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  • About IUCN

    IUCN is a membership Union composed of both government and civil society organisations. It harnesses the experience, resourcesand reach of its 1,300 Member organisations and the input of some 15,000 experts. IUCN is the global authority on the statusof the natural world and the measures needed to safeguard it.

    www.iucn.orghttps://twitter.com/IUCN/

    IUCN The Species Survival Commission

    The Species Survival Commission (SSC) is the largest of IUCNs six volunteer commissions with a global membership of more than10,000 experts. SSC advises IUCN and its members on the wide range of technical and scientific aspects of species conservationand is dedicated to securing a future for biodiversity. SSC has significant input into the international agreements dealing withbiodiversity conservation.

    http://www.iucn.org/theme/species/about/species-survival-commission-ssc

    IUCN Global Species Programme

    The IUCN Species Programme supports the activities of the IUCN Species Survival Commission and individual Specialist Groups,as well as implementing global species conservation initiatives. It is an integral part of the IUCN Secretariat and is managed from IUCNsinternational headquarters in Gland, Switzerland. The Species Programme includes a number of technical units covering SpeciesTrade and Use, the IUCN Red List Unit, Freshwater Biodiversity Unit (all located in Cambridge, UK), the Global Biodiversity AssessmentInitiative (located in Washington DC, USA), and the Marine Biodiversity Unit (located in Norfolk, Virginia, USA).

    www.iucn.org/species

    IUCN Centre for Mediterranean Cooperation

    The Centre was opened in October 2001 with the core support of the Spanish Ministry of Agriculture, Fisheries and Environment,the regional Government of Junta de Andaluca and the Spanish Agency for International Development Cooperation (AECID). IUCNhas over 203 members in the Mediterranean region, including 15 governments. Its mission is to influence, encourage, and assistMediterranean societies to conserve and use sustainably the natural resources of the region and work with IUCN members andcooperate with all other agencies that share the objectives of the IUCN.

    www.iucn.org/mediterranean

  • THE STATUS AND DISTRIBUTIONOF MEDITERRANEANBUTTERFLIES

    Catherine Numa, Chris van Swaay, Irma Wynhoff, Martin Wiemers, Violeta Barrios,David Allen, Catherine Sayer, Miguel Lpez Munguira, Emilio Balletto, Dubi Benyamini,Stoyan Beshkov, Simona Bonelli, Robert Caruana, Leonardo Dapporto, Filip Franeta,Patricia Garcia-Pereira, Evrim Karaetin, Ahmad Katbeh-Bader, Dirk Maes, Nikola Micevski,Rebecca Miller, Eva Monteiro, Riadh Moulai, Ana Nieto, Lazaros Pamperis, Guy Peer,Andrew Power, Martina aic, Katy Thompson, Elli Tzirkalli, Rudi Verovnik, Martin Warrenand Hilary Welch

    The IUCN Red List of Threatened SpeciesTM Regional Assessment

  • Compilers:

    Catherine Numa Mediterranean Species Programme, IUCN Centre for Mediterranean Cooperation, SpainChris van Swaay De Vlinderstichting, Butterfly Conservation Europe, NetherlandsIrma Wynhoff De Vlinderstichting, NetherlandsMartin Wiemers Helmholtz Centre for Environmental Research - UFZ, GermanyVioleta Barrios Mediterranean Species Programme, IUCN Centre for Mediterranean Cooperation, SpainDavid Allen Red List Unit, IUCN Global Species and Key Biodiversity Areas Programme, United KingdomCatherine Sayer Freshwater Biodiversity Unit, IUCN Global Species and Key Biodiversity Areas Programme, United KingdomMiguel Lpez Munguira Department of Biology, Universidad Autnoma de Madrid, SpainEmilio Balletto Department of Life Sciences and Systems Biology, University of Turin, ItalyDubi Benyamini Israeli Lepidopterists Society, IsraelStoyan Beshkov Bulgarian National Museum of Natural History, BulgariaSimona Bonelli Department of Life Sciences and Systems Biology, University of Turin, ItalyRobert Caruana University of Malta, MaltaLeonardo Dapporto Freelance, ItalyFilip Franeta Institute for field and vegetable crops, Novi Sad, SerbiaPatricia Garcia-Pereira Museu Nacional de Histria Natural e da Cincia and Tagis, PortugalEvrim Karaetin Doga Koruma Merkezi, TurkeyAhmad Katbeh-Bader Department of Horticulture and Plant Protection, Faculty of Agriculture. University of Jordan, JordanDirk Maes Research Institute for Nature and Forest, BelgiumNikola Micevski Macedonian Entomological Society, MacedoniaRebecca Miller IUCN Global Ecosystem Management Programme, United KingdomEva Monteiro Museu Nacional de Histria Natural e da Cincia and Tagis, PortugalRiadh Moulai Universit de Bjaia, AlgeriaAna Nieto European Biodiversity conservation, IUCN Global Species and Key Biodiversity Areas Programme, BelgiumLazaros Pamperis Freelance, GreeceGuy Peer Department of Conservation Biology, Helmholtz Centre for Environmental Research - UFZ, GermanyAndrew Power School of Life Sciences, Nottingham University, United KingdomMartina ai Department of Zoology, Croatian Natural History Museum, CroatiaKaty Thompson School of Life Sciences, Nottingham University, United KingdomElli Tzirkalli Cyprus Butterfly Study Group, CyprusRudi Verovnik Department of Biology University of Ljubljana, SloveniaMartin Warren Butterfly Conservation UK, United KingdomHilary Welch Doga Koruma Merkezi, Turkey

    The designation of geographical entities in this report and the presentation of the material do not imply the expression of any opinion whatsoever on the part of IUCNor other participating organizations concerning the legal status of any country, territory, or area, or its authorities, or concerning the delimitation of its frontiers orboundaries.

    The views expressed in this publication do not necessarily reflect those of IUCN or other participating organizations.

    Published by: IUCN, Malaga, SpainCopyright: 2016 International Union for Conservation of Nature and Natural Resources

    Reproduction of this publication for educational or other non-commercial purposes is authorized without prior written permissionfrom the copyright holder provided the source is fully acknowledged.

    Reproduction of this publication for resale or other commercial purposes is prohibited without prior written permission of thecopyright holder.

    Red List logo: 2008Citation: Numa, C., van Swaay, C., Wynhoff, I., Wiemers, M., Barrios, V., Allen, D., Sayer, C., Lpez Munguira, M., Balletto, E.,

    Benyamini, D., Beshkov, S., Bonelli, S., Caruana, R., Dapporto, L., Franeta, F., Garcia-Pereira, Karaetin, E., Katbeh-Bader, A.,Maes, D., Micevski, N., Miller, R., Monteiro, E., Moulai, R., Nieto, A., Pamperis, L., Peer, G., Power, A., ai, M., Thompson, K.,Tzirkalli, E., Verovnik, R., Warren, M. and Welch, H. (2016). The status and distribution of Mediterranean butterflies. IUCN,Malaga, Spain. x + 32 pp.

    ISBN: 978-2-8317-1790-6DOI: http://dx.doi.org/10.2305/IUCN.CH.2016.MRA.6.enCover photo: The Bollands Blue (Polyommatus bollandi) only occurs in Hatay (Turkey) and is considered Critically Endangered. It is threatened

    by urbanization, quarrying and overgrazing. Ali Atahan

    Layout by: Simtrica, S. L.Printed by: Solprint S. L.Available from: IUCN Centre for Mediterranean Cooperation

    C/ Marie Curie 2229590 Campanillas, Malaga, Spain. Tel: +34 952 028430 Fax: +34 952 028145 http://www.iucn.org/mediterranean http://www.iucn.org/resources/publications

    The text of this book is printed on 115 gsm environmentally-friendly paper.

  • Table of contents

    Acknowledgements . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . vForeword . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . viiExecutive summary . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ix

    Chapter 1. Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11.1. The Mediterranean region . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11.2. Overview of the regional butterfly fauna . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 11.3. Objectives of the regional assessment . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3

    Chapter 2. Assessment methodology . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 42.1. The IUCN Red List of Threatened Species .

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