evolution, biodiversity, and population ecologytext+ch+3...copyright © 2006 pearson education,...

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Copyright © 2006 Pearson Education, Inc., publishing as Benjamin Cummings PowerPoint ® Slides prepared by Jay Withgott and Heidi Marcum PowerPoint ® Slides prepared by Jay Withgott and Heidi Marcum Ch 5 Evolution, Biodiversity, and Population Ecology Part 1: Foundations of Environmental Science Copyright © 2008 Pearson Education, Inc., publishing as Benjamin Cummings

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Copyright © 2006 Pearson Education, Inc., publishing as Benjamin Cummings

PowerPoint® Slides prepared by Jay Withgott and Heidi Marcum

PowerPoint® Slides prepared by Jay Withgott and Heidi Marcum

Ch 5Evolution, Biodiversity, and Population Ecology

Part 1: Foundations of Environmental Science

Copyright © 2008 Pearson Education, Inc., publishing as Benjamin Cummings

Copyright © 2008 Pearson Education, Inc., publishing as Benjamin Cummings

This lecture will help you understand:

• Natural selection • How evolution influences

biodiversity • Reasons for species extinction • Ecological organization • Population characteristics • Population ecology • Conservation biology

Copyright © 2008 Pearson Education, Inc., publishing as Benjamin Cummings

Striking gold in Costa Rica

• Golden toads were discovered in 1964, in Monteverde, Chile

• The mountainous cloud forest has a perfect climate for amphibians

• Unfortunately, they became extinct within 25 years - Due to global

warming’s drying effect on the forest

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Early Earth was a very different place Theory- dispersed bits of material whirling through space around our sun were drawn together by gravity

• 4.5 billion years ago, Earth was a hostile place - Severe volcanic and tectonic activity - Intense ultraviolet energy from the sun - No oxygen existed in the atmosphere, until photosynthesis

developed in microbes - No life existed

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Several hypotheses explain life’s origin

• Primordial soup (the heterotrophic hypothesis) - life originated from a “primordial soup” of simple inorganic chemicals in the oceans - First life forms used organic compounds for energy

• “Seeds” from space (the panspermia hypothesis) - microbes from space traveled on meteorites to Earth

• Life from the depths (the chemoautotrophic hypothesis) - life originated in deep-sea hydrothermal vents, with abundant sulfur - First organisms were chemoautrotrophs

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The fossil record teaches about life’s history

• Single-celled bacteria occurred on Earth 3 billion years ago

• Fossil - an imprint in stone of a dead organism

• Fossil record - gives information about the history of past life

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Present-day organisms help decipher history• Biologists use present-day

organisms to get information about evolution

• Archea - single-celled prokaryotes very different from bacteria

• The tree of life now consists of 3 prongs: bacteria, archaea, eukaryotes

• Prokaryote- a typically unicellular organism that lacks organelles and nucleus

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Evolution: the source of Earth’s biodiversity

• Biological evolution - genetic change in populations of organisms across generations - May be random or directed by natural

selection - Natural Selection – the process by which

traits that enhance survival and reproduction are passed on more frequently to future generations than those that do not

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Understanding evolution is vital

• It alters the genetic makeup of a population • It is important for understanding antibiotic

and pesticide resistance, agricultural issues, production, medicines, etc.

• Organisms adapt to their environment and change over time

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Natural selection shapes organisms

• In 1858, Darwin and Wallace both proposed natural selection as the mechanism of evolution - Organisms face a constant struggle to survive and reproduce - Organisms tend to produce more offspring than can survive - Individuals of a species vary in their characteristics due to genes and the

environment - Some individuals are better suited to their environment and will survive and

pass their genes on in their offspring

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Genetic variation

• Adaptive Trait (Adaptation) - a trait that promotes reproductive success

• Mutations - accidental changes in DNA that may be passed on to the next generation - Non-lethal mutations provide the genetic variation on which natural selection acts

• Sexual reproduction also leads to variation

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Natural selection acts on genetic variation

• Directional selection - drives a feature in one direction (a-thick shell protects snail)

• Stabilizing selection - produces intermediate traits, preserving the status quo (b-preserves extra resources for feeding and reproduction an favors an int. shell)

• Disruptive selection - traits diverge in two or more directions (c- two traits are favored. Thin shells conserve resources and out reproduce and thick shells are well protected)

If the environment changes, a trait may no longer be adaptive

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Evidence of natural selection is everywhere

• It is evident in every adaptation of every organism

• Evident in bacteria and fruit flies in laboratories

• Selective breeding of animals

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Natural selection

• Selective breeding has produced tremendous diversity among organisms

• Adaptive radiation- burst of species formation due to natural selection

Statistics 1.5 to 1.8 million species have been discovered Estimated 100 million species mat exist

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Artificial selection

• Artificial Selection - the process of selection conducted under human direction - For example, artificial selection has led to the great variety of dog breeds, and

agricultural crops

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Evolution generates biodiversity

• Biological Diversity - An area’s sum total of all organisms - The diversity of species - Their genes - Their populations - Their communities

• Species - a population or group of populations whose members share characteristics and can freely breed with one another and produce fertile offspring

• Population - a group of individuals of a species that live in the same area

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Speciation produces new types of organisms

• The process of generating new species - A single species can generate

multiple species • Allopatric speciation - species

formation due to physical separation of populations over some geographic distance

• DNA mutation arises in the isolated population - Can be separated by glaciers, rivers,

mountains, migration - The main mode of species creation

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Genetic Drift occurs when changes in the frequency of a gene in a population are due not to mutation, selection or migration but simply to chance

Founder Effect (allopatric) - occurs when a small number of individuals are isolated from a larger population - They may have a smaller genetic variation than the original species Both genetic drift and founder effect may not be better adapted to their environment

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Another type of speciation

• Sympatric speciation - species form from populations that become reproductively isolated within the same area - Feed in different areas - Mate in different seasons - Hybridization between two species

- Mutations

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The fossil record shows that…

• Earlier organisms evolved into later ones • The vast majority of species are extinct • Numbers of species increase over time • Earlier organisms were smaller and simpler • Several mass extinctions have occurred • Large, complex organisms occurred 600 million years

ago - Plants, animals, fungi

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Speciation results in diverse life forms

• Speciation generates complex patterns of diversity above the species level

• Phylogenetic trees (Cladograms) - Represents the history of species divergence - Scientists can trace when certain traits evolved - Show relationships between species

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Extinction

• Species generally evolve from simple to complex and small to big, but the opposite can occur, and some even disappear

• Extinction - the disappearance of a species from Earth - Occurs when a species cannot

adapt quickly enough to a changing environment

- Speciation and extinction affect species numbers

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Extinction is a natural process

• Extinction is irreversible: once a species is lost, it is lost forever

• Humans profoundly affect rates of extinction

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Some species are more vulnerable to extinction

• Extinction occurs when the environment changes too rapidly for natural selection to keep up

• Endemic species - a species only exists in a certain, specialized area (golden toad to the Monteverde cloud forest) - Very susceptible to extinction - These species usually have small populations

• Many other factors also cause extinction - Severe weather - New species - Specialized species

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Earth has had several mass extinctions

• Background extinction rate –the average rate of extinction that occurred before the appearance of humans

• Fossil record indicates that for both bird and mammals one species in the world typically becomes extinct every 500-1000 years or

• One to five species per million species on earth extinction usually occurs one species at a time

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Extinction

• Mass extinction events – a catastrophic widespread event where large groups of species are wiped out in a geological period lasting up to 5 million years

• five events in Earth’s history that killed off massive numbers of species at once (20-60 million years apart during the past 500 million years)

• 250 million years ago, 50-95% of all species went extinct at one time

• Humans are causing the sixth mass extinction event - Resource depletion - Population growth - Development

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Causes of Extinction

• Risk- the chance that a species of population will become extinct owing to one of these causes

1- Population Risk- random variations in population rates (in birth rates and death rates) can cause a species low in abundance to become extinct (butterfly in Colorado mountains only lay their eggs in the buds of a lupine tree. Their was a late snow killing all the buds)

2- Environmental Risk- changes in the environment that occur. This includes variations in the physical and biological environment

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Causes of Extinction – cont.

3- Natural Catastrophe- sudden changes in the environment that is not a result of human activities (volcano, earthquake, flood)

4- Genetic Risk- changes in genetic characteristics, not caused by external environmental changes (species lacks variety and may be vulnerable)

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Species’ ranges can be severely restricted

Some U.S. salamander species live on top of single mountains

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Endangered Species- Red List• The International Union for Conservation of Nature (IUCN) is

the world's main authority on the conservation status of species. A series of Regional Red Lists are produced by countries or organizations, which assess the risk of extinction to species

• The percentage of species in several groups which are listed as red critically endangered, orange endangered, or yellow vulnerable on the 2007 IUCN Red List

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Ecology is studied at several levels

• Ecology and evolution are tightly intertwined

• Biosphere - the total living things on Earth and the areas they inhabit

• Ecosystem - communities and the nonliving material and forces they interact with

• Community - interacting species that live in the same area

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Levels of ecological organization

• Population ecology - investigates the quantitative dynamics of how individuals within a species interact

• Community ecology - focuses on interactions among species

• Ecosystem ecology - studies living and nonliving components of systems to reveal patterns - Nutrient and energy flows

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Organismal ecology: habitat

• Habitat - the environment in which an organism lives - Includes living and nonliving elements - Scale-dependent: from square meters to miles - each organism thrives in certain habitats, but not in others (habitat use)

• Habitat selection - the process by which organisms actively select habitats in which to live - Availability and quality of habitat are crucial to an organism’s well-being - Human developments conflict with this process

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Organismal ecology: niche

• Niche - an organism’s use of resources and its functional role in a community - total way of life or role of a species in an ecosystem - Habitat use, food selection, role in energy and nutrient flow - Interactions with other individuals

• Specialists - species with narrow niches and very specific requirements - Extremely good at what they do, but vulnerable to change

• Generalists = species with broad niches that can use a wide array of habitats and resources - Able to live in many different places

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Population characteristics

• All populations show characteristics that help scientists predict their future dynamics

• Population size - the number of individual organisms present at a given time - Numbers can increase, decrease, cycle

or remain the same

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Population characteristics

• Population density - the number of individuals within a population per unit area - High densities make it easier to find mates, but increase competition, and vulnerability

to predation - Low densities make it harder to find mates, but individuals enjoy plentiful resources

and space Population dynamics – the study of how the characteristics of the population change in

response to changes in the environmental conditions

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Population characteristics

• Population distribution (dispersion) - spatial arrangement of organisms within an area - Random – haphazardly located individuals,

with no pattern - Uniform – individuals are evenly spaced due

to territoriality - Clumped – arranged according to availability

of resources - Most common in nature

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• Sex ratio - proportion of males to females - In monogamous species, a 50/50 sex ratio maximizes

population growth • Age Structure - the relative numbers of organisms of each age

within a population - Age structure diagrams (pyramids) - show the age structure

of populations

Population characteristics

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Birth and death rates

• Crude birth/death rates - rates per 1000 individuals

• Survivorship curves - the likelihood of death varies with age - Type I: More deaths at older ages - Type II: Equal number of deaths at all

ages - Type III: More deaths at young ages

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Four factors of population change

• Natality - births within the population • Mortality - deaths within the population • Immigration =-arrival of individuals from outside the

population • Emigration =-departure of individuals from the

population • Growth rate formula -

- (Crude birth rate + immigration rate) - (Crude death rate + emigration rate) - Growth rate

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Exponential population growth

• Steady growth rates cause exponential population growth - Something increases by a fixed percent - Graphed as a J-shaped curve

• Exponential growth cannot be sustained indefinitely - It occurs in nature with a small population

and ideal conditions

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Limiting factors restrain growthIntrinsic rate of increase (r) is the rate at which the population of a

species would grow if it had unlimited resources - Reproduce early in life - Have short generation times - can reproduce many times and have many offspring

• Limiting factors - physical, chemical and biological characteristics that restrain population growth - Water, space, food, predators, and disease

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Carrying capacity

• Carrying capacity (K)- the maximum population size of a species that its environment can sustain - An S-shaped logistic growth curve - Limiting factors slow and stop

exponential growth

• Carrying capacity changes

Humans have raised their carrying capacity by decreasing the carrying capacity for other species

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Perfect logistic curves aren’t often found

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Population density affects limiting factors

• Density-dependent factors - limiting factors whose influence is affected by population density - Increased risk of predation and competition for mates occurs with increased density

• Density-independent factors - limiting factors whose influence is not affected by population density - Events such as floods, fires, and landslides

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Biotic potential and reproductive strategies vary• Biotic potential - the ability of an organism to produce

offspring • K-selected species - animals with long gestation periods

and few offspring - Have a low biotic potential - Stabilize at or near carrying capacity - Good competitors

• r-selected species - animals which reproduce quickly - Have a high biotic potential - Little parental care

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K-selected vs. r-selected species

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Population changes affect communities

• As population in one species declines, other species may appear

• Human development now displaces other species and threatens biodiversity - As Monteverde dried out, species from lower, drier habitats appeared - But, species from the cloud-forest habitats disappeared

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Challenges to protecting biodiversity

• Social and economic factors affect species and communities - Nature is viewed as an obstacle to development - Nature is viewed as only a source of resources - Human population growth pressures biodiversity

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Preserving biodiversity

• Natural parks and protected areas help preserve biodiversity - Often, they are underfunded - Ecotourism brings jobs and money to developing areas