96 2004 ,4 Éj 1j* 112 t!ljâjiwebagris.inra.org.ma/doc/awamia/11208.pdf · 2011-02-11 · 96 2004...
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96 2004 ,4 ÉJ 1J* 112 t ! " l jâJ I
30- Saneoka, H., Shir:ta, K., Kurban, H., Chaudhary, M.1., Premachandra, G. S., and K.Fuiita 1999. Hffect of Slirrity on Growth and Solute Accumulation in Two Wheat LinesDiffering in Salt'folerance. Soil Sci. Plant Nutrition. 45 (4), 873-880.
3l- Santivieri, F., Romagosa, I., Royo, C. 2001. Assessing genotypic variability fbr plantdevelopment in spring and winter triticale. Cereal Research Communications 29 (3/4)359-366.
32- Siddiqi, MY., Glass ADM. 1983. Studies on the growth and mineral nutrition of barleyvarieties. I. Effect of potassium supply on the uprake of porassium and growth.Canadian Journal ol'Botenica. 61: 67I-678.
33- Slama, F. 1986. Effet du nitrate d'ammonium sur Ie degré de tolérance à une forte dosede NaLll de dix variétés de blé. Colloque sur les Végétaux cn Milieu Aride. FST/ACCT.Tunisie (Jerba), pp. a60-a73.
34- Stewart, C.R., Lee, J.A. 1974. The role of proline acôumulation in tralophytes. Planta;rZ0:279-289.
35- Steel, C. ancl Torrie, J.H. 1960. Principles and procedures of statistics. Mc Grow- HillBook Co., Inc., New York. 200pp.
36- Storey, IL., Wyn Jones. RG. 1978. Salt stress and comparative physiology in theGratninaea. L Ion relations of two salt- and water-stressed barley cultivars, CalifbrniaMariout and Arimar. Australian Jorlmal Plant Physiology . 5: 801-816.
37- Touraine, B. et Ammar, M" 1985- Etude comparée dc la sensibilité arr sel d'un triticaleet d'une orge. Agronomie, 5; 39i-395.
38- Wyn Jones, R. G. and Gorham, J. Mcl)onnell E. 1984. Organic and inorganic solutecontents as selection criteria fbr salt tolerance in the Triticeae. In: Staples RC,Toennienssen GA (eds.) : Salinity 'l'olerance in Plants. Strategies for Crophnprovement. Wiley, New Yor:k, pp. l89-?03.
39- Wyn Jones, R. G. and Gorham, J. 1989. Use of physiological traits in breeding forsalinity tolerance. In: Drought Resistance in Cereals, Baker F.W.G. (ed.). CABInternational, ïValli ngford.
40- Wyn.Iones, R. G., and R. Storey. 1978. Salt Stress and Comparative Physioiogy in theGramineae. IL Glycinebetaine and Proline Accumulation in Two Salt and Water-strcssed BarJey Cuitivars. Australian Joru'nal Plant Physiology. 5: 817-829.
4l-Zid, E., et Grignon, C.l99l- Les testrj de sélection précoce pour la résistance desplanles aux stress. Cas des strsss salin et hydrique. In: L amelioration des plantes pourI'adaptation aux nrilicux alides. Ed. ALIPELF-UREF. John Libbey Eurotext. Paris,I 1 - r08 .
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3- Ali Dib, T., and Monneveux, Ph. 1992. Adaptation à la sécheresse Êt notion d'idéotypechez le blé drrr:. I. Caractôrcs morphologiques, onracinement. Agronomie, I2, 371-319.
4- Arbaoui, M. Benkhelifa, M. and M. Belkhodja. 2000. Réponses physiologiques dequelques variéïés de blé dur à la salinité au stade juvénile, In Option Meditelraneennes.L amelioration du blé dur dans la région mediterraneenne; Nouvear"rx defis : Eds C.Royo, M.M. Nachit, N.DI Fonzo, J.L. Àraus. (CIHËAM;Centre Udl - IRIA, CIMMYIICARDA), p267-270.
5- Benlaribi, M., et Monneveux, Fh. 1988. Etude comparée du comportement en situationde déficit hydrique de deux variétds algériennes de blé dur (T'riticum durum Desf.)adaptées à la sécheresse. C" R. Acad. Agri. France, 74:'73-83.
6-Bizid, E,Zid, E., Grignon, C. 1988. Tolérance à NaCl et sélectivité K+/Irla+ chez lestritiÇales. Agronomie; 8: 23-27 .
7- Cheeseman, J. M. 1988. Mechanisrrs of salinity tolerance in plants. Plant Physiology.87:547-550.
8- Cholick, F.A., Welsh, J.R., Vernon-Cole, C. 1977. Rooting pattelns of semi-dwarf andtall wheat cultivars r-rnder dryland field conditions. Crop Scienc e. 17 , 637 -640.
9-Ë,hert, D.L., Redmann, R.E., Harvey, B.L. et Cipywnyk, A. 1990- Salinity-inducedcalcium deficiencies in wheat and barley. Plant Soil, 128: 143-I51.
t0- Erdei, L. and Taleisnik, E. 1993- Changes in water relation parameters under osmoticand stresses in maize and sorghum. Plant Physiology, 89, 38 1-387.
11- Francois, L. E., and Maas, E. V. 1985. Plant responses to salinity; An indexedbibliography. Science and Edr-rcation Administration, U.S. Department of Agricuiture,Berkeley, Califomia.
12-Fowler, D. B. and Hamm, J. W. I9B0- Crop response to saline soil conditions in theparkland area ofSaskatchewan. Canadian Journal ofSoil Science. 60:439-449.
13- Greenway, H. and Munns, R. 1980, Mechanism of salt tolerance in non halophytes.Annual Review Plant Physiology. 31: 149-190.
14- Jensen, P., Petterson, S. 1980" Varietal variation in uptake and utilisation of potassillm(rubidiurn) in high-salt secdling of barley. Plant Physiol<lgy. 48: 4l I-415.
t5- Koryo, H. W. 1997. Ultrastructurai and physiological changes in root cells of sorghumplants (Sorghum bicolor x Sorghum sudânesis cv. Sweet Sioux) induced by NaCl. J.Exu. Bot." 48.693-706.
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13 ,4 _ oia gétaliyl 4J.É jgl i'.L C C,+= ol . (ratlt dg.,,r, r:T 45JJJ| g!* gl il;H!)
dJr;..^ +Li dLra 6{s9 . 07 Drira Cross $.c (VoL69*fi NDITYN 22 r3.e a/o 141309 C ÉiÊ. (2 J,gr+) {^ el"Jl Ërt--rj rljl . Àr.i.. Çls+ Ël.lr?r-l g,T)tsJl ôb J*d+ Ls (5jiÂ'i {agJa x âJ}r'*
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i-/iJ"JlJ" 2,-5 (Flogiand solution) $)tÊJl J3'l+7 control 'raLill -l
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.(p*/ir"+l+"z1 ,-J! .Jr-$ saL J#5r + u)'tÀ5ra J$5") f,-/iJ"JlJ" 2l ÉJisâ -4
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JJb,/ {i,irâJJt c.Li3a'Yl3 d;cr çiLâJl dî=aJl J+lta-t.ÉEÀôl À.é,-ôÀêll crt-r5-ll q,o]5'^*.ùSJ=lf c.r,r;kfu; ,3J!+ )t* i+,lr-aJl crlii1Il Ji.tÂ3
"É 'a*s ' ;;[3ç CI+IJ 2B
ili.é3ll +* g..rtr .J+rd ûS ç. 4ôf*sJr ltâJLr-ôlJ ï'll3 ;r^-yl Jlf'â3Jl t'-Ë' *i. ârÀ.3$sJl
2004 ,4 ÉJ l J.rF* ll2 J+"lt-ll
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olrr! p.rÀ3a;J . e3ÂJl \rrj+S ëf 4+Jrr= ffi ,t+,c. ,*itt 45,1i- ,.f ,.J-+ J4J . ,-till qiilJ LÉ+
,.éÀi ,-iJrs fr+ Ai ,-lÉ+J ù-al3Jl #X* d 4+J;s dir.3.a3 tJ .aÇSJ^r.llg;3tratl if" +rdl
urrlta fi)B $ll3s i,{ôJ"Yl Êl-rJ ChJl d ,Jt5$g'13Jl+.t"S.rtst 4sLMJl Ëri! .Àr,nLôJl 43ir't,+r"i ,.lt*iis, f+, errJl J+rJeJl ,-Fj li.,o .r+r*Jir l+s-rr t=r-lsl3+,J l4olâe-a tsr, .rtjsro
,li<a siii +0 ÂsS;Fl 4sLsôJl Ëri+ ++= d-ôæJl 13.è 4.ÊlJi (cy-.g.f -Élj+Jl .u*.iJl).(2001 Santiveri, et al,)
.Uf;" jSI i+'.+l,1s.a*Jl lia ;airç3 ,JtJâ.oTl 4+15 élbÉrJl ,p ++,54r. $'t+S fl+à-ill 9â+4irirl dÈt-r-oll d'^*,.,f
'tr:t*lir"i alri.tt fpC}.tiyl ÊSliiJ+5+J.6JÀyl çe+sJl J+,ol-.û"
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d e+JldôsiJ 4+sl#IlJÂ'fi- l l p.ai.r-ltre+;+r +++#s*;#r
ùl+J'*Jl ù. Cl_ _91 Na+ ol-r3'ri pl .ri..,l crBbiJl uéâ+ e+b3*: ,''.f- (Saneoka et al, i999)
6;l Jtar'.yl ËrLrl3 .JJHI *. ùp.Jls .JJ"+ll t+ÏÀ L"ix+.ri" eÉÀâ.ô é.ir3 J)tÀ Ûiô,:lJlJ.(Errtei and Taleisnik, 1993); (Koryo et al 1993); (Koryo, 1997) èlJÉl
,3f+ït9 ç5LJl ûjlJill J+ïri+ +IJ,ig À.sSJdJ.i.IôsiiJl J+Ê ÉËl!.ill $Ê J"ôiJl {=+tâJl ,"Llii
furalit Ô. Â*jr3. Ëratrill oia.g .élJJIl .Efu,rJs (Greenway etMunns, 1980) À-+.É)tJ
lâ4JJlJ Ét +-rll (]++ $Jl*Jl r+-5J' JrJj d qlËtl ';'À o ,(Et-l-uVt1 r$j=Jr ,'l*;g l3a .e.tntl +r d-'l+ y Jâ5rl elâ+ #*séÀJl u"t+ clS.,*t+tt ,L-ai [evigner et al, 199-5)
$+ LdS e.êi.Jl r:Ê .i.!sÂiJl3 J+tiJi 6"o.ù;{"iolL'r le."JËi elalJ eéÀll J"=I;'l+ !f.c;,lUi!teJJ.l Oa 4"lLi" ,iSTt *t'.,^Ïl : +Na.r3l3*ii.rb:irol g 6Jall .i"SLii" û++ Ë.lSi, È'Ë}tÂJl ùl . t+--ÀllJr i,^l urÀriil Ê+ e+= r.r,Àr..rl ËJUl sJÊ l,]a ,*Ë+h4g t+sl;Sl ,J +Na r;lo rll "rglgi çJfi ,.#l
FJtéJ AiJÉ q#l {:il L,rL*l .rJ*3 Na+ .,rlJù +o J+S ,1;J .5Js $'L*i .,J+ ÀsJJôJJ .Uô.s.3i
À.r.t'lrJl ut+r.3J r+t ,r.r. ^ alr[n ,:lJ.i r.," $lr3U 15J.ur.o ÉJq ùl dÉ# ùi! . +K pl+sEJÉJl .r;13*ii
a++ cl. . +]r[n gét*E3il *r l€i..,*J 4s.ÉYl S* {ÊJ- U++ JIL3.JI dhÀll r..r+-'+ tilsl ,y"tt ,iStg
dJ"rsâ oJs,,,i.lS.A+.i.,ll .53+r*â {r.l$ ,ciJÂJl .5Js.+ ,,iJ+.tSJl li-ô 4:i+ "uF
.,JJ-l
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elq Â,#sJl
rlj;Tl u+l L-+iS À.=-"iTlg ,(+$tSo'tFJ ,..rSi+) r'r,À.,!1 ,;+r +t-iy"Tl 45j+ 4*l.oe ';'r!titrgos;a:r: e+.s .o.sJrJl .rÉi")u.âffltl c,tfli d;râ! eÉ ô,lL+Ê (lj:l+Srl"" ,Ë.g+") Â+glÂJl
.4pl ,.,-ll cr|.ltJ.iJl {.ji{Sr-r AJâsirJl *|jlgJl ri,c . Ë q+ÀJ' çf.s'tl '4-o' Jls{fu ôlj-+r p3çl3r,eJl 'r;13*ii
105
AL AWATIIA I12. Vol. t-2. 200,1 106
ti+Jlf.fulg* d flÉ.ilf o.r r.rlÉ.rJl Jh/Él-r.t, 4+lJ.
+t . ol+J1 * d SJ
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u* a*l;j jb;t f,l .oriryl tiÀ ù,â..â t+;f.rÉ iroT a-3'l"tl 4.tÉ" 4Jlt* plrÀj""1 JËùt-i élJlJ
d3uai 4Jâ.J-ë qt+ ,.f K+/ Na+ 4lLirii$lg +Na p3A.r3r,eJl .r;lJ.il $l;gTt uk-r3ul3 rtrJl 4JLr
4dâJ. ';$lts a*.;i e-c {$l-r*Jl3 -lJilÂll û'd Ë'lI*r ÂJrÉÂJ (oJl-rJ- ill'rl 28 JJb) ùtsJl. AsJJÀU J*'iJl Jl=,r- J;Jlo3J +*L*Tl s,t{llgyl dl,pl 4-r-i-r (f/iS"+L 2I ,I4 ,7 . .,Ât*È)
.f,.Il.:ÂôJt Ë'y
sàl dl+loll q*gJçJiddll i{l+i"ùUl
(,Ct^*-t;Jrlt) Fl;r*.ell rr,D Ër$Ua dsrË
.8;slrll it*p s-.i Triticosecale WittmaclcX f
J+.e-Jl ,!.c. JLâ.Ê'J ++u ule .!;Ue
i,ln=Jl "l='-t*Jl f*3 .,l=,lrjJl irE
ir:Jfr Â*l;'
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:'
,,1,I
' f i :
:
, i f i.,:'-
' . .
2004 ' 4 o-!J 1 J-*-" ll2 i.*.lt;l
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