1½ inches wide, ellipticto ovate with a rounded or heart-shaped base.

SE

RV

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150

There are two native species ofmountain ash found in Maine:

the American mountain ash SorbusAmericana Marsh, also called round-wood, and the showy or Northernmountain ash Sorbus decora (Sarg.)Schneid.

American mountain ash occursstatewide; it is not a true ash, but isclosely related to the apple. It rarelyreaches over 20 feet in height. It is par-ticularly common in mountainousregions and along the coast. The leavesare alternate, compound 13–17 incheslong, tapered, and have 11–17 finelytoothed leaflets. The leaflets are 2–4inches long, ⅝–1 inch wide, and with-out hairs.

The small creamy-white flowersare borne in cymes. The berry-likefruit is bright red, and about ¼ inch indiameter. These remain on the tree lateinto the winter; they make good bird

M A Sorbus spp.

Showy mountain ash is usuallybetter balanced in outline thanthe American mountain ash andhas a well-rounded crown.M O U N T A I N A S H

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M O U N T A I N A S H 151

This photo is of American mountain ash fruit.Showy mountain ash fruit is larger.

food. In the past, they were sometimesused as an astringent in medicine. Thebud scales are hairless and sticky. Thepale brown wood has little valuebecause it is soft and weak.

Showy mountain ash is most com-monly found in northern and westernparts of the state. It is usually betterbalanced in outline than the Americanmountain ash and has a well-roundedcrown. The leaves are alternate, com-pound, and differ from the precedingspecies in having leaflets which are only1½–3 inches long, and ⅝–1⅝ incheswide. The fruit is larger, up to ½ inchin diameter, and matures later in theseason. The outer bud scales are sticky;the inner scales are hairy.

MO

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IN

AS

H

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152 S H A G B A R K H I C K O R Y

S HCarya ovata (P. Mill.) K. Koch Shagbark hickory is most common-

ly found in southern Maine onmoist but well-drained soil. It has acylindrical head and a straight, gradu-ally tapering trunk. It reaches a heightof 70 feet and a diameter of 2 feet.

The bark is light gray on the trunkand separates into long, loose plates,giving it a shaggy appearance.

The leaves are compound, alter-nate, 8–14 inches long; most oftenthere are 5 leaflets, rarely seven. The 3terminal leaflets are the largest. Leafletmargins are serrate.

The fruit has the thick outer huskdeeply grooved at the seams. The huskseparates along these grooves whenripe. The fruit is globose and is bornesingly or in pairs. The edible kernel issweet. The twigs are hairy or smoothand olive-gray to dark red-brown. Pithis star-shaped in cross section. Budscales are hairy.

Shagbark hickorywood is primarilyused to make pallets.

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153S H A G B A R K H I C K O R Y

The wood is very strong, close-grained, heavy, hard, tough and flexi-ble. It was formerly used in the manu-facture of agricultural implements, axeand tool handles, carriages and wag-ons, especially the spokes and rims ofthe wheels. Its principal uses are nowpallets, pulp and firewood.

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8Shagbark Hickory*

Circumference: 116''/133''Height: 84'/72'

Crown Spread: 60'/58'Location:

Falmouth/Westbrook *TIE

Shagbark hickory is easily distinguished by itsbark, which separates into long, loose plates.

HI

CK

OR

Y

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154 B I T T E R N U T H I C K O R Y

B HCarya cordiformis (Wangenh.) K. Koch

Although common further south,bitternut hickory is rare in

Maine, occurring only in the extremesouthwestern corner of the state at thesouthern tip of York County. Bitternuthickory will grow on a variety of sites,but makes its best growth on moistbottomland soils.

The bark of young trees is silvery-gray and smooth; older trees have graybark with tight, shallow, interlacing fur-rows. The bark remains tightly attachedon old trees and does not become shag-gy. The leaves are 8–10 inches long,alternate, pinnately compound with 7–9leaflets. The terminal leaflet is similarin size to the adjacent ones.

The flowers occur in spring; maleflowers are in catkins and female flow-ers are in a terminal spike. The fruit isa nut; it is nearly round and onlyslightly flattened. It is covered by a thin green husk with 4 small wingsdescending from a sharp point to themiddle. As the name bitternut implies,

the meat is very bitter and not eaten byhumans, although some wildlife utilizeit as food. The twigs are somewhatstout (although much less so thanother hickories) and have distinctivesulfur-yellow buds.

The wood is hard; further south,it used for making tool handles, furni-ture, paneling and pallets, as well as forfuel. It is a choice wood for smokingmeats. In Maine, due to its rarity, thewood is not used commercially.

HI

CK

OR

Y

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B W Juglans nigra L.

Black walnut is not native toMaine, but is planted occasional-

ly as an ornamental tree. In forestedsituations in its native range, it cangrow to be up to 100 feet tall with along straight trunk free of branches.In Maine, it is usually planted in theopen and exhibits an open-grown formwith wide-spreading branches. Blackwalnut’s natural range extends over alarge portion of the eastern UnitedStates from western Vermont andMassachusetts to southeastern SouthDakota, south into Texas and theFlorida panhandle.

The bark is brown, with furrowedridges forming a diamond pattern. If thebark is cut with a knife, the cut surfacewill be dark brown. The leaves are alter-nate, pinnately compound 12–24 incheslong with 10–24 leaflets; a terminalleaflet is often lacking.The fruit is roundand composed of a nut enclosed in athick green husk. The twigs are stout,light brown, with a chambered pith.Thebuds are large and tan.

The wood is so valuable that, in

M A I N E R E G I S T E R O FB I G T R E E S 2 0 0 8

Black WalnutCircumference: 158''

Height: 100' Crown Spread: 87'Location: Limerick

WA

LN

UT

155B L A C K W A L N U T

some parts of the country, trees havebeen stolen in the dead of night fromfront lawns and city parks. It is a rich,dark brown and takes a good polish,making it valuable for furniture, cabi-nets and gunstocks. Much of the woodharvested today is turned for veneer.

The nuts are edible, but must begathered before the animals harvestthem all. Ground nut shells have hadnumerous uses, including as a carryingagent for insecticidal dusts and forcleaning aircraft engine parts; while thefruit husks have been used to makefabric dye.

Page 158: Forest Trees of Maine

156 B U T T E R N U T

B Juglans cinerea L.

Butternut, also known as whitewalnut, occurs naturally or in

cultivation to some extent statewide. Itgrows on rich, moist soil and on rockyhills, especially along fencerows. It fre-quently has stout, spreading limbsextending horizontally from the trunkto form a low, broad, rounded head. Itgrows to 30–40 feet high and a diame-ter of 1–2 feet. Currently, butternut isunder severe threat from butternutcanker, Sirococcus clavigignenti-juglan-dacearum. This fungus was most likelyintroduced from outside of NorthAmerica and is now killing butternutsthroughout much of Maine.

The bark of young trees and of thebranches is gray. On old trees, it is broad-ly ridged on the trunk and light brown.

The leaves are compound with aterminal leaflet, alternate, 15–30 inch-es long, and consist of 11–17 leaflets.The leaflets have serrate margins.

Butternut is sometimesused for furnitureand cabinetwork andtakes a high polish.

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157B U T T E R N U T

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Butternut Circumference: 186'' Height: 74' Crown Spread: 88' Location: Durham

The fruit is composed of a nutenclosed by a fleshy husk covered withsticky hairs. It is about 2½ inches longand oval shaped. Fruit is produced indrooping clusters of 3–5. The nut isthick-shelled with sharp ridges on thesurface. American Indians used the oilfrom the nuts to make butter. Browndye was made from the husk.

The twigs are stout, greenish andhairy, with chocolate-brown, cham-bered pith. The large leaf scars have aconspicuous, buff-colored hairy pad atthe top; the buds are also hairy.

The wood is coarse-grained, light,soft and weak. It is sometimes used forfurniture and cabinetwork and takes ahigh polish.

BU

TT

ER

NU

T

Page 160: Forest Trees of Maine

158 H O R S E C H E S T N U T

Not related to the native chestnut,the horsechestnut comes from

Asia and the Balkan Peninsula and isgenerally planted as a shade and orna-mental tree.

It is symmetrically round or ovalin outline with a stiff branch habit. Thetips of the branches curve slightly whenmature. It has heavy, luxuriant, deepgreen foliage which changes to bronzein early autumn. The large, oppositeleaves with 5–7 leaflets, are arrangedpalmately on a single stalk; and distin-guish it from any of Maine’s nativetrees. With the pyramids of white flow-ers blossoming in the early spring andthe large, bur-like, leathery huskenclosing one or more smooth,mahogany-colored nuts, the horsech-estnut is not easily confused with anyother species. The nuts are poisonouswhen ingested. It makes a good shade

tree, but requires rich soil for bestdevelopment. It is prone to a leaf blight.

The buds are large, sticky andnearly black. The wood is soft, lightand close-grained. In Europe, it is usedfor carving and veneer. In the past inthe U.S., it was burned as firewood.

H Aesculus hippocastanum L.

HO

RS

EC

HE

ST

NU

T

Page 161: Forest Trees of Maine

159A M E R I C A N S Y C A M O R E

diameter. It contains a large number ofsmall wedge or shoenail-shaped nut-lets, and usually remains on the treeuntil spring.

The twigs are zigzag in shape andare encircled by conspicuous stipules.The winter buds have a single, wrin-kled, cap-like scale.

The wood is hard, firm, very perish-able when exposed to the weather, andliable to warp. In the past, it was used forfurniture and interior finish of houses.

There are historic records ofAmerican Sycamore occurring

along streams and on rich bottomlands in southern Maine. Currentlythere are no known native populationsin the state. However, sycamore isplanted here as an ornamental. Farthersouth and west it grows to be an enor-mous tree, often 4-6 feet in diameterand 120 feet tall, trees in Maine how-ever do not attain great size.

The bark on the trunk and largelimbs is greenish-gray and flakes off inbroad scales exposing white patchesbeneath.

The leaves are simple, alternate,3–5 lobed and light green. The base ofthe leaf-stalk is hollow, swollen andcovers the bud.

The fruit head generally occurssingly, is round and about 1 inch in

A S Platanus occidentalis L.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8American Sycamore Circumference: 160'' Height: 105' Crown Spread: 93' Location: Waterford

SY

CA

MO

RE

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160 B L A C K T U P E L O

B T Nyssa sylvatica Marsh.

Black tupelo, or blackgum, is foundin Sagadahoc, Androscoggin,

Cumberland and York counties and asfar north as Southern Oxford Countyand Waterville in Kennebec County.However, it is not commonly foundexcept in very wet areas. Trees 2 feet indiameter are found in the town ofCasco on an island in Sebago Lake.Large specimens have also been report-ed on the south side of PleasantMountain in Denmark on a flat, open,wet area. Easily distinguished at a dis-tance by its numerous slender horizon-tal branches, the tree rarely reachesmore than 50 feet in height. It occurs inrich moist soils, such as swamps or bor-ders of rivers. Black tupelo can live to avery old age. Trees over 500 years oldhave been found in New Hampshire.

The bark on young trees issmooth, grayish and flaky, later becom-ing reddish to grayish-brown. On oldtrees, it forms coarse blocks or ridges.

While black tupelo wood isheavy, fine-grained and verytough, it is not durable and isused principally for pulp.

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161B L A C K T U P E L O

The leaves are alternate, oval toobovate, 2–5 inches long, wedge-shaped at the base and pointed at thetip. The edges are usually entire. Theleaves are dark green, shiny above,occasionally hairy below, and turnbright crimson in autumn.

The fruit is dark blue, fleshy,approximately ½ inch in length, andborne in clusters of 1–3 on long, slen-der stems. The fruit has an acid taste,but is edible.

The twigs are moderately stoutwith a diaphragmed pith.

The wood is heavy, fine-grained,very tough but not durable. It was for-merly used for the hubs of wheels andsoles of shoes. It is now used principal-ly for pulp.

Black tupelo hasa characteristichorizontalbranching pattern.

TU

PE

LO

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162 B L A C K L O C U S T

The locust is a rapid grower,frequently attaining a heightof 20 feet in 10 years, butincreasing much more slowlythereafter.

Black locust is not a native of thisstate, but is extensively planted.

It is abundant in some localities, andis found mostly near dwellings or onabandoned farmlands, where it oftenbecomes naturalized. The locust is arapid grower, frequently attaining aheight of 20 feet in 10 years, butincreasing much more slowly there-after. It reaches a height of about 50feet and a diameter of 8–20 inches.The branches are small, brittle, occa-sionally multi-angled, and at first arearmed with stipular spines. The top isnarrow and oblong. It is one of the lasttrees to send out foliage in the spring.

The bark on old trees is darkbrown, deeply furrowed and brokeninto small scales.

The leaves are alternate, oncecompound, 8–14 inches long, with7–19 leaflets that are about 2 incheslong with an entire margin and aslightly notched tip.

B L Robinia pseudoacacia L.

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163B L A C K L O C U S T

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Black Locust Circumference: 205'' Height: 80' Crown Spread: 66' Location: Belfast

The flowers are borne in looseracemes 4–5 inches long. Showy andvery fragrant, they appear in June.

The fruit is a smooth, flat, darkpurplish-brown pod about 3–4 incheslong, containing 1–8 bean-like seeds.

The wood is heavy close-grained,strong, and very durable when in con-tact with the soil. It is used for fenceposts, firewood and planking for boats.In the past, it was used to make pegsfor use with glass insulators.

The buds of the black locust are almostcompletely hidden.

LO

CU

ST

Page 166: Forest Trees of Maine

164 H O N E Y L O C U S T

Honeylocust is not native toMaine, but has been frequently

planted in urban areas in the southernand central portions of the state. Thetrees most commonly planted andthose that have escaped cultivation inMaine are a thornless variety, Gleditsiatriacanthos f. inermis (L.) Zabel.

Honeylocust has somewhat pen-dulous, slender, spreading branches thatform an open, broad, flat-topped head.It attains a height of 75 feet and adiameter of 20 inches. Simple or (usu-ally) three-forked spines, 1½–3 incheslong or longer, occur on the branchesand trunk; but spines are lacking on thecommonly planted variety.

The bark is divided into long, nar-row ridges by deep fissures; and thesurface is broken into small scales thatare persistent. The leaves are alternate,both once and twice compound, 4–8inches long and have from 18–28leaflets. The margins of the leaflets arefinely blunt-toothed.

The flowers are borne in slenderclusters 2–2½ inches long. Theyappear in June when the leaves areabout fully-grown. Staminate and pis-tillate flowers are produced separatelyon the same tree.

The fruit is a shiny, reddish-brown, flattened pod 8 inches or morein length. The pod is curved, withirregular wavy edges, and is oftentwisted. The walls are thin and tough.

The twigs are smooth and dis-tinctly zigzag in shape. Winter budsbarely protrude from the leaf scar.

The wood is coarse-grained, hard,strong, and very durable in contact withthe soil. It is used for firewood and boatdecking in Maine. In the past, it was usedto manufacture the wooden pegs thatglass insulators were screwed onto whenglass insulators were used with telegraph,telephone and electrical power lines.

H Gleditsia triacanthos L.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Honeylocust Circumference: 178'' Height: 50'

Crown Spread: 80' Location: Bath

LO

CU

ST

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165S A S S A F R A S

Sassafras occurs in southern Mainein eastern Cumberland, southern

Oxford and York counties, and issometimes planted for ornament.Excellent specimens may be seen inthe York Village cemetery.

The bark on young stems is thinand reddish-brown. On older stems, itbecomes thick and scaly.The inner barkis very fragrant and sometimes chewed.

The leaves are alternate, veryhairy when they first appear, losing thehair at maturity except on the midrib.They are light green and of 3 shapes:entire, mitten-shaped and three-lobed.

The flowers open in early springwith the first leaves, in racemes con-taining about 10–15 flowers.

The fruit ripens in September andOctober and is a blue, lustrous drupethat is supported on a fleshy, red stalk.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Sassafras Circumference: 82'' Height: 66' Crown Spread: 21' Location: York

The twigs are green in color,smooth and aromatic when broken.

The wood is soft, weak, brittle,very aromatic, light brown and verydurable in the soil. Historically, theroots and bark were distilled for oil ofsassafras, used to perfume toiletries.The oil has been banned from use infoods in the US.

S Sassafras albidum (Nutt.) Nees

SA

SS

AF

RA

S

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166 N A N N Y B E R R Y

N Viburnum lentago L.

Nannyberry occurs statewide as ashrub or small tree reaching a

height of 10–30 feet. It frequently isfound growing in moist soils, oftenalong the borders of swamps or streams.

The leaves are opposite, ovate,abruptly pointed, with fine sharp teeth.The upper surface is a lustrous deepgreen. The undersurface is lighter. Thepetiole is conspicuously flanged with awarty, wavy margin.

The dark blue fruit ripens in fall.It is about ½ inch long, ellipsoid, edible,sweet, tough-skinned, with a nipple-like tip. The fruit occurs in smalldrooping clusters on red-stemmedstalks, and does not shrivel or shrinkwhen ripe.

The terminal buds are shaped likea pair of rabbit ears and bulge at thebase. The 2 large bud scales extendbeyond the end of the bud. They arenearly smooth and are purplish-brownto lead-colored. The smooth twigs ofthe season are gray to gray-brown. Thewood is orange-brown and emits anunpleasant odor.

NA

NN

YB

ER

RY

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167W I T C H - H A Z E L

W - H Hamamelis virginiana L.

Witch-hazel occurs as a smalltree or shrub in most parts of

Maine except in the far north. It isfound on borders of the forest in lowrich soil or on rocky banks of streams.

The bark is gray-brown and some-what scaly on older stems. The leavesare alternate, broadly obovate, non-symmetrical at the base, and have awavy margin.

It has bright yellow flowers withthread-like petals in autumn or earlywinter. The fruit is a woody capsule,usually two in a cluster. The seeds aredischarged fiercely when ripe.

The twigs are gray, zigzag, withgray or rust-colored hair and scalpel-shaped buds.

An extract from the bark is mixedwith alcohol and used as an astringent.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Witch-hazel Circumference: 18'' Height: 32' Crown Spread: 17' Location: Rockport

WI

TC

H-

HA

ZE

L

Page 170: Forest Trees of Maine

168 S T A G H O R N S U M A C

Staghorn sumac is a shrub or smalltree that grows throughout most

of the state. It can grow to about 25feet tall and about 8 inches in diameter,although it is usually smaller.Occurring mostly on disturbed sitessuch as road sides and old fields,staghorn sumac sprouts readily fromthe roots. It often forms thickets thathave a characteristic domed-shapedappearance with the tallest stems in thecenter. Unlike the unrelated poisonsumac, staghorn sumac is not poison-ous to the touch.

The bark is grayish-brown andhas numerous lenticels. The leaves are16–24 inches long, alternate, pinnatelycompound with 11–31 opposite, ser-rate leaflets. The leaves turn a brilliantred in fall. The flowers form in earlysummer in large, compact, yellow pan-icles 2–8 inches long. The fruit ripensin August as a spire of showy, red, vel-vety berries that often remain into the

winter. The twigs are stout and veryhairy. The winter buds are not coveredby scales.

The wood has a greenish cast withdark grain. It is not used commerciallyexcept for small specialty items.Because the wood has a chamberedpith which can be easily cleaned out, itwas used for sap spiles (tubes for col-lecting sap in a sugarbush).

S SRhus hirta (L.) Sudworth

SU

MA

C

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169M O U N T A I N L A U R E L

M L Kalmia latifolia L.

Mountain laurel is an erect-stemmed low shrub or small

tree that grows in rocky woods or on lowground. Mountain laurel occurs rarely insouthern and western Maine and is list-ed as a species of special concern.

The leaves are evergreen, green onboth sides, elliptical, up to 3 incheslong and 1 inch wide. They are flat,thick and leathery with an entire mar-gin, and narrowed at both ends.Arrangement is mostly alternate,grouped at the tip of the twig, some-times opposite and rarely in threes.

The flowers are pink with varia-tions possible. They are borne in erect,terminal clusters.

The fruit are globose, woody cap-sules borne on erect, hairy, sticky stalksthat are many times longer than thediameter of the capsules. The capsuleshave long, persistent styles.

The twigs are rounded and stickyat first, but later become smooth.

LA

UR

EL

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170 R O S E B A Y R H O D O D E N D R O N

R R Rhododendron maximum L.

Rosebay rhododendron, or greatlaurel, is a shrub or straggling

tree up to 30 feet high. It is a very rarespecies found locally in parts ofSomerset, Franklin, Cumberland and

Rosebay rhododendronis listed as a threatenedspecies in Maine.

York counties in damp woods or nearpond margins. It is listed as a threat-ened species in Maine.

The leaves are evergreen, ovate tooblong, alternate, entire, 4–8 incheslong, thick and leathery, with the marginfrequently rolled under.They are smoothand dark green above, pale below. Theflowers are bell-shaped and occur indense clusters. They are generally whitewith a pinkish tinge with other varia-tions possible. The fruit is an oblong,woody capsule covered with sticky hairs.It is borne terminally in erect clusters onstalks several times longer than the cap-sule. The twigs are hairy.

RH

OD

OD

EN

DR

ON

Page 173: Forest Trees of Maine

171F L O W E R I N G D O G W O O D

F D Cornus florida L.

Flowering dogwood is an unusual-ly beautiful shrub or small tree

and occurs naturally only in YorkCounty. Planted specimens generally areonly hardy in the southern and coastalareas of the state. Unfortunately flower-ing dogwood is under serious threat fromDogwood anthracnose Discula destruc-tive, a non-native fungal disease.

Flowering dogwood reaches aheight of 12–20 feet. The bark is grayand smooth on younger stems; on oldertrees it becomes black and finely blocky,as if broken into small squares. Theleaves are opposite, entire, ovate toelliptic, bright green and smooth above,pale green with hairs on the veinsbeneath. They are 3–6 inches long.

The flowers are conspicuous andappear early in the spring. They aregreenish-white or yellowish and arearranged in dense umbels surrounded

by 4 large, white, petal-like bractswhich give the appearance of largespreading flowers. The fruit is a brightred, ellipsoid drupe about ½ inch longthat occurs in clusters.

The twigs are smooth, greenishand angular. The buds are covered bytwo valve-like scales. The flower budsare large and button-shaped.

The wood is hard and close-tex-tured. In the past, it was widely usedfor the manufacture of shuttles for tex-tile weaving. It is not used commercial-ly in Maine due to its rarity.

M A I N E R E G I S T E R O F B I G T R E E S 2 0 0 8Flowering Dogwood Circumference: 35'' Height: 37' Crown Spread: 30' Location: Portland

DO

GW

OO

D

Page 174: Forest Trees of Maine

172 A LT E R N A T E - L E A F D O G W O O D

A- D Cornus alternifolia L. f.

Alternate-leaf or blue dogwoodoccurs throughout the state as a

shrub or small tree up to 20 feet tall.The leaves are alternate, entire,

elliptic-ovate and tend to be crowdedat the ends of the twigs. They are2½–4½ inches long, yellowish-green,smooth above and have appressed hairsbeneath.

The creamy white flower clustersappear in June after the leaves havedeveloped. The fruit is a bluish-blackdrupe, somewhat round, about ⅓ inchin diameter, that ripens in Septemberand October.

The twigs are often lustrous andgreenish-brown. Dead twigs becomebright yellow-green.

DO

GW

OO

D

Page 175: Forest Trees of Maine

173R E D O S I E R D O G W O O D

Red osier dogwood is an abundantcolonial shrub usually less than

10 feet tall. On very rare occasions it

may reach the size of a small tree. Itoccurs throughout the state. It growson the edges of fields and streams, andin wet areas. It readily invades fields,where it is considered a pest.

The leaves are opposite, entire,lance-shaped to elliptic to ovate, 2–4inches long and whitened underneath,with 5–7 lateral veins.

The flowers are in flat-toppedclusters. The fruit is white and ¼–⅓inch in diameter. The twigs are brightred to green and minutely hairy. Theyare used for handles for cemetery bas-kets. The buds lack scales and aredensely hairy.

R O D Cornus sericea L.

Red osier dogwood has opposite leaves andbright red twigs.

DO

GW

OO

D

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174 S E L E C T E D R E F E R E N C E S

S R T S

GDwelley, Marilyn J. Trees and Shrubs of New England, Down East Books, printed by TwinCity Printery, Lewiston, Maine, 1980.Elias, Thomas S. The Field Guide to North American Trees, Van Nostrand Reinhold Co.,New York, 1991.Fernald, Merritt L. Gray’s Manual of Botany. (Illus.) 8th ed. (1950) American Book Co., 1987.Graves, Arthur H. Illustrated Guide to Trees and Shrubs, Harper & Row Publishers, NewYork, 1956. (Revised by Dover Publ.)Harlow, William M., J. W. Hardin and F. M. White. Textbook of Dendrology. (Illus.) 7th ed.McGraw-Hill Book Co., Inc., New York, 1991.Hyland, Fay. Conifers of Maine, (Illus.) U. of Maine Ext. Ser. Bull. 345. (Revised) 1961.Hyland, Fay and Ferdinand H. Steinmetz. Trees and Other Woody Plants of Maine, Orono.University of Maine Press, 1978.Little, Elbert. The Audubon Society Field Guide to North American Trees: Eastern Region,Alfred A. Knopf, New York, 1980.McMahon, Janet S., G.L. Jacobson and Fay Hyland. An Atlas of the Native Woody Plantsof Maine; A Revision of the Hyland Maps. Maine Agriculture Expt. Sta. Bul. 830. 1990.Petrides, George A. A Field Guide To Eastern Trees, The Peterson Field Guide Series:Houghton Mifflin Co., Boston, 1988.Symonds, George, W.D. The Tree Identification Book, William Morrow & Co., Inc, 1958.USDA, NRCS. 2005. The PLANTS Database, Version 3.5 (http://plants.usda.gov). Datacompiled from various sources by Mark W. Skinner. National Plant Data Center, BatonRouge, LA 7087-4490 USA.

SCampbell, C.S. et al. Revised Checklist of Vascular Plants of Maine. Orono. MaineAgriculture and Forestry. Expt. Sta. Bul. 844. 1995.Hyland, Fay and Barbara Hoisington. The Woody Plants of Sphagnous Bogs of NorthernNew England and Adjacent Canada. Illustrated by Laurel Smith. Orono, University ofMaine Press, 1981.

K C L Britton, N.L., and A. Brown. 1913. Illustrated flora of the northern states and Canada.(Source of some of the images).Campbell, C.S., Fay Hyland, and M.L.F. Campbell. Winter Keys to Woody Plants of Maine,Orono. University of Maine Press. Revised Ed. 1978.Dearborn, Richard G. Key to the Woody Alpine and Subalpine Flora of Mount Katahdin. Me.Field. Nat. 19 (6): 83–90, 1963.Haines, A., and T.F. Vining. Flora of Maine. V.F. Thomas Co. Bar Harbor, Maine. 1998.

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Abies balsamea (L.) P. Mill. . . . . . . . . . . . 46–47Acer negundo L. . . . . . . . . . . . . . . . . . . . . . 84–85Acer pensylvanicum L. . . . . . . . . . . . . . . . . 78–79Acer platanoides L. . . . . . . . . . . . . . . . . . . 82–83Acer rubrum L. . . . . . . . . . . . . . . . . . . . . . 72–73Acer saccharinum L. . . . . . . . . . . . . . . . . . 76–77Acer saccharum Marsh . . . . . . . . . . . . . . . . 74–75Acer spicatum Lam. . . . . . . . . . . . . . . . . . . 80–81Aesculus hippocastanum L. . . . . . . . . . . . . . . 158Alder, Speckled . . . . . . . . . . . . . . . . . . . . . 102Alnus incana ssp.Rugosa (Du Roi) Clausen . . 102Amelanchier arborea Wieg . . . . . . . . . . 148–149Amelanchier laevis (Michx.f.) Fern . . . 148–149Amelanchier spp. Medik. . . . . . . . . . . . 148–149Arborvitae, Eastern . . . . . . . . . . . . . . . . . 54-55Ash, Black (Brown) . . . . . . . . . . . . . . 132–133Ash, Green (Red) . . . . . . . . . . . . . . . 130–131Ash, Mountain (Roundwood and

Showy Northern) . . . . . . . . . . . . . . 150–151Ash, White . . . . . . . . . . . . . . . . . . . . 128–129Aspen, Bigtooth . . . . . . . . . . . . . . . . . . 64–65Aspen, Quaking (Trembling) . . . . . . . . 62–63Balm-of-Gilead . . . . . . . . . . . . . . . . . . . 66–67Balsam Fir . . . . . . . . . . . . . . . . . . . . . . . 46-47Balsam Poplar . . . . . . . . . . . . . . . . . . . . 66–67Basswood, American . . . . . . . . . . . . . 134–135Beech, American . . . . . . . . . . . . . . . . 104–105Betula alleghaniensis Britt. . . . . . . . . . . . . 92–93Betula cordifolia Regel. . . . . . . . . . . . . . . . . . . 96Betula lenta L. . . . . . . . . . . . . . . . . . . . . . 94–95Betula papyrifera Marsh. . . . . . . . . . . . . . 88–89Birch, Gray . . . . . . . . . . . . . . . . . . . . . . 90–91Birch, Mountain Paper . . . . . . . . . . . . . . . . 96Birch, Paper (White or Canoe) . . . . . . . 88–89Birch, Sweet (Black or Cherry) . . . . . . . 94–95Birch, Yellow . . . . . . . . . . . . . . . . . . . . . 92–93Boxelder . . . . . . . . . . . . . . . . . . . . . . . . 84–85Butternut . . . . . . . . . . . . . . . . . . . . . . 156–157Carpinus caroliniana Walt. . . . . . . . . . 100–101Carya cordiformis (Wengenh.) K. Koch . . . 154Carya ovata (P. Mill) K. Koch . . . . . . 152–153Castanea dentate (Marsh.) Borkh. . . . . 124–125Cedar, Atlantic White . . . . . . . . . . . . . . 52–53Cedar, Eastern Red . . . . . . . . . . . . . . . . 56–57Cedar, Northern White . . . . . . . . . . . . . 54–55Chamaecyparis thyoides (L.) B.S.P. . . . . . 52–53Cherry, Black . . . . . . . . . . . . . . . . . . . 142–143Cherry, Pin (Fire) . . . . . . . . . . . . . . . 140–141Chestnut, American . . . . . . . . . . . . . 124–125Chokecherry, Common . . . . . . . . . . 144–145Cornus alternifolia L. f. . . . . . . . . . . . . . . . . . 172Cornus florida L. . . . . . . . . . . . . . . . . . . . . . . 171Cornus sericea L. . . . . . . . . . . . . . . . . . . . . . . 173Crataegus spp. L. . . . . . . . . . . . . . . . . . . . . . . 147Dogwood, Alternate-Leaf (Blue) . . . . . . . 172

Dogwood, Flowering . . . . . . . . . . . . . . . . . 171Dogwood, Red Osier . . . . . . . . . . . . . . . . . 173Elm, American . . . . . . . . . . . . . . . . . 136–137Elm, Slippery . . . . . . . . . . . . . . . . . . . . . . 137Fagus grandifolia Ehrh. . . . . . . . . . . . . 104–105Fir, Balsam . . . . . . . . . . . . . . . . . . . . . . 46–47Fraxinus americana L. . . . . . . . . . . . . . 128–129Fraxinus nigra Marsh. . . . . . . . . . . . . . 132–133Fraxinus pennsylvanica Marsh. . . . . . . 130–131Gleditsia triacanthos L. . . . . . . . . . . . . . . . . . 164Gum, Black . . . . . . . . . . . . . . . . . . . . 160–161Hackmatack . . . . . . . . . . . . . . . . . . . . . 50–51Hamamelis virginiana L. . . . . . . . . . . . . . . . 167Hawthorn (Thorn-Apple) . . . . . . . . . . . . . 147Hemlock, Eastern . . . . . . . . . . . . . . . . . 48–49Hickory, Bitternut . . . . . . . . . . . . . . . . . . . 154Hickory, Shagbark . . . . . . . . . . . . . . . 152–153Honeylocust . . . . . . . . . . . . . . . . . . . . . . . 164Hophornbeam, Eastern (Ironwood) . . . 98–99Hornbean, American (Blue-beech,

Musclewood) . . . . . . . . . . . . . . . . . 100–101Horsechestnut . . . . . . . . . . . . . . . . . . . . . . 158Ironwood . . . . . . . . . . . . . . . . . . . . . . . . 98–99Ivy, Poison (Mercury) . . . . . . . . . . . . . . . . . . 6Juglans cinerea L. . . . . . . . . . . . . . . . . . 155–156Juglans nigra L. . . . . . . . . . . . . . . . . . . . . . . . 157Juniper, Common . . . . . . . . . . . . . . . . . . . . 58Juniperus communis L. . . . . . . . . . . . . . . . . . . 58Juniperus virginiana L. . . . . . . . . . . . . . . 56–57Kalmia latifolia L. . . . . . . . . . . . . . . . . . . . . 169Larch, Eastern . . . . . . . . . . . . . . . . . . . . 50–51Larix laricina (Du Roi) K. Koch . . . . . . 50–51Laurel, Mountain . . . . . . . . . . . . . . . . . . . 169Linden . . . . . . . . . . . . . . . . . . . . . . . . 134–135Locust, (Honeylocust) . . . . . . . . . . . . . . . . 164Locust, Black . . . . . . . . . . . . . . . . . . . 162–163Maple, Ashleaf (Boxelder) . . . . . . . . . . . 84–85Maple, Mountain . . . . . . . . . . . . . . . . . 80–81Maple, Norway . . . . . . . . . . . . . . . . . . . 82–83Maple, Red (Soft, White, Swamp) . . . . 72–73Maple, Silver . . . . . . . . . . . . . . . . . . . . 76–77Maple, Striped (Moosewood) . . . . . . . . 78–79Maple, Sugar (Rock or Hard) . . . . . . . . 74-75Nannyberry . . . . . . . . . . . . . . . . . . . . . . . . 166Nyssa sylvatica Marsh. . . . . . . . . . . . . . 160–161Oak, Bear (Scrub) . . . . . . . . . . . . . . . 114–115Oak, Black (Yellow) . . . . . . . . . . . . . . 110–111Oak, Bur . . . . . . . . . . . . . . . . . . . . . . 120–121Oak, Chestnut . . . . . . . . . . . . . . . . . . 118–119Oak, Northern Red . . . . . . . . . . . . . . 108–109Oak, Scarlet . . . . . . . . . . . . . . . . . . . . 112–113Oak, Swamp White . . . . . . . . . . . . . . 122–123Oak, White . . . . . . . . . . . . . . . . . . . . 116–117Ostrya virginiana (P. Mill.) K. Koch . . . 98–99Picea abies (L.) Karst. . . . . . . . . . . . . . . . . . . . 44

S I

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Rhododendron maximum L. . . . . . . . . . . . . . 170Rhododendron, Rosebay (Great Laurel) . . 170Rhus hirta (L.) Sudworth . . . . . . . . . . . . . . 168Robinia pseudoacacia L. . . . . . . . . . . . . 162–163Salix spp. L./Salix nigra Marsh . . . . . . . . . . . 68Sassafras albidum (Nutt.) Nees . . . . . . . . . . 165Sassafras . . . . . . . . . . . . . . . . . . . . . . . . . . 165Serviceberry . . . . . . . . . . . . . . . . . . . . 148–149Sorbus spp. . . . . . . . . . . . . . . . . . . . . . . 150–151Sorbus americana Marsh. . . . . . . . . . . . 150–151Sorbus decora (Sarg.) . . . . . . . . . . . . . . . 150-151Spruce, Black . . . . . . . . . . . . . . . . . . . . . 38–39Spruce, Blue . . . . . . . . . . . . . . . . . . . . . . . . 45Spruce, Norway . . . . . . . . . . . . . . . . . . . . . . 44Spruce, Red . . . . . . . . . . . . . . . . . . . . . . 40–41Spruce, White (Cat) . . . . . . . . . . . . . . . 42–43Sumac, Poison . . . . . . . . . . . . . . . . . . . . . . . . 6Sumac, Staghorn . . . . . . . . . . . . . . . . . . . . 168Sycamore, American (Buttonwood,

Planetree) . . . . . . . . . . . . . . . . . . . . . . . . 159Tamarack (Eastern Larch) . . . . . . . . . . . 50–51Thuja occidentalis L. . . . . . . . . . . . . . . . . . 54–55Tilia cordata Mill. . . . . . . . . . . . . . . . . . . . . . 135Tilia europaea L. . . . . . . . . . . . . . . . . . . . . . . 135Tilia americana L. . . . . . . . . . . . . . . . . 134–135Tsuga canadensis (L.) Carr. . . . . . . . . . . . 48–49Tupelo, Black (Black Gum) . . . . . . . . 160–161Ulmus Americana L. . . . . . . . . . . . . . . . 136–137Ulmus rubra Muhl . . . . . . . . . . . . . . . . . . . . 137Viburnum lentago L. . . . . . . . . . . . . . . . . . . . 166Walnut, Black . . . . . . . . . . . . . . . . . . . . . . 155Willows/Black Willow . . . . . . . . . . . . . . . . 68Witch-Hazel . . . . . . . . . . . . . . . . . . . . . . . 167

Picea glauca (Moench) Voss . . . . . . . . . . . . 42–43Picea mariana (P. Mill.) B.S.P. . . . . . . . . 38–39Picea pungens Engelm. . . . . . . . . . . . . . . . . . . 45Picea rubens Sarg. . . . . . . . . . . . . . . . . . . . 40–41Pine, Eastern White . . . . . . . . . . . . . . . 26–28Pine, Jack (Gray) . . . . . . . . . . . . . . . . . . 32–33Pine, Pitch . . . . . . . . . . . . . . . . . . . . . . . 34–35Pine, Red (Norway) . . . . . . . . . . . . . . . . 30–31Pine, Scots (Scotch) . . . . . . . . . . . . . . . . . . . 36Pinus banksiana Lamb. . . . . . . . . . . . . . . 32–33Pinus resinosa Soland. . . . . . . . . . . . . . . . 30–31Pinus rigida P. Mill. . . . . . . . . . . . . . . . . . 34–35Pinus strobes L. . . . . . . . . . . . . . . . . . . . . . 26–28Pinus sylvestris L. . . . . . . . . . . . . . . . . . . . . . . 36Platanus occidentalis L. . . . . . . . . . . . . . . . . . 159Plum, Canada (Red) . . . . . . . . . . . . . . . . . 146Poplar (see Aspen)Poplar, Balsam . . . . . . . . . . . . . . . . . . . 66–67Populus balsamifera. L. . . . . . . . . . . . . . . . 66–67Populus grandidentata Michx. . . . . . . . . . 64–65Populus tremuloides Michx. . . . . . . . . . . . 62–63Prunus nigra Ait. . . . . . . . . . . . . . . . . . . . . . 146Prunus pensylvanica L. . . . . . . . . . . . . . 140–141Prunus serotina Ehrh. . . . . . . . . . . . . . 142–143Prunus virginiana L. . . . . . . . . . . . . . . 144–145Quercus alba L. . . . . . . . . . . . . . . . . . . . 116–117Quercus bicolor Willd. . . . . . . . . . . . . . 122–123Quercus coccinea Muenchh. . . . . . . . . . 112–113Quercus ilicifolia Wangenh. . . . . . . . . . 114–115Quercus macrocarpa Michx. . . . . . . . . . 120–121Quercus prinus L. . . . . . . . . . . . . . . . . . 118–119Quercus rubra L. . . . . . . . . . . . . . . . . . . 108–109Quercus velutina Lam. . . . . . . . . . . . . . 110–111Redcedar, Eastern . . . . . . . . . . . . . . . . . 56-57

Maine’s river drives continued until the 1970’s.

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Leaf measurements do not include the petiole of simple leaves.

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A N D R O S C O G G I N

A R O O S T O O K

C U M B E R L A N D

F R A N K L I N

H A N C O C K

K E N N E B E C

K N O XL I N C O L N

O X F O R D

P E N O B S C O T

P I S C ATA Q U I S

S O M E R S E T

WA L D O

WA S H I N G T O N

Y O R K

DENNIS BRENNAN

KEN CANFIELD

MORTON MOESSWILDE

J IM ECKER

STEVE MACDONALD

DAVE ROCHESTER

DAN JACOBS

PATTY CORMIER

GORDON MOORE

MERLE RING

SAGA-DAHOC

M A I N E D I S T R I C T F O R E S T E R SDennis Brennan [email protected] 324-7000Ken Canfield [email protected] 657-3535Patty Cormier [email protected] 474-3499Jim Ecker [email protected] 827-1811Dan Jacobs [email protected] 463-3653Steve MacDonald [email protected] 434-2622Morten Moesswilde [email protected] 549-9003Gordon Moore [email protected] 695-3721Merle Ring [email protected] 674-3787Dave Rochester [email protected] 435-7963

www.maineforestservice.gov

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